The Genus Salix
in Alaska and the Yukon
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The Genus Salix
in Alaska and the Yukon
National Museum of Natural Sciences Musée national des Sciences naturelles
Publications in Botany, No. 2 Publications de botanique, n° 2
Published by the Publié par les
National Museums of Canada Musées nationaux du Canada
Staff editor
Frances Smith
The Genus Salix
in Alaska and the Yukon
George W. Argus
©Crown copyrights reserved
Available by mail from the
National Museums of Canada
Marketing Services
Ottawa
K1A OM8
Catalogue No. NM95-9/2
National Museum of Natural Sciences
National Museums of Canada
Ottawa, Canada
1973
PO987654321
Y79876543
Printed in Canada
The research on which this study is based was
carried out prior to 1969.
Contents
List of Plates, vi
List of Figures, viii
List of Maps, ix
List of Tables, xii
Résumé, xiv
Summary, xv
Biographical Note, xvi
Preface, 1
Taxonomic Relationships, 4
Classification of Salix in Alaska and the Yukon, 5
Hybridization, 6
Polyploidy, 10
Ecology, 13
Eco-geographical Key, 18
Method of Study, 20
Systematic Treatment, 23
Key to the Species, 25
Section 1.Pentandrae (Borr.) Schneid., 35
Section 2. Maccallianae Argus, 38
Section 3. Subalbae Koidz., 41
Section 4. Longifoliae Anderss., 43
Section 5. Chamaetia Dumort., 45
Section 6. Retusae Kern., 49
Section 7. Glaucae Pax, 81
Section 8. Myrtosalix Kern., 107
Section 9. Myrtilloides Koehne, 113
Section 10. Hastatae Kern., 119
Section 11. Cordatae Barr. ex Hook., 121
Section 12. Vetrix Dumort., 149
Section 13. Arbuscella Ser. ex Duby, 157
Section 14. Lanatae Koehne, 171
Section 15. Villosae Rouy, 177
Section 16. Sitchenses Bebb, 191
Appendices
A. Key to Codes, 193
B. Specimens Cited, 197
Bibliography, 271
Index to Salix, 275
List of Plates
|
Salix lasiandra Benth., 36
II
Salix maccalliana Rowlee, 40
III
Salix interior Rowlee, 42
IV
Salix reticulata L., 46
V
Salix polaris Wahl., 50
VI
Salix rotundifolia Trautv., 54
VII
Salix nummularia Anderss., 60
VIII
Salix phlebophylla Anderss., 62
IX
Salix ovalifolia Trautv., 64
X
Salix stolonifera Cov., 74
XI
Salix setchelliana Ball, 78
XII
Salix arctica Pall., 80
XIII
Salix sphenophylla Skvortsov, 86
XIV
Salix brachycarpa Nutt., 90
XV
Salix glauca L., 96
XVI
Salix athabascensis Raup, 102
XVII
Salix chamissonis Anderss., 106
XVIII
Salix arctophila Cock. ex Heller, 108
XIX
Salix fuscescens Anderss., 112
XX
Salix pedicellaris Pursh, 114
vi
List of Plates
XXI
Salix hastata L., 118
XXIl
Salix rigida Muhl., 122
XXIII
Salix monticola Bebb, 126
XXIV
Salix barclayi Anderss., 128
XXV
Salix hookeriana Barr., 132
XXVI
Salix commutata Bebb, 136
XXVII
Salix novae-angliae Anderss., 138
XXVIII
Salix myrtillifolia Anderss., 142
XXIX
Salix pyrifolia Anderss., 144
XXX
Salix bebbiana Sarg., 148
XXXI
Salix scouleriana Barr., 152
XXXII
Salix planifolia Pursh ssp. planifolia, 156
XXXIll
Salix planifolia Pursh ssp. pulchra (Cham.) Argus, 160
XXXIV
Salix arbusculoides Anderss., 168
XXXV
Salix lanata L. ssp. richardsonii (Hook.) Skvortsov, 170
XXXVI
Salix barrattiana Hook., 174
XXXVII
Salix candida Fliigge ex Willd., 176
XXXVIII
Salix alaxensis (Anderss.) Cov., 180
XXXIX
Salix drummondiana Barr., 184
XL
Salix sitchensis Sanson, 190
vii
List of Figures
1
Summary data card, 21
2
The variation of some morphological characteristics in Salix
ovalifolia, 72
3
The variation in pistil indumentum in Salix ovalifolia, 73
4
The variation in stipule and petiole length in Salix barclayi, 134
5
A comparison of stipule and petiole length variation in Salix
planifolia, 165
6
A representative chromatogram of crude leaf extracts in Salix
planifolia, 166
Vili
List of Maps
1
Salix Hybrids, 7
2
Salix Hybrids and Intergrades, 8
3
Salix Intergrade, 9
4
Salix lasiandra, 38
5
Salix maccalliana, 41
6
Salix interior, 44
f
Salix reticulata ssp. reticulata, 48
8
Salix reticulata ssp. glabellicarpa, 49
9
Salix polaris, 52
10
Salix rotundifolia ssp. rotundifolia, 56
11
Salix rotundifolia ssp. dodgeana, 57
12
Salix nummularia, 61
ia
Salix phlebophylla, 65
14
Salix ovalifolia var. ovalifolia, 67
5
Salix ovalifolia var. arctolitoralis, 68
16
Salix ovalifolia var. cyclophylla, 69
17
Salix ovalifolia var. glacialis, 70
18
Salix stolonifera, 76
19
Salix setchelliana, 79
20
Salix arctica, 83
List of Maps
21
Salix sphenophylla, 89
22
Salix brachycarpa ssp. brachycarpa, 92
23
Salix brachycarpa ssp. niphoclada, 94
24
Salix glauca, Beringia phase (var. glauca) , 98
25
Salix glauca, Western phase (var. acutifolia) , 99
26
Salix glauca, Rocky Mountain phase (var. villosa) , 100
27
Salix athabascensis, 104
28
Salix chamissonis, 110
29
Salix arctophila, 111
30
Salix fuscescens, 116
31
Salix pedicellaris, 117
32
Salix hastata, 121
33
Salix rigida, 124
34
Salix monticola, 127
35
Salix barclayi, 130
36
Salix hookeriana, 135
37
Salix commutata, 139
38
Salix novae-angliae, 141
39
Salix myrtillifolia, 146
40
Salix pyrifolia, 150
List of Maps
41
Salix bebbiana, 151
42
Salix scouleriana, 155
43
Salix planifolia ssp. planifolia, 159
44
Salix planifolia ssp. pulchra var. pulchra, 162
45
Salix planifolia ssp. pulchra var. yukonensis, 163
46
Salix arbusculoides, 172
47
Salix lanata ssp. richardsonii, 173
48
Salix barrattiana, 178
49
Salix candida, 179
50
Salix alaxensis var. alaxensis, 182
51
Salix alaxensis var. longistylis, 183
52
Salix drummondiana, 187
53
Salix sitchensis, 192
List of Tables
1
Chromosome numbers of Salix indigenous to Alaska
and the Yukon, 11
2
Comparison table: Salix polaris, S. rotundifolia, S. stolonifera,
S. phlebophylla and S. arctica, 53
3
Comparison table: Salix rotundifolia ssp. rotundifolia
and ssp. dodgeana, 58
4
Comparison table: Salix rotundifolia, S. phlebophylla
and S. nummularia, 58
5
Comparison table: Salix arctica, S. glauca and S. brachycarpa, 87
6
Comparison table: Salix sphenophylla, S. arctica
and S. ovalifolia, 87
7
Comparison table: Salix brachycarpa ssp. brachycarpa,
ssp. niphoclada and S. glauca, 95
8
Comparison table: Salix athabascensis and S. pedicellaris, 105
9
Comparison table: Salix arctophila and S. arctica, 111
10
Comparison table: Salix pedicellaris and S. myrtilloides, 119
11
Comparison table: Salix hastata and S. barclayi, 121
12
Comparison table: Salix rigida, S. monticola and S. barclayi, 125
13
Comparison table: Salix commutata, S. novae-angliae,
S. myrtillifolia and S. hastata, 140
14
Comparison table: Salix novae-angliae and S. myrtillifolia, 145
15
Comparison table: Salix scouleriana and S. sitchensis, 157
16
Comparison table: Salix planifolia ssp. planifolia
and ssp. pulchra, 167
sw
Comparison table: Salix planifolia ssp. planifolia, ssp. pulchra
and S. phylicifolia, 167
xii
List of Tables
18
Comparison table: Salix Janata ssp. richardsonii, S. barclayi
and S. planifolia ssp. pulchra, 175
19
Comparison table: Salix drummondiana and S. alaxensis var.
longistylis, 188
xiii
Résumé
Le présent examen du genre Salix s’appuie sur des études
effectuées sur le terrain et visant a interpréter les espéces
en fonction des critéres suivants: 1) variations au sein des
populations, 2) variations reliées au développement, 3) varia-
tions provoquées par le milieu, 4) préférences écologiques et
5) modifications résultant de |’hybridation et de |'introgression.
ll se référe aussi aux spécimens d’herbiers et a la documenta-
tion existante.
Trente-neuf espéces aborigénes, deux acclimatées, neuf
sous-espéces et onze variétés composent cette flore. Chaque
espece est décrite en détail et les groupes taxonomiques
intraspécifiques sont comparés a l’espéce elle-méme. Les des-
criptions, qui se veulent le plus complétes possible, expliquent
largement les variations naturelles. L’exposé des problemes
taxonomiques, écologiques et évolutifs se compléte d'un ré-
sumé des caractéristiques permettant l'identification défini-
tive. En outre, l’ouvrage décrit dans ses grandes lignes la
répartition écologique et géographique de chaque groupe
taxonomique et la reporte sur une carte appropriée. Une an-
nexe issue de l’ordinateur 6énumére les spécimens qui font
‘objet des descriptions et des cartes. L’ordinateur permet de
stocker et de récupérer les données pour ensuite imprimer
la liste systématique des spécimens. Voila une innovation
dont on tire € bon compte toute la documentation que com-
portent les travaux de recherche.
L’un des principaux objectifs du présent ouvrage est de
faciliter l'identification des spécimens. Cette opération devient
beaucoup plus simple si l’on connait bien leur répartition
et leur écologie. L’auteur a établi une clé éco-géographique
qui se fonde sur la localisation. Avec la clé traditionnelle
reflétant les traits morphologiques, elle facilitera grandement
les travaux d'identification. De plus, des tableaux comparatifs
mettent en évidence les caractéristiques qui différencient les
espeéces apparentées et exposent les difficultés d’ordre taxo-
nomique qu’elles soulévent.
Les espéces du genre Salix forment un ensemble polyploide
dont le nombre des chromosomes somatiques varie de 38
(diploide) a 224 (dodecaploide). On connait le nombre chro-
mosomien de 33 des 50 espéces qui composent cette flore.
C’est une donnée précieuse permettant d’établir ou de certifier
l'identification dans de nombreux cas. Les tableaux contien-
nent tous les nombres chromosomiens des espéces de I’Alaska
et du Yukon.
L'hybridation naturelle semble beaucoup moins fréquente
qu'on ne l’avait cru. Bien que 12 hybrides interspécifiques et
plusieurs populations introgressives soient notés, |'influence
de lI’hybridation ne semble que relativement importante.
On traite également du réle des espéces du genre Salix
dans la succession écologique des moraines glaciaires du
sud-est de |'Alaska; dans celle des plaines de fusion de la
chaine de |’Alaska; dans celle des alluvions de |’Alaska central
et du gravier des plaines alluviales de |’'Alaska septentrional.
XiV
Summary
The revision of the genus Salix which is presented here is
based on field studies which attempt to understand the species
in terms of the following: 1) population variation, 2) devel-
opmental variation, 3) environmentally induced variation, 4)
ecological preferences, and 5) modification as a result of
hybridization and introgression, as well as on a study of
herbarium specimens and the literature.
Thirty-nine native species, two introduced species, nine sub-
species, and eleven varieties are recognized in the flora. A
detailed description is provided for each species and intra-
specific taxa are compared and contrasted with the species.
The descriptions attempt to describe the species fully and to
account for much of their natural variation. A discussion of
taxonomic, ecological, and evolutionary problems includes a
summary of diagnostic characteristics. In addition, a general
statement of the ecological and geographical distribution of
each taxon is included as well as a distribution map. The speci-
mens on which the maps and descriptions are based are cited
in a computer-generated appendix. The use of computerized
data storage and retrieval methods to prepare this list of
specimens is an innovation which permits the complete docu-
mentation of research materials at a modest cost.
One of the main objectives of the revision is to facilitate
the identification of specimens. Identification can be aided
greatly by a knowledge of the distribution and ecology of the
specimens. An eco-geographical key, based on the geograph-
ical and ecological occurrence of the species, has been
devised. The use of this key along with the conventional key
based on morphological characteristics will permit much easier
identifications than previously has been possible. In addition,
species are compared, in tabular form, with related and similar
species, and taxonomic problems are discussed.
Salix species form a polyploid complex with somatic chromo-
some numbers ranging from 38 (diploid) to 224 (dodecaploid).
Chromosome numbers are known for 33 of the 50 native taxa
in the flora and provide data valuable in making and cor-
roborating taxonomic decisions. All chromosome numbers
known for the Salix of Alaska and the Yukon are tabulated.
Natural hybridization is apparently less common than has
previously been suspected and although 12 interspecific hy-
brids and several introgressive populations are recognized,
the influence of hybridization on species variation is not
considered to be of major importance.
The role that Salix species play in successional develop-
ment on glacial moraine in southeastern Alaska, on glacial
outwash plains in the Alaska Range, on river alluvium in
central Alaska, and on gravel floodplains in arctic Alaska is
discussed. Their success as colonizers is based on their growth
forms and on their reproductive, evolutionary, and ecological
characteristics.
XV
Biographical Note
George William Argus was born and raised in Brooklyn, New
York. In 1949 his interest in the North led him to Alaska where
he studied geology and biology at the University of Alaska. He
received a B.S. degree in 1952. Following two years of military
service in Alaska, he entered graduate school at the University
of Wyoming and later at Harvard University where he was
awarded a Ph.D. degree in biology (plant taxonomy) in 1961.
Field research for his dissertation on the taxonomy of the
Salix glauca-complex in North America took him to Churchill,
Manitoba and to the Gaspé Peninsula of Quebec. After gradua-
tion he held a National Research Council of Canada _ post-
doctorate fellowship at the University of Saskatchewan for
two years.
In 1963 Dr. Argus became a professor in the Departments
of Plant Ecology and Biology and an associate of the Institute
for Northern Studies at the University of Saskatchewan. During
his eight years in Saskatoon he continued his studies on the
taxonomy of Salix and published papers on the willows of
Wyoming and of Wisconsin as well as on the chromosome
numbers and taxonomy of Alaskan and Canadian willows. At
this time he developed an interest in the boreal flora and con-
ducted research into botanical endemism in the sand dune
region of Lake Athabasca. In 1970, after a sojourn as curator
of the herbarium at the University of Oregon, he returned to
Canada where he was engaged in plant systematics research
in the Forest Ecology Institute, Canadian Forestry Service of the
Department of the Environment, Ottawa. He is now Associate
Curator in the Vascular Plant Section of the National Herbarium
(CAN), National Museum of Natural Sciences. In 1971 he
became a naturalized Canadian and now lives in Ottawa with
his family.
xvi
Preface
The genus Salix in the flora of Alaska and the Yukon is one of
the largest, most widespread, and most taxonomically complex
genera. Its antiquity, wide distribution, perennial habit, poly-
ploidy, ability to hybridize, wide ecological amplitude, and
morphological plasticity all contribute to its complexity and
confer upon it great taxonomical, ecological, and evolutionary
interest. However, because of its taxonomic complexities and
uncertainties it is often avoided by taxonomists and ecologists
and thereby valuable information is lost. The taxonomic prob-
lems in Salix are not wholly inherent and many can be traced
to taxonomists who failed to understand the degree and impor-
tance of population variability, or who overemphasized the
occurrence of natural hybridization, or who studied specimens
as individuals rather than as samples of populations. It was
through a desire to contribute to the solution of some of the
taxonomic problems that this study was undertaken.
The only world-wide monograph of Sa/ix was written in 1868
by N. J. Andersson for the A.P. de Candolle Prodromus. Since
that time increased collecting and field study, modern taxon-
omic methods, and new concepts of taxa have made this
work obsolete. More recently, monographs have been written
for the genus in North America (Schneider 1918-1921; Raup
1943, 1959) and in Europe and Asia (Schneider 1917; Skvortsov
1957, 1961, 1966, 1968; Rechinger 1964). Numerous floristic
and monographic works dealing with smaller areas or with
portions of the genus have also been prepared. Over the
years, Salix in Alaska and the Yukon has received considerable
attention from taxonomists and phytogeographers. Coville
(1900, 1901) published two important papers on the willows
of Alaska; Floderus contributed a treatment of the genus to
Hultén, Flora of the Aleutian Islands (1937); Kimura prepared
a treatment for Tatewaki and Kobayashi, A Contribution to the
Flora of the Aleutian Islands (1934). Through the years, Eric
Hultén (1940, 1943, 1967) has paid close attention to the genus,
and his most recent treatment contained in his monumental
Flora of Alaska and Neighboring Territories (1968) has made
a significant contribution to our understanding of the taxonomy
and phytogeography of Sa/ix. Hugh Raup, on the basis of field
studies conducted along the Alaska Highway, on his wide
experience, and on extensive herbarium collections, prepared
a treatment of the willows of boreal western America (1959)
which summarized our understanding of the genus up to that
time. In 1964 Cronquist published a treatment of Salix in
Vascular Plants of the Pacific Northwest, treating a large num-
ber of species which occur in Alaska and contributing a fresh
point of view to their taxonomy. Monographs dealing with
sections or with species complexes have been prepared by
Crovello (1968) for the section Sitchenses and by Argus
(1965a) for the Salix glauca-complex.
However, in spite of this attention, many of the Salix spe-
cies in Alaska and the Yukon are still not well understood.
The present work grew out of a study started in 1966 which
was designed to resolve some of the taxonomic problems as-
Preface
sociated with the dwarf, arctic species. At that time | was also
asked to contribute a treatment of Salix to the revision of
Anderson's Flora of Alaska and Adjacent Parts of Canada be-
ing prepared by S.L. Welsh. Initially the treatment was to be
brief — more a compilation than a revision. However, as spec-
imens were studied and field-work was undertaken it became
evident that a revision would not only be possible but highly
desirable. The primary purposes of this study are: 1) to provide
a workable taxonomic treatment of Salix in Alaska and the
Yukon, and 2) to contribute to the understanding of each taxon,
including its morphology, variation, ecology, and distribution.
More than 3,000 herbarium specimens were studied in this
investigation and over 2,000 specimens were studied and
collected in the field. The collections were studied at the Uni-
versity of Saskatchewan (SASK). Loans of all Sa/ix from Alaska
and the Yukon were obtained from the University of Alaska
(ALA) and lowa State University, including the J.P. Anderson
collection (ISC); selected specimens and types were obtained
from the Plant Research Institute, Ottawa (DAO), the Uni-
versity of Minnesota (MIN) and the Rocky Mountain Herbarium,
University of Wyoming (RM); visits were made to the Arnold
Arboretum (A), the National Herbarium of Canada (CAN), the
Chicago Natural History Museum (F), the Gray Herbarium
(GH), the University of Michigan (MICH), the U.S. National
Arboretum (NA), the New York Botanical Garden (NY), and
the U.S. National Museum (US) to study type specimens, to
obtain range data, and to select loans for special study. All
specimens cited in Appendix B as located in GWA (Argus
personal herbarium) have been deposited in CAN, and repre-
sentative specimens are located at SASK, ALA, US, GH, the
Naturhistoriska Riksmuseum, Stockholm, Sweden, and the Bo-
tanical Garden Herbarium, University of Moscow, U.S.S.R. 1 am
indebted to the curators of these institutions for making avail-
able materials and facilities for study. | am also indebted to
L. Viereck, D.B. Murray, W. Klein, W. Drury, S.L. Welsh, and M.
Williams for placing their recent Alaska and Yukon collections
at my disposal.
My experience with the willows of Alaska dates from 1952
when | collected the alpine flora on nunataks of the Juneau
Icefield. This was followed by trips to the Kennecott region in
1955, and by extensive collecting in central Alaska in 1956
and 1957, in Mount McKinley Park in 1956, and along the Alaska
Highway in 1956 and 1966. In 1966 my assistant Walter Chunys
and | were accompanied by Dr. Yutaka Suda, the cytologist,
to the Arctic Research Laboratory at Barrow, Alaska. There,
through the invaluable logistical support provided by Drs. M.
Brewer and J. Schlindler, we were able to visit Barrow, Meade
River, Cape Beaufort, Nuvagapak Point, and Bullen on the Arctic
slope of Alaska. In 1967 we visited southeastern coastal Alaska,
travelling to Yakutat with the assistance of R. Hurd, U.S. Forest
Service; to Glacier Bay, where we received logistical support
from R. Howe of the U.S. National Park Service; and to Haines,
Juneau, Sitka, Petersburg, Wrangell, and Ketchikan.
Preface
As is usual in studies covering a large geographical area, it
was necessary to rely heavily on the knowledge, experience,
and hospitality of many people. | would particularly like to
acknowledge Les and Teri Viereck, College, Alaska; Keith and
Betty Lou Hart, Juneau; M. Perensovich, Juneau; and C. Jonda,
Glacier Bay; Walter Chunys, whose accomplished outdoor
skills and fine companionship did much to ensure the success
of the field research in 1966 and 1967; Yutaka and Sumi Suda,
who accompanied us to Alaska in 1966; Jo Whitehorn, who
spent many long hours looking up specimen latitude and lon-
gitude to upgrade the herbarium label data and supervising
the keypunching of the summary data cards; and my wife,
Mary, who undertook yeoman service while we were en route
to Alaska in 1966.
The base map is used with the permission of the Canada
Department of Mines and Technical Surveys. Assistance in
mapping data was provided by Eric Argus, Michael Argus, and
M.L. Anderson. The final maps were drafted by the Canada
Department of the Environment, Cartographic Section, under
the supervision of J. Brittain. The computer print-out used in
the Appendices was produced by the Canada Department of
the Environment, Biometrics and Computer Science Branch,
under the supervision of J. Barnabe.
The research was supported by grants from the National
Research Council of Canada (A1955), the Arctic Institute of
North America (ONR-390) under contractual arrangements with
the U.S. Office of Naval Research, and from the Institute for
Northern Studies, University of Saskatchewan.
Taxonomic Relationships
One of the weakest links in the clas-
sification of Salix is the organization
of species into sections. | am not at
all confident that it is possible, with
our present knowledge, to recognize
meaningful sectional groupings, and a
strong argument could be made for
following the method used by Raup
(1959) and simply group related spe-
cies, assigning them informal “group”
names. However, in this study | used
sectional nomenclature, recognizing
the difficulties and pitfalls involved
in this method. | have not attempted
to typify the sectional names, but
rather selected names in common use
by Schneider (1921) and Skvortsov
(1968).
The sections are arranged accord-
ing to a reductional series in stamen
number, starting with section Pentan-
drae having more than two stamens,
followed by the sections with two
stamens and ending with section Sit-
chenses having one stamen. Within
the sections with two stamens, sec-
tion Maccallianae is first, indicating
a presumed relationship with section
Pentandrae, followed by sections Sub-
albae and Longifoliae. The latter sec-
tion is assumed to be among the more
primitive sections because of the oc-
currence of branched aments (Argus
1964). These sections are followed by
those containing dwarf or low-growing
species and their relatives, and then
by sections containing taller, erect
shrubs. This arrangement of sections
cannot be interpreted in phylogenetic
terms — except perhaps for the re-
ductional series in stamen number
— but only as a convenient way to
arrange groups of presumably related
species.
Classification of Salix in Alaska and the Yukon
Section 1. Pentandrae (Borr.) Schneid.
1. S. pentandra L.
2. S. lasiandra Benth.
Section 2. Maccallianae Argus
3. S. maccalliana Rowlee
Section 3. Subalbae Koidz.
4. S. babylonica L.
Section 4. Longifoliae Anderss.
5. S. interior Rowlee
Section 5. Chamaetia Dumort.
6. S. reticulata L.
ssp. reticulata
ssp. glabellicarpa Argus
Section 6. Retusae Kern.
7. S. polaris Wahl.
8. S. rotundifolia Trautv.
ssp. rotundifolia
ssp. dodgeana (Rydb.) Argus
.nummularia Anderss.
. phlebophylla Anderss.
. ovalifolia Trautv.
var. ovalifolia
var. arctolitoralis (Hult.) Argus
var. cyclophylla (Rydb.) Ball
var. glacialis (Anderss.) Argus
12. S. stolonifera Cov.
13. S. setchelliana Ball
NNN
Section 7. Glaucae Pax
14. S. arctica Pall.
15. S. sphenophylla Skvortsov
16. S. brachycarpa Nutt.
ssp. brachycarpa
ssp. niphoclada (Rydb.) Argus
17. S. glauca L.
var. glauca
var. acutifolia (Hook.) Schneid.
var. villosa (Hook.) Anderss.
18. S. athabascensis Raup
Section 8. Myrtosalix Kern.
19. S. chamissonis Anderss.
20. S. arctophila Cock.
Section 9. Myrtilloides Koehne
21. S. fuscescens Anderss.
22. S. pedicellaris Pursh.
Section 10. Hastatae Kern.
23.«S.hastatark:
Section 11. Cordatae Barr. ex Hook.
24. S. rigida Muhl.
25. S. monticola Bebb
26. S. barclayi Anderss.
27. S. hookeriana Barr.
28. S. commutata Bebb
29. S. novae-angliae Anderss.
30. S. myrtillifolia Anderss.
31. S. pyrifolia Anderss.
Section 12. Vetrix Dumort.
32. S. bebbiana Sarg.
33. S. scouleriana Barr.
Section 13. Arbuscella Ser. ex Duby
34. S. planifolia Pursh
ssp. planifolia
ssp. pulchra (Cham.) Argus
var. pulchra
var. yukonensis (Schneid.)
Argus
35. S. arbusculoides Anderss.
Section 14. Lanatae Koehne
36. S. Janata L.
ssp. richardsonii (Hook.)
Skvortsov
37. S. barrattiana Hook.
Section 15. Vi/losae Rouy
38. S. candida Fliigge ex Willd.
39. S. alaxensis (Anderss.) Cov.
var. alaxensis
var. longistylis (Rydb.)
Schneid.
40. S.drummondiana Barr.
Section 16. Sitchenses Bebb
41. S. sitchensis Sanson
Hybridization
The genus Salix is frequently cited as
an example of a taxonomically com-
plex genus in which hybridization is
at the root of many, if not most, of
its taxonomic problems. Evidence
supporting the occurrence of hybrid-
ization in Salix is based on: 1) the
extensive hybridization experiments
conducted by Heribert-Nilsson (1918,
1930); and 2) the occurrence of
polyploidy, presumed to be largely of
the alloploid type.
Heribert-Nilsson’s artificial hybrid-
izations and experiments in species-
making were very extensive and
proved that hybrids are easily formed
between species, particularly those
with the same chromosome number
and within the same section. Chromo-
some homology between species in
the same section was subsequently
shown to be high (Hakansson 1955),
and remarkably good chromosome
pairing was found even in 10- and
12-hybrids. Heribert-Nilsson also
demonstrated the potential for new
stabilized species to arise by the seg-
regation and recombination which
occurs during hybridization, citing
the production of the ‘‘trigena-type”’
from the cross (S. repens x_phylici-
folia) x (S. viminalis x phylicifolia)
and the natural occurrence of S. arc-
togena Flod. in northern Sweden de-
rived from the cross S. herbacea xX
polaris.
In 1918 he stressed the observa-
tion that sometimes the offspring of
crosses did not resemble one or
either of the parents and that con-
clusions concerning the origin of
these “extravagant” forms based on
external morphology alone were like-
ly to be erroneous. He cited as ex-
amples, certain shrubs of the cross
S. viminalis * caprea which morpho-
logically resemble the hybrids S. vi-
minalis x cinerea, S. viminalis x
aurita, S. aurita XX repens and S.
repens X viminalis; specimens of the
hybrid (S. cinerea X purpurea) X
S. caprea that could be mistaken for
a S. phylicifolia hybrid and one plant
of the cross (S. cinerea x purpurea)
x (S. purpurea xX viminalis) which
diverged so widely from the parents
that its origin would be doubtful.
Although Heribert-Nilsson was very
successful in producing artificial
hybrids, he was sceptical about the
permanence of the new forms, be-
lieving that they would be rapidly
exterminated by nature.
The recognition and taxonomic
treatment of natural hybridization
has been the subject of considerable
dispute and is not yet resolved.
Standing at one extreme was the
Swedish salicologist, B. Floderus
(1926), who tended to treat most
arctic and subarctic Sa/ix populations
as large, polymorphic hybrid popula-
tions, restricting species rank to more
or less pure populations. Du Rietz
(1930) noted that there were obvious
dangers in assigning triple and quad-
ruple hybrid names, as Floderus had
done, to complex populations even if
the occurrence of ‘‘exaggerant”’
forms was exaggerated by Heribert-
Nilsson. He suggested as an alter-
native approach that highly poly-
morphic populations could be treated,
without reference to their hybrid
origin, as a single species subdivided
into subspecies, varieties and forms.
He noted, however, that this could re-
sult in the undesirable lumping to-
gether of widely divergent types. A
second alternative, used widely today,
is to hold a less rigid species concept
than the Floderan one and to recog-
nize as hybrids only those forms
clearly intermediate between the
‘pure’ species.
In 1959 Raup argued that since
the recognition of most hybrids was
based on the inspection and com-
Hybridization
MAP 1
® arctica x glauca
() arctica x ovalifolia
& arctica x barclayi
O arctica x stolonifera
Salix Hybrids.
parison of small numbers of spec-
imens, which could not be subjected
to modern studies of introgression
and hybridization, it is more appro-
priate to group such specimens with
the species they most nearly resem-
ble. He writes,
The recognition of hybrids among the willows
by simple comparison of a few specimens, if
carried to its logical conclusion, can lead to
utter taxonomic confusion where this is not
warranted.
In this study | have attempted to
appraise hybridization and introgres-
sion in terms of: 1) discordant varia-
tion in several morphological char-
acteristics, 2) signs of infertility, and
@ brachycarpassp. niphoclada x glauca
O brachycarpa ssp. niphoclada
x glauca (Argus 1965a)
3) the sympatric occurrence of the
putative parents. Where hybrids or
introgressants are recognized | pro-
vide documentation to support such
a conclusion. In the flora of Alaska
and the Yukon the following hybrids
are recognized: Salix arctica x glau-
ca, S. arctica x ovalifolia, S. arctica
x barclayi, S. arctica x stolonifera
and S. brachycarpa ssp. niphoclada X
glauca (Map 1); S. phlebophylla x
rotundifolia, S. athabascensis X pedi-
cellaris, S. barclayi * commutata, S.
barclayi X lanata ssp. richardsonii, S.
barclayi xX stolonifera, S. planifolia
ssp. pulchra xX scouleriana and S.
Hybridization
~~
MAP 2. Salix Hybrids and Intergrades.
@ philebophylla x rotundifolia
© rotundifolia > phlebophylla
a pPhlebophylia x rotundifolia and
rotundifolia > phlebophylla
@ athabascensis x pedicellaris
pedicellaris > athabascensis
alaxensis var. alaxensis < lanata ssp.
richardsonii (Map 2). Four of these
hybrids are based on studies of large
population samples in the field and
in the herbarium (S. phlebophylla x
rotundifolia, S. arctica x stolonifera,
S. athabascensis x _ pedicellaris, S.
barclayi <* stolonifera), whereas the
others are based on isolated spec-
imens.
The following intergrades, which
are apparently the result of hybridi-
zation and introgression, are recog-
nized: S. rotundifolia > phlebophylla
and S. pedicellaris > athabascensis
(Map 2); S. alaxensis var. alaxensis>
0° 135° 130°
& barclayi x commutata
Q barclayi x lanatassp. richardsonii
© barclayi x stolonifera
LJ planifoliassp.pulchra x scouleriana
& alaxensis var. alaxensis x
lanata SSDP. richardsonii
alaxensis var. longistylis (Map 3).
| am unable to recognize most of
the hybrids recorded for Alaska and
the Yukon by Hultén (1968). His
suggestions that S. glacialis (S. ova-
lifolia var. glacialis) = S. arctica xX
ovalifolia, that S. myrtillifolia var.
pseudomyrsinites (S. novae-angliae)
= hybrid forms, that the glabrescent
forms of S. commutata and S. arbus-
culoides are probably of hybrid origin,
and that S. reticulata ssp. glabelli-
carpa is a possible hybrid are unsub-
stantiated and highly speculative. In
my field experience natural hybrid-
ization was encountered infrequently,
Hybridization
170°
MAP 3. Sa/ix Intergrade.
* alaxensis var. alaxensis >var.
longistylis
supporting the view that hybridization
and introgression, although present
in Salix, are not the major causes of
its variability.
Polyploidy
Chromosome numbers based on the
species in the flora of Alaska and
the Yukon reveal a polypoid series
including: 2x, 3x, 4x, 5x, 6x, 8x, 10x
and 12x (Table 1). These numbers
are primarily derived from the basic
number 19, although a parallel series
based on number 22 is known in Salix
and the count reported for S. rigida is
based on this number. The polyploid
series probably represents, for the
most part, alloploidy, a conclusion
supported by the experimental pro-
duction of tetraploid S. cinerea from
crosses between the diploids S. cap-
rea and S. viminalis (Hakansson
1955); but the possible occurrence
of segmental allopolyploidy and auto-
polyploidy cannot be discounted.
Intraspecific variation in chromo-
some number is known, in our flora,
for Salix maccalliana, S. rotundifolia,
S. arctica, S. pedicellaris, S. hastata,
S. sphenophylla and S. scouleriana,
and intrapopulation variation is known
to occur in S. glauca, S. athabascensis
and S. planifolia ssp. planifolia. This
variation in chromosome _ number,
when unaccompanied by significant
morphological differences, may be
the result of autopolyploidy (Hakans-
son 1955). However, this remains to
be proved. The taxonomic importance
of this variation is that it emphasizes
the need for many more chromosome
counts, even for those species that
have already been counted, before
these data can be used with confi-
dence in classification.
Many of the polyploid species, such
as Salix glauca, S. arctica, S. scou-
leriana and S. planifolia are highly
variable morphologically and have
wide distributional ranges. This varia-
bility may be explained in terms of
a rich environmental diversity (see
S. arctica for discussion), or by re-
peated intergenomic exchange result-
ing in new linkage relationships and
new position effects (Mosquin 1966),
or may be due to polyphylesis. It is
very likely that many Salix poly-
ploids have arisen several times
(polychronism) and in several places
(polytopism) in the past, and it is
probable that some are actively evolv-
ing now. The possibility of the poly-
phyletic origin of species is receiving
serious attention in the taxonomic
literature (see Davis and Heywood
1963 and Cain 1944 for review) and
the proof of its occurrence in Salix
will have important taxonomic impli-
cations.
In this study chromosome numbers
have been of assistance in making
certain decisions concerning species
relationships. For example, the rela-
tionship between Salix brachycarpa
ssp. brachycarpa and ssp. niphoclada
has been supported by their common
diploid chromosome number, and their
differentiation from the tetraploid-
hexaploid S. glauca has been affirmed.
Similarly, S. planifolia ssp. planifolia
and ssp. pulchra share a common
tetraploid chromosome number in
contrast to the hexaploid (or tetra-
ploid on basic number 22) S. phylici-
folia. It is the heuristic value of chro-
mosome number data that makes
them so important to the taxonomists,
and our hopes for an improved classi-
fication of Salix depend upon obtain-
ing further chromosomal and cytolog-
ical data.
Polyploidy
Table 1
Species
Sect. Pentandrae
S. lasiandra
Sect. Maccallianae
S. maccalliana
Sect. Longifoliae
S. interior
Sect. Chamaetia
S. reticulata
Sect. Retusae
. polaris
. rotundifolia
. nummularia
. phiebophylla
Nn NAH DH
. ovalifolia
var. ovalifolia
Sect. Glaucae
S. arctica
S. sphenophylla
S. brachycarpa
ssp. brachycarpa
ssp. niphoclada
S. glauca
2n
76
ca. 190, 224
38
ca. 38
76, 95, 114
ca. 76
S. athabascensis ca. 76, 95, 114
Sect. Myrtosalix
S. chamissonis
S. arctophila
76
114
114
76
Geographic
Origin
California(?)
Saskatchewan
Saskatchewan
Alaska
N.W.T.
Manitoba (7?)
U.S.S.R
Scandinavia
Scandinavia
Alaska
U.S.S.R.
U:S:5.R-
Alaska
U.S.S.R.
Alaska
Alaska
N.W.T.
Greenland
Alaska
U.S.S.R.
U:S.S:R:
U.S.S.R.
Manitoba
Saskatchewan
Alaska
Alaska
Alaska
Alberta
Europe
Scandinavia
Scandinavia
Manitoba(?)
Iceland
Manitoba
Yukon
Saskatchewan
U.S.S.R.
Greenland
Chromosome Numbers of Salix Indigenous to Alaska and the Yukon
References
Wilkinson 1944
Suda and Argus 1968
Suda and Argus 1968
Suda and Argus 1969, Johnson and
Packer 1968
Hedberg 1967
Love 1954
Sokolovskaja and Strelkova 1941, 1960,
Zhukova 1967
Holmberg 1931
Holmberg 1931
Johnson and Packer 1968
Zhukova 1968
Zhukova 1969
Suda and Argus 1969
Zhukova 1967
Suda and Argus 1969, Johnson and
Packer 1968
Johnson and Packer 1968
Mosquin and Hayley 1966
Suda and Argus 1968, Holmen 1952
Suda and Argus 1969
Sokolovskaja and Strelkova 1948
Zhukova 1969
Zhukova 1968
Argus 1965a
Suda and Argus 1968
Johnson and Packer 1968
Suda and Argus 1969
Johnson and Packer 1968
Suda and Argus 1968
Love 1954
Holmberg 1931, L6ve and Léve 1948
Wilkinson 1944
Léve 1954 (as cordifolia s.1.)
Love and Love 1956 (as callicarpaea)
Argus 1965a (as glauca)
Suda and Argus 1969
Suda and Argus 1969
Zhukova 1968
Jorgensen, Sorensen, and
Westergaard 1958
11
Polyploidy
Species
Sect. Myrtilloides
S. fuscescens
S. pedicellaris
Sect. Hastatae
S. hastata
Sect. Cordatae
S. rigida
S. monticola
S. myrtillifolia
Sect. Vetrix
S. bebbiana
S. scouleriana
Sect. Arbuscella
S. planifolia
ssp. planifolia
ssp. pulchra
S. arbusculoides
Sect. Lanatae
S. lanata
ssp. richardsonii
Sect. Villosae
S. candida
S. alaxensis
var. alaxensis
S. drummondiana
Sect. Sitchenses
S. sitchensis
152
Geographic
Origin
Alaska
U.S.S.R.
Manitoba (?)
Yukon
Manitoba
Scandinavia
U.S.S.R.
U.S.S.R.
?
Saskatchewan
Saskatchewan
Manitoba (?)
Saskatchewan
B.C.
Yukon
Saskatchewan
New Hampshire
Manitoba (7?)
Alaska
Alaska
U:S.S.R:
Alaska
Saskatchewan
U.S.S.R.
Saskatchewan
Alaska
U.S.S.R.
Saskatchewan
B.C.
References
Suda and Argus 1969, Johnson and
Packer 1968
Zhukova 1967
Love 1954
Suda and Argus 1969
L6ve and Ritchie 1966
Holmberg 1931
Zhukova 1967
Sokolovskaja and Strelkova 1960
Wilkinson 1944 (as cordata)
Suda and Argus 1968
Suda and Argus 1968
Love 1954
Suda and Argus 1968
Taylor and Mulligan 1968
Suda and Argus 1969
Suda and Argus 1968
Love and Léve 1964
Love 1954
Johnson and Packer 1968
Suda and Argus 1969
Zhukova 1967, 1968, 1969
Suda and Argus 1969
Suda and Argus 1968
Zhukova 1969
Suda and Argus 1968
Suda and Argus 1969, Johnson and
Packer 1968
Zhukova 1967, 1969
Suda and Argus 1968 (as subcoerulea)
Taylor and Mulligan 1968
Ecology
In Alaska and the Yukon Salix species
occupy a wide variety of habitats
from forests to tundra and from
wet bogs and marches to dry sandy
forests, talus slopes and rock out-
crops. These habitats are extremely
diverse in terms of moisture and
nutrient regimen and in species com-
position, but they have one important
common attribute, namely, they are
subject to change. This change may
take the form of physical disturbance
or of a labile successional stage. The
relationship of Sa/ix to habitat insta-
bility has long been known and wil-
lows are characteristically associated
with seasonally disturbed river flood-
plains, gravel outwash plains, glacial
moraines, fire disturbance, frost dis-
turbed arctic and alpine tundra, and
sites of human disturbance. They are
also known to be components of
labile successional stages often ap-
pearing in the earliest stages and
becoming less and less important as
vegetational succession leads to a
relatively stable, mature vegetation.
An example of the role Salix plays
in successional development on gla-
cial moraine in southeastern Alaska,
on glacial outwash in the Alaska
Range, and on river alluvium in cen-
tral and northern Alaska will illus-
trate this generalization.
Succession on Glacial Moraine
This account of the successional
development on glacial moraine in
southeastern Alaska is based on ob-
servations made at Muir Inlet, Glacier
Bay National Monument in 1967, and
on work done by Decker (1966).
Stage 1. Recently uncovered glacial
moraine near the ice front. No vegeta-
tional development except for seed-
lings of Epilobium J/atifolium, Dryas
drummondii, Salix reticulata, S. stolon-
ifera, S. arctica, S. commutata, S. ala-
xensis, S. barclayi and S. sitchensis.
Stage 2. Glacial moraine vegetated
by separate, often circular, Dryas
mats interspersed with lichens and
the willows Salix stolonifera, S. reti-
culata (in lichen mats), S. arctica
(usually in Dryas mats), S. commu-
tata, S. alaxensis, S. barclayi, and
S. sitchensis.
Stage 3. Dryas forms an almost
continuous mat interspersed with
shrubs of Alnus incana and Salix
commutata, S. alaxensis, S. barclayi
and S. sitchensis. The dwarf species,
Salix arctica, often occurs under
clumps of A/nus and under taller
Salix.
Stage 4. A young Al/nus-Salix thick-
et about 3 m tall. The same Salix
species are present as in Stage 3
but S. arctica and S. commutata are
less frequent.
Stage 5. A dense Aj/nus_ thicket
with saplings of Picea and Populus.
The only Salix present are S. barclayi
and S. sitchensis.
Stage 6. In this stage succession
has led to a young Picea sitchensis
forest in which Salix sitchensis is the
last surviving species of willow. As
this forest type matures even Salix
sitchensis is eliminated except for its
occurrence in forest openings and
along stream margins.
In general all the willow species
in this example occur in the pioneer
stages of vegetational development.
As taller and taller shrubs invade the
area, the dwarf willows are the first to
be eliminated with only Salix arctica
able to survive even under tall alder
(Stage 4). The next to be eliminated
is the low shrubby species, S. com-
mutata, followed by S. a/axensis and
S. barclayi and finally by S. sitchensis.
The latter species is the only willow
that occurs in this area from the fresh
moraine up to the young Picea forest
stage. In the early stages of vegeta-
tional succession, Salix sitchensis
Ecology
may be a prostrate or a very low-
growing shrub, later producing erect
branches and finally reaching a height
of about 4 m tall. In the mature Picea
sitchensis-Tsuga heterophylla forests
few shrubs of any kind occur. How-
ever, the Salix that do occur there
are always associated with forest
openings, water courses and human
disturbance.
The characteristics of these spe-
cies that permit their rapid establish-
ment on glacial moraine include the
production of abundant, wind-borne
seeds and rapid seed germination
(Argus 1965a). Their elimination from
the later successional stages is prob-
ably related, at least in part, to their
growth habit. The dwarf species are
eliminated as soon as they are over-
topped by taller shrubs which shade
them and which lead to a more rapid
aggradation of the soil surface due to
the accumulation of organic debris.
The smaller shrubs are next shaded
out and the longest surviving species,
such as Salix sitchensis, are those
which are able to remain in the
canopy for the longest time.
Succession on Glacial Outwash
Viereck (1966) describes the vegeta-
tional succession on gravel outwash
in Mount McKinley National Park,
Alaska.
1. Pioneer Stage. Outwash plain
vegetated by mats of Dryas drum-
mondii and D. integrifolia and the
willows, Salix alaxensis var. alaxen-
sis, S. myrtillifolia, S. brachycarpa
ssp. niphoclada, S. glauca and S. set-
chelliana. On the basis of my ex-
perience elsewhere, | would suspect
that in this stage seedlings of all the
Salix species that are to follow would
be present; but the identification of
willow seedlings is difficult and they
may be easily missed.
2. Meadow Stage. E/ymus-Festuca-
Poa meadow. The two low-growing
species, S. myrtillifolia and S. setchel-
liana, present in stage 1 are absent
and several taller species, including
S. lanata ssp. richardsonii, S. plani-
folia ssp. pulchra and S. barrattiana
are present for the first time. The
dwarf species, S. reticulata, is also
reported in this stage but its occur-
rence is erratic and its frequency is
low.
3. Early Shrub Stage. In the early
Betula glandulosa shrub stage, two
additional tall Sa/ix enter the picture,
S. arbusculoides and S. barclayi.
4. Late Shrub Stage. In general,
all the Salix in stages 2 and 3 persist
through the late shrub stage although,
as the Betula glandulosa thicket be-
comes more dense, several of these
species decline in importance.
5. Climax Tundra. In the Eriopho-
rum-Betula glandulosa tundra the
only willow present is Salix glauca;
however, this species is now erratic
and infrequent.
Succession on River Alluvium in
Central Alaska
In central Alaska, the role of Salix
in succession on silty-sandy alluvial
deposits follows a similar pattern of
invasion and dominance in the early
successional stages, followed by de-
cline and elimination. Pioneer vegeta-
tion on silty-sandy bars is dominated
by S. alaxensis in association with S.
arbusculoides, S. lasiandra and S.
planifolia ssp. pulchra var. yukonen-
sis. As the site becomes more stabi-
lized and the thicket increases in
height and density, S. /asiandra and
S. planifolia are eliminated. Salix ala-
xensis and S. arbusculoides may dom-
inate such a site for a considerable
period of time, reaching a height of 7-
9 m and obtaining a circumference
near the base of 5-10 dm. Populus
balsamifera and Picea glauca then in-
Ecology
vade the site, eventually dominating
it and eliminating the willows. In ma-
ture Picea glauca forests, Salix beb-
biana may occur in forest openings
produced by fallen trees or snow
breakage but the original, pioneer
Salix are absent.
Succession on River Alluvium in
Northern Alaska
Bliss and Cantlon (1957) studied the
vegetational succession on the Col-
ville River at Umiat, Alaska, recogniz-
ing four plant communities.
1. Perennial Herb Community. Suc-
cession is initiated on pockets of
sand and silt on the bare floodplain
gravels. The pioneer species include
the herbs, Crepis nana, Erigeron pur-
puratus, Epilobium latifolium, et al.,
and the willows, Salix alaxensis var.
alaxensis, S. arbusculoides and S.
glauca.
2. Young Felt-Leaf Willow Com-
munity. On the next terrace vigorous
stands of Salix alaxensis develop, in
association with S. glauca, S. hastata,
S. arbusculoides and S. lanata ssp.
richardsonii.
3. Decadent Felt-Leaf Willow Com-
munity. In this community Salix ala-
xensis is still the dominant species
but there are many dead stems pres-
ent. There is an increase in the im-
portance of S. arbusculoides and S.
glauca; S. hastata is absent and S.
brachycarpa ssp. niphoclada is pres-
ent for the first time.
4. Terrace Communities. The vege-
tation on the higher terraces is com-
posed of a mixture of shrubby, dwarf
heath meadow and marsh types. Salix
planifolia ssp. pulchra is the domi-
nant shrub and is associated with S.
lanata ssp. richardsonii, S. glauca and
S. arbusculoides. The willows S. ala-
xensis, S. hastata and S. brachycarpa
ssp. niphoclada are absent in this rel-
atively undisturbed vegetation type.
Succession on Arctic Floodplain
On the arctic coast of Alaska at Cape
Beaufort, a transect from the sandy-
gravel floodplain of a creek to the
upper terrace reveals a change in the
Salix species composition (Argus, un-
published).
1. Floodplain. On the floodplain of
the creek, frequently disturbed by
flooding and the resulting erosion and
siltation, Salix alaxensis var. alaxen-
sis is the dominant species in thickets
1-2 m tall and is associated with
occasional low shrubs of S. hastata.
2. First Terrace. At the level of the
first terrace an organic turf develops
over the gravel, and the vegetation
consists of an open Salix alaxensis
thicket with the shrubby S. /anata
ssp. richardsonii and the dwarf wil-
lows, S. rotundifolia, S. reticulata
and, rarely, S. arctica. The dwarf wil-
lows grow in the turf within openings
in the Salix thicket.
3. Second Terrace. The drier second
terrace is dominated by a Salix
glauca, S. planifolia ssp. pulchra
thicket, 3-6 dm tall, with the dwarf
S. phlebophylla and S. reticulata in
the openings.
4. Tussock Meadow. An _ Eriopho-
rum, Arctagrostis tussock meadow
develops on the upper terrace. The
dominant shrub is Salix planifolia ssp.
pulchra, and S. fuscescens grows in
the wet depressions between the tus-
socks.
In the Cape Beaufort area the taller
Salix occupy the open floodplains
where drainage is better and where
the depth to permafrost is greater.
The dwarf species occur in the rela-
tively stable vegetation in which dis-
turbance and siltation are minimal
and where drainage is still good. In
the poorly drained vegetation only
Salix planifolia ssp. pulchra is com-
mon and Salix fuscescens is restrict-
ed to the wettest habitats.
Ecology
Colonizing Attributes of Salix
Ehrendorfer (1965) developed three
models of successful colonizers based
on habit and on reproductive, evolu-
tionary, and ecological factors. The
habitat requirements of Salix corre-
spond very closely to those of Ehren-
dorfer's type 1 colonizer, the peren-
nial polyploid type which occurs in
“labile successional stages or open
facies of otherwise more or less clos-
ed associations” or in secondary com-
munities opened by man. A compari-
son of Salix with the characteristics
of Ehrendorfer’s type 1 colonizer may
help explain its great success as a
colonizer and its rapid geographical
expansion. In the following discus-
sion | will cite the factor to be con-
sidered, followed by the general
characteristics exhibited by Ehren-
dorfer's type 1 colonizer in brackets
and a discussion of the factor as it
occurs in Salix.
1. Predominant form of fertiliza-
tion (type 1, allogamy). Salix is dioe-
cious and therefore obligately out-
crossed.
2. Differentiation of seed or fruit
structure and dispersal (type 1, weak,
conservative). The seeds of Salix are
specialized for wind dispersal by
their light weight and the presence of
an arillate comae of fine trichomes
(Argus 1965a).
3. Vegetative reproduction (type 1,
often present). Most Salix propagate
readily from cuttings, and the dis-
persal of branch fragments by water
is common. Some species, such as the
North American S. hookeriana, S. sit-
chensis and S. lasiandra, and the Eu-
ropean S. fragilis, are characterized
by highly brittle branchlets, an at-
tribute which confers an adaptive ad-
vantage, facilitating dispersal, to spe-
cies occupying riverine habitats. Salix
interior, a very aggressive colonizer
of sandy and silty river bars and is-
lands, reproduces by shoot buds on
roots, enabling it to spread rapidly
and to form large colonies. This
mode of vegetative reproduction is
uncommon in Salix and is’ only
known in section Longjfoliae.
4. Chromosome number (type 1,
often polyploid). About 40 per cent
of Salix species are polypoid (Suda
and Argus 1968).
5. Chromosome structural differen-
tiation (type 1, weak). In general,
the structural variation in Salix chro-
mosomes is small. Wilkinson (1944),
in describing the morphology of chro-
mosomes in Salix, notes that the typ-
ical pattern consists of four long
chromosomes with secondary con-
striction, four satellited chromo-
somes (two of these with median
constrictions and two either con-
stricted or unconstricted) and gener-
ally four to eight unconstricted chro-
mosomes in a short length class.
6. Chromosome centromere posi-
tion (type 1, relatively symmetrical).
The majority of the somatic chromo-
somes in Salix have a median centro-
mere (Wilkinson 1944).
7. Population isolation (type 1, pri-
marily geographical and ecological).
Salix species are differentiated ecolo-
gically and geographically and such
isolation probably plays an important
role in speciation. However, many Sa-
lix communities contain several spe-
cies, and hybrids are not as abundant
as they theoretically could be (Heri-
bert-Nilsson 1918, 1930), suggesting
that other isolating mechanisms, such
as differences in flowering time (Ar-
gus 1965a), are also operative.
8. Population differentiation (type
1, mostly allopatric). The geographi-
cal differentiation within Salix sug-
gests that this pattern is also dis-
played by Salix.
9. Hybridization (type 1, in contact
areas, often extensive). Hybridization
Ecology
between Salix species is theoretically
possible (see Hybridization, p. 6)
and occurs in some areas. However,
of greater importance is the hybridiza-
tion between incompletely differenti-
ated geographical populations which
were isolated by Pleistocene glacia-
tions and have in _ post-Pleistocene
time subsequently re-established their
ranges. The great population variation
in Salix glauca in the central Cana-
dian Arctic and Subarctic is thought
to be due to this process (Argus
1965a).
10. Population variability (type 1,
populations relatively variable). In
Salix population variation is relatively
high (cf. S. arctica, S. glauca, S. ova-
lifolia and S. scouleriana), as would
be expected in an outcrossing taxon
occurring in habitats which exert rel-
atively low selection pressures (Baker
1959). However, at the same time,
polyploidy, acting as a buffer against
genetic change (Stebbins 1950, and
Mosquin 1966), and vegetative re-
production may lead to a reduction
in variability.
The comparison of Salix with
Ehrendorfer’s type 1 colonizer leads
to the conclusion that the general
success of Salix species as colonizers
is due to an interrelated complex of
ecological, reproductive and evolu-
tionary characteristics. A comparative
study of Salix species in relation to
these characteristics could lead to
hypotheses explaining the narrow
ecological amplitude of some species
and the limited geographical distri-
bution of others.
Eco-geographical Key
A knowledge of the distribution and
ecology of unknown specimens of Sa-
lix can be very useful in arriving at
their identification. The following key
is an attempt to organize the Salix
species of Alaska and the Yukon in
relation to these attributes. The ar-
rangement of the species is based
largely on my field experience and
on the herbarium specimens that |
have studied. Further exploration and
study are sure to result in some modi-
fication of this key, but its major out-
line is unlikely to be changed.
|. Forested Regions
A. Central Alaska and Southern Yukon
1. Treed vegetation
a.
b.
Upland forests
bebbiana, scouleriana, planifolia ssp. planifolia, arbusculoides
Floodplain forests
monticola, bebbiana, arbusculoides, alaxensis
. Picea mariana muskegs
maccalliana, glauca var. acutifolia, fuscescens, pedicellaris, monti-
cola, novae-angliae, myrtillifolia, bebbiana, scouleriana, planifolia ssp.
planifolia, ssp. pulchra var. yukonensis, arbusculoides
2. Treeless vegetation
a.
Fens
maccalliana, brachycarpa ssp. brachycarpa, athabascensis, pedicella-
ris, myrtillifolia, planifolia ssp. planifolia, ssp. pulchra, candida
. Floodplains
lasiandra, maccalliana, interior, brachycarpa ssp. brachycarpa, ssp.
niphoclada, glauca var. acutifolia, var. villosa, rigida, novae-angliae,
planifolia ssp. pulchra var. yukonensis, lanata ssp. richardsonii, ala-
xensis, drummondiana
B. Southern Coastal Alaska
1. Treed vegetation
a.
b.
Forest openings
barclayi, scouleriana, sitchensis
Floodplains and glacial outwash plains
lasiandra, barclayi, commutata, alaxensis, sitchensis
2. Treeless vegetation
a.
b.
Sand dunes
hookeriana, alaxensis var. alaxensis
Fens
lasiandra, barclayi, commutata, hookeriana, scouleriana
Il. Alpine and Subalpine Regions
A. Yukon-Tanana Hills, Alaska Range, Coast Ranges and Northern Rocky
Mountains
1. Alpine vegetation
a.
Tundra
reticulata ssp. reticulata, ssp. glabellicarpa, rotundifolia ssp. rotundi-
folia, ssp. dodgeana, phlebophylla, stolonifera, arctica, chamissonis,
fuscescens, hastata, arbusculoides
Eco-geographical Key
b. Snowbeds
reticulata, polaris
2. Shrubby subalpine vegetation
brachycarpa, glauca var. acutifolia, var. villosa, hastata, barclayi, com-
mutata, bebbiana, planifolia ssp. pulchra, lanata ssp. richardsonii,
barrattiana, drummondiana
3. Floodplain vegetation
setchelliana, brachycarpa, hastata, monticola, commutata, bebbiana,
scouleriana, planifolia ssp. pulchra, arbusculoides, lanata ssp. richard-
sonii, alaxensis
B. Brooks Range and Northern Mackenzie Mountains
1. Alpine vegetation
a. Tundra
reticulata, rotundifolia, phlebophylla, arctica, chamissonis, arctophila,
fuscescens, planifolia ssp. pulchra, arbusculoides
b. Snowbeds
reticulata, polaris
2. Subalpine vegetation
brachycarpa ssp. niphoclada, glauca var. glauca, planifolia ssp. pul-
chra, lanata ssp. richardsonii, barrattiana
3. Floodplain vegetation
brachycarpa ssp. niphoclada, glauca var. glauca, var. acutifolia, has-
tata, pyrifolia, alaxensis
ll. Arctic Regions
1. Northern Alaska and Yukon, Western Alaska
a. Herbaceous tundra
reticulata, polaris, rotundifolia ssp. rotundifolia, ssp. dodgeana, phle-
bophylla, arctica, sphenophylla, chamissonis, arctophila, fuscescens
b. Shrubby tundra
glauca var. glauca, planifolia ssp. pulchra, arbusculoides, lanata ssp.
richardsonii, barrattiana
c. Fell-fields and talus slopes
reticulata, phlebophylla, arctica
d. Coastal beaches and marshes
ovalifolia var. ovalifolia, var. arctolitoralis, var. glacialis, arctica
e. Floodplains and sand dunes
brachycarpa ssp. niphoclada, glauca var. glauca, hastata, arbuscu-
loides, alaxensis var. alaxensis, barrattiana
2. Bering Sea Islands
reticulata, polaris, rotundifolia ssp. rotundifolia, ssp. dodgeana, num-
mularia, phlebophylla, ovalifolia var. ovalifolia, var. cyclophylla, arc-
tica, sphenophylla, chamissonis, fuscescens, planifolia ssp. pulchra,
alaxensis var. alaxensis
3. Aleutian Islands
reticulata, rotundifolia ssp. rotundifolia, ovalifolia var. ovalifolia, var.
cyclophylla, arctica, barclayi, commutata
Method of Study
This study began by an _ examina-
tion of herbarium specimens and the
literature in order to obtain a pre-
liminary understanding of the units
(usually corresponding to species or
intraspecific taxa) occurring in Alaska
and the Yukon. These units were then
studied as natural populations in the
field during 1966 and 1967 at which
time 36 of the 41 species in the flora
were encountered. In the field an
effort was made to gain an appre-
ciation of the following: 1) population
variation, 2) developmental variation,
3) environmentally induced variation,
4) ecological preferences, and 5) the
occurrence of hybridization or intro-
gression. This information, along with
chromosome numbers and geograph-
ical distribution, was then used to
delimit the taxa. Finally, the appro-
priate nomenclature was determined
through a study of original descrip-
tions and type specimens.
Descriptions
A detailed description of each taxon
based on specimens from Alaska and
the Yukon was compiled. In some taxa
(Salix nummularia, S. sphenophylla,
S. maccalliana, S. pyrifolia and S.
rigida) there were insufficient speci-
mens from Alaska and the Yukon, and
therefore extraterritorial specimens
were used. Descriptive data were ob-
tained from a number of specimens
and generally the more variable the
characteristic, the more specimens
that were examined. For example, in
S. planifolia ssp. pulchra the relative-
ly invariable bract length and anther
length were based on five to eight
observations, but leaf length and
width, petiole length and_ stipule
length were based on 71 observa-
tions. Every effort was made to
characterize the mean and the ex-
tremes of variation of each character-
istic; for example, leaf length in S.
20
planifolia ssp. pulchra is given as
(2.2) 3.2-6 (7.5) cm, indicating that
the general variation in leaf length
is between 3.2 and 6 cm, but ex-
tremes of 2.2 and 7.5 cm also occur.
Measurements were made using a
millimetre scale or an ocular micro-
meter in a dissecting microscope at
22X.
Terminology
The terminology used for simple, sym-
metrical, plane shapes follows the
system proposed by the Systematics
Association Committee for Descrip-
tive Biological Terminology (1962).
Other terminology generally follows
W.T. Stearn, Botanical Latin, 1966.
In the case of certain commonly
used terms for indumentum types,
the following standards were set up
within Salix.
Sericeous: the type of indumentum
on the lower side of the leaves of
S. arbusculoides.
Villous: the type of leaf indumen-
tum in S. commutata.
Densely white lanate: the type of
indumentum on the underside of
leaves of S. alaxensis var. alaxensis.
Sericeous-lanate: the type of indu-
mentum on the lower surface of the
leaves of S. barrattiana.
Villous-woolly: the type of indu-
mentum on the young branchlets of
S. alaxensis var. alaxensis.
Some morphological terminology
conventionally used in Salix is evi-
dently incorrect and several new
terms have been used in this paper:
Stipe: the pedicel of authors. If
the nectary represents a_ reduced
perianth as was suggested by Fisher
(1928), then this structure cannot
be a pedicel, which is the stalk of
a flower, but rather a stipe or gyno-
phore.
Floriferous branchlet: the peduncle
of authors. In 1965 | proposed using
Method of Study
GENUS tsi
TAXON
00000/00 ARG DON OOOO OOO ODOC OOO000HDHOODDONODO0NDD 010 000000000000000
1 2.3 4 5/6 7 8/39 ;10/ 99 12 13/14 15]96 17 18)19 20 21 22/23 24 25 25 27:28 29 30 3) 32 33 34 35 36 37 38 39 40 41
SEs OB GAN ON Shy hon Coca
eS AA hs AA Die AE ital tl
Beer ere Lib bee
000000/0000000000
42 43 44 45 46 47 48145 50 51 52 S354 55 96 57 58 59 60 6 62 63 64/65 66 G7 68 69 70) 71 72 73 74 7S 76 77 78 19 BO
REE REE ERERERESRRSEREE SERRE RRR R EERE
2222222
3333333
22 22222222222222)222222)2222222222
as a aah cin a Sued ada
SOE aa een econ Cee CROnR ES CCD ceca CCE coe RCC
sssssssssbassspsspssspssssssssssssssssssssossuspsssssssossanssssssssgsssssssss
sasesisasisesesesspess
|
i
RESTO SUMO E SERS CHR SRO SLES
$99 99,99 9\919|9 9 5/9 919991999 919999 9\999999999993
R234 sls 7? len EE GAS MAL CRS SEE ES
Fig. 1 Summary data card
Columns __— Data
1-5 Genus code number
6-8 Species code number
9 Subspecies code number
10 Variety code number
11-13 Hybrid code; code number
of second parent
14-15 Province/State code number
16-18 Quadrangle code number;
1:250,000 map sheet
19-22 Latitude
23-27 Longitude
28-41 Locality place name
42-48 Date
49-64 Collector’s name;
first collector only
65-70 Collection number
71-74 Herbarium acronym
75-80 Herbarium accession number
the term reproductive branchlet for
the leafy branchlet terminating in an
ament. However, this term is not
completely appropriate and | have
changed it to floriferous branchlet,
that is, a branchlet bearing flowers.
The length of the floriferous branch-
let is measured from the proximal
end of the branchlet to the lowermost
(often sterile) bract. A true peduncle
does occur (Argus 1965a) but its
limits are often doubtful and this
term is not used here.
Bract: the scale of authors. Usually
this term is applied to the foliar struc-
tures on the floriferous branchlet, but
| see no reason not to refer to them
BEG EEC EECEGK EC EE HGF CECH GK OG GG BC EES EG E6G EEE G GEG E6G HG EGE EGEGES
Pan IDA TIAMAT
SASe eee ee ees ieee ee eae ee ow
SSO EEL CLE LEU ELLE
Authority
Dalla Torre and Harms 1900-1907
Arbitrary
Arbitrary
Arbitrary
Arbitrary
Arbitrary
Orth 1967 (Alaska), Arbitrary
(Canada)
Lanjouw and Stafleau 1964
as leaves, and | use the term bract
for the foliar structure subtending
each flower.
Specimen Citations
Herbarium label data were recorded
on 3 x 5 inch index cards as the
research proceeded. After these data
were supplemented by geographical
and taxonomic code numbers and,
where necessary, by latitude and lon-
gitude, the data were key-punched on
IBM-type summary data cards (Fig.
1). The cards were patterned after
those used by Soper (1964) to ma-
chine map species distribution. Sum-
mary data cards were produced for
21
Method of Study
the Argus & Chunys 1967 collections
as a by-product of a computerized
herbarium labelling system. The sum-
mary data cards were machine sorted
in terms of taxa and geography, and
a print-out of the information on these
cards is included as Appendix B.
Appendix A contains a list of code
numbers, acronyms, and _ abbrevia-
tions used in Appendix B and in the
text.
The use of computerized data stor-
age and retrieval methods to prepare
a list of specimen citations is an in-
novation which permits the complete
documentation of research materials
at a modest cost.
Nomenclature
No attempt was made to cite com-
plete synonymy for each taxon, and
names used in reference to the spe-
cies in northwestern North America
are emphasized. Wherever possible
type material was studied and all the
types cited have been examined. In
cases where type specimens were
not seen, the evaluation of names
was based on the original descrip-
tions and on common usage.
Distribution Maps
The specimens examined in this study
(listed in Appendix B) were hand-
plotted on outline maps. The map
projection is Lambert azimuthal equal
area. On the species maps dots rep-
resent specimens and circles repre-
sent literature reports or doubtful
identifications. The distribution maps
of Alaska and the Yukon are relatively
comprehensive. However, all possible
sources of collections from the west-
ern Northwest Territories and north-
western British Columbia were not
consulted and the maps in these
regions are therefore less complete.
22
Systematic Treatment
23
Salicaceae
SALIX L. Sp. Pl. 1015. 1753.
Description of genus
Plants dioecious; dwarf or trailing arctic-alpine shrubs, erect shrubs, or
occasionally trees; buds covered with a single scale; leaves alternate, simple
and usually stipulate, the margins entire or variously toothed; flowers uni-
sexual, borne in spike-like aments which are either sessile or are borne on
floriferous branchlets (short shoots) on the branches of the previous year;
each flower subtended by an entire bract (scale), and by one to several
nectaries (glands); staminate flowers usually consist of two stamens, rarely
one, or three to five, the filaments distinct or partly united; pistillate flowers
consist of a single pistil which may be sessile or borne on a stipe (pedicel),
the single style sometimes bifurcate, the two stigmas usually bifurcate or
horseshoe-shaped; fruit a two-valved capsule with several seeds surrounded
by a coma of fine hairs.
24
Key to the Species
1. Dwarf or prostrate, trailing shrubs under 2 dm tall.
2. Pistillate and staminate flowers with two nec-
taries, one on either side of stipe; leaves prom-
inently reticulate and pale beneath; aments borne
on prominent, subterminal floriferous branchlets.
3. Pistils densely sericeous; stipes sericeous;
bract apex rounded or retuSe ...............cccceeesseeeeeees
3. Pistils glabrous and glaucous or with patchy
pubescence; stipes glabrous; bract apex re-
tuse
POPPE HEHEHE THH HT OHHH EE EH THEE OHHH EEE ET EEHEH FES OHHHHEEE EHO HEE EE EEE HEHEHE
2. Pistillate flowers with one nectary between the
stipe and the ament axis; aments borne on lateral,
floriferous branchlets.
4. Pistils pubescent, sometimes only on beak.
5. Leaves distinctly and minutely serrulate ..........
5. Leaves entire or toothed only on the lower
half.
6. Leaves green or pale green beneath, non-
glaucous.
7. Branches clothed with persistent, skel-
etonized leaves; leaf margins usually
ciliate; nectaries usually shorter than the
SRM So wn Picacho tact dad ck ae, - GES Mee eeeo s N
. Branches without persistent, skeleton-
ized leaves; leaf margins rarely ciliate;
nectaries 2-5 times as long as_ the
iT | 1) = peal a eal eer cence. eldadal > Seana ae
6. Leaves glaucous beneath.
8.
Leaves 0.9-1.5 cm long, margins promi-
nently ciliate; aments globose ..................
Leaves longer than 1.5 cm, margins not
ciliate; aments cylindrical.
S. reticulata ssp.
reticulata, p. 45
S. reticulata ssp.
glabellicarpa, p. 47
S. chamissonis,
BS. 107
S. phlebophylla,
p. 61
S. polaris, p. 49
S. ovalifolia var.
glacialis, p. 68
25
Salix L.
9. Styles 0.1-0.5 mm long.
10. Leaves obovate to elliptic 1.7-2.7
times as long as wide, glabrous
above, margins distinctly toothed on
lower half; pistils sparsely pubes-
cent with ferruginous trichomes ...... S. fuscescens,
o- TT3
10. Leaves narrowly elliptic to narrowly
obovate, 2-3.7 times as long as wide,
pubescent on both sides, margins
entire; pistils densely pubescent
with white trichomes. ...................00. S. brachycarpa ssp.
niphoclada, p. 91
9. Styles longer than 0.5 mm.
11. Pistils sparsely pubescent with
crinkly, refractive trichomes, nec-
taries shorter than the stipes,
branchlets slender and trailing, gla-
brous; leaves glabrous ..................0. S. arctophila,
p. 109
11. Pistils sparsely or densely pubes-
cent with non-refractive trichomes;
nectaries equal to or longer than
the stipes; branchlets and leaves
various.
12. Pistils densely pubescent; leaves
dark green and usually glossy
above, cuneate to rounded at
base; branchlets trailing to erect. S. arctica, p. 81
12. Pistils glabrous or sparsely pu-
bescent on beak; branchlets trail-
ing and rooting.
13. Leaves dull above, base usually
cuneate, sometimes acute ........ S. sphenophylla,
p. 85
13. Leaves glossy above, base
acute to rounded or subcordate_ S&S. stolonifera,
p. 72
26
Key to the Species
4. Pistils glabrous.
14. Leaves green (non-glaucous) beneath.
15.
15.
Decumbent or trailing forest shrubs, leaves
narrowly elliptic to narrowly obovate, 2-5
cm long, margins crenate to crenate-
peeratiateane.20. us. URIS, A6. 2 Baines
Dwarf, sometimes trailing, arctic shrubs;
leaves circular to obovate or narrowly el-
liptic, 0.5-1.4 cm long, margins entire or
toothed only at base.
16. Leaves subcircular, prominently reticu-
late, margins glandular toothed on lower
half, non-ciliate; branchlets more or less
trailing; styles 0.2-0.4 mm long ................
16. Leaves circular or sometimes narrowly
elliptic, not prominently reticulate, mar-
gins entire and ciliate; branchlets erect,
not trailing; styles 0.5-1 mm long.
17. Pistillate aments 4-12 (15)-flowered;
leaves 5-10-14 mm long, 0.9-1.3 times
as long as wide; petioles 1.4-2-3 mm
ONG so... 82etlle...cavede-.csmeems..4
17. Pistillate aments 2-4 (9)-flowered;
leaves 4-6 mm long, 1.7-2.7 times as
long as wide; petioles 0.8-1.6 mm
LONG: 2.8). :1.ebee Nis... ebscawen.: eatvan.t:..3
14. Leaves glaucous beneath.
18.
Té.
Branchlets usually densely woolly; leaves
lemon green, coriaceous, obovate to nar-
rowly obovate and tapering to a short
petiole; petioles 0-0.3 mm long; bracts
tawny, the apex often retuse; pistils brick
PM a iosne asa ya coe sees as ies eset cae waceyuswes Soa,
Branchlets glabrous to sparsely pubescent;
leaves thin, elliptic to subcircular; petioles
2-20 mm long; bracts brown to blackish;
pistils reddish, purplish or greenish.
S. myrtillifolia,
p. 145
S. nummularia, p. 59
S. rotundifolia ssp
rotundifolia, p. 53
S. rotundifolia ssp
dodgeana, p. 55
S. setchelliana,
p. 77
27
Salix L.
19. Pistillate nectaries shorter than the
stipes; leaf margins distinctly toothed
on lower half; petioles 2-5.6 mm long ....
19. Pistillate nectaries longer than or equal
to the stipes; leaf margins usually en-
tire; petioles usually 4-20 mm long.
20. Leaves dull above, base usually cu-
Neane 4 SNe IMHO: | Beh Ora
20. Leaves glossy above, base acute to
rounded or subcordate.
21. Branches short and erect, some-
times trailing, often glaucous;
plants often rhizomatous; styles
0,:8-1.6 ‘mom feng) £02. Pico ORS .
21. Branches long and trailing, non-
glaucous; styles 0.2-0.8 mm long.
22. Leaves) Ssubciréular. W238. Ni2.,. J 25:
22. Leaves obovate to narrowly ellip-
tic.
23. Leaves obovate, elliptic or
broadly elliptic; pistillate am-
ents generally shorter; pistils
2 WANT TONIG p32) 1i2) oven ecateetes
23. Leaves narrowly elliptic, 2.5-
4.6 cm long; pistillate aments
2.2-5 cm long; pistils 5.2-9.6
UAE aN 2 Speci dads cepen nacrtastai ansaid
28
S. fuscescens,
p. 113
S. sphenophylla,
p. 85
S. stolonifera, p. 72
S. ovalifolia var.
cyclophylla, p. 67
S. ovalifolia var.
ovalifolia, p. 65
S. ovalifolia var.
arctolitoralis, p. 66
Key to the Species
1. Erect shrubs, exceeding 2 dm tall, or trees.
24. Flowering precocious.
25. Pistils glabrous.
26. Stipules absent; branchlets brittle and with
persistent, long villous hairs at the base ...... S. hookeriana,
p: 131
26. Stipules present, often persistent; branchlets
tenacious, without long, villous hairs at the
base.
27. Stipules persistent for several years, linear
to ovate, the apex attenuate; styles longer
than 1.2 mm; nectaries 2-3 times as long
aepbnetstipee:):... crouched: .eecte..nleni: S. lanata ssp.
richardsonii, p. 171
27. Stipules not persistent for more than
one year, elliptic to broadly ovate, apex
rounded; styles shorter than 1.2 mm; nec-
taries shorter.
28. Aments' precocious, on_ floriferous
branchlets 0-0.5 cm long; styles 0.8-
1.2 mm long; branchlets sparsely pu-
bescent; leaves elliptic or obovate ...... S. monticola, p. 124
28. Aments subprecocious, on floriferous
branchlets 0.3-1.3 cm long; styles 0.5-
0.75 mm long; branchlets glabrescent;
leaves narrowly oblong-obovate ............ S. rigida, p. 121
25. Pistils pubescent.
29. Leaves densely white lanate beneath, bright
green above; stipes 0-0.4 mm long.
30. Branchlets densely white yellow, villous-
WOOIIY SHG NOM-QIANCOUS scccceciceiScccsssescceredle S. alaxensis var.
alaxensis, p. 181
30. Branchlets glabrescent and glaucous ........ S. alaxensis var.
longistylis, p. 182
29. Leaves sericeous or densely villous to
sparsely pubescent or glabrescent beneath;
stipes 0.2-2 mm long.
29
Salix L.
31. Branchlets with thick glaucescence ...........
31. Branchlets non-glaucous (rarely thinly so
in S. planifolia).
32. Buds and stipules oily; stipules broad-
ly ovate, margins prominently glandular;
leaves white or grey sericeous-lanate
BeNGat dish. B>.2t dete. SRE Gx.
32. Buds and stipules not oily; stipules
ovate to linear; leaves glabrous, gla-
brate or sericeous beneath.
33. Branchlets velutinous; styles 0.2-0.5
mm long, stipes 0.8-2 mm long ............
33. Branchlets pubescent to _ villous;
styles 0.5-1.8 mm long; stipes 0.2-
0.8 mm long.
34. Stipules narrowly elliptic, not per-
sistent for more than one year, 0.8-
2 BOMATAITIONG 2:50... 00 BERR sens!
34. Stipules linear, often persistent for
two to four years, 3.5-14, up to
32 mm long.
35. Branchlets glabrous or sparsely
DERDESBE OE 2.09R Ol OUa..c0 0)
35. Branchlets densely white grey,
VINOUS les cdprigencuse es Se ae:
24. Flowering coetaneous or serotinous.
36. Pistils glabrous.
37. Introduced species cultivated in some south-
ern centres.
30
S. drummondiana,
p. 186
S. barrattiana, p. 175
S. scouleriana,
py53a
S. planifolia ssp.
planifolia, p. 157
S. planifolia ssp.
pulchra var.
pulchra, p. 159
S. planifolia ssp.
pulchra var.
yukonensis, p. 161
Key to the Species
38. Leaves broadly ovate to narrowly elliptic,
green or pale beneath; branches not
pendulous; staminate flowers with five
SESS, SHOES Sie cco tandevwescepcasw awe
38. Leaves narrowly ovate, glaucous beneath;
branchlets pendulous; staminate flowers
LEED EO -OEMINIGINS aac) asadinesornvidcsadentzecseketenes
37. Indigenous species.
39. Leaves green or pale beneath, non-glau-
cous.
40.
40.
Leaves linear, 7-18 times as long as
wide, margins distantly denticulate;
aments often branched; bracts decid-
uous after flowering; stipes pubescent
Leaves not linear, only 2-5 times as long
as wide, margins serrulate or crenate;
aments unbranched; bracts persistent;
stipes glabrous.
41. Leaves coarsely villous on both sides,
margins glandular serrulate or partly
enbihe-ci ev ee eee! .2c.. dtoomad: tne
41. Leaves glabrous or glabrescent, mar-
gins glandular, crenate to crenate-
serrulate.
42. Shrubs decumbent, 0.1-0.9 m tall;
stipules minute to 1-2 m_ long;
styles 0.3-0.5 mm lontiasses-nai-:
42. Shrubs erect, 0.6-4 m tall, stipules
1-5 mm long; styles 0.5-0.9 mm
lONGiteito nae. <.cccmmtantss<<ceadange oie
39. Leaves glaucous beneath.
43. Immature leaves membranaceous and
translucent, glabrate and green on
both sides becoming glaucous beneath;
leaves and buds with persistent balsam-
like fraarance:+.&.2.@..6aduien «enn
S. pentandra, p. 35
S. babylonica, p. 41
S. interior, p. 43
S. commutata,
p. 135
S. myrtillifolia,
p. 145
S. novae-angliae,
p. 140.
S. pyrifolia, p. 147
31
Salix L.
43. Immature leaves thicker and opaque,
glabrous to pubescent, lacking a balsam-
like fragrance.
44. Petioles glandular near base of leaf;
stamens 5; leaf apex acuminate to
Gardatenw 2 QAR Renee.
44. Petioles non-glandular; stamens 2;
leaf apex acute to rounded.
45. Styles 0.1-0.4 mm long.
46. Stipes 2-3.2 mm _ long, gla-
brous; leaves coriaceous, gla-
brous styles; 0.1-0.2 mm long ......
46. Stipes 0.4-1.2 mm long, pubes-
cent; leaves thin, glabrescent
with ferruginous trichomes per-
sistent along midrib; styles 0.2-
D4 qT ORG: 22.2.2 eR OR eee.
45. Styles 0.5-2 mm long.
47. Stipules absent; leaves pubes-
cent beneath, at least on mid-rib;
branchlets brittle, with long, per-
sistent, villous hairs at base;
styles red ‘in clife AR
47. Stipules present; leaves glabrous
beneath; branchlets tenacious,
lacking persistent hairs at base;
styles greenish.
48. Leaves elliptic or obovate, the
immature green and opaque;
petioles green; branchlets
densely to sparsely villous;
styles 0.6-1.6 mm long ...............
48. Leaves narrowly oblong to nar-
rowly obovate, the immature
reddish and translucent; peti-
oles reddish; branchlets gla-
brescent; styles 0.5-0.75 mm
[1s Sie aeicrnt ee NA viet ROR AU TEE L Nets
32
S. lasiandra, p. 35
S. pedicellaris,
py tts
S. hastata, p. 119
S. hookeriana,
p. 131
S. barclayi, p. 129
S. rigida, p. 121
Key to the Species
36. Pistils pubescent.
49. Stipes 2.8-4.8 mm long, about 10 times as
FONE AS TNE THECIATIOS oo oc.ecs csi sesecccncees. SRIF
49. Stipes 0-2 mm long, 1-2 times as long as
the nectaries.
50. Leaves sericeous beneath, margins glan-
dular serrulate to distantly so.
o1.
of.
Leaves narrowly ovate, 5-7 times as
long as wide, sericeous beneath with
short, white or ferruginous trichomes
oriented toward the apex, margins prom-
inently glandular serrulate, styles 0.3-
O25 (nie TONG: 0.20. fen. Bk eh ee
Leaves narrowly elliptic to obovate, 2.5-
3 times as long as wide, appearing
satiny beneath with matted sericeous
trichomes, margins distantly and incon-
spicuously glandular serrulate to glan-
dular crenate; styles 0.5-0.8 mm long.
50. Leaves densely pubescent to glabrescent
beneath, not sericeous.
D2.
52.
Leaves pale green (non-glaucous) and
glabrescent beneath, margins prom-
inently glandular dotted to glandular
serrulate; immature leaves sericeous
with a mixture of white and ferruginous
trichomes; stipes 0.8-2 mm long ............
Leaves glaucous beneath, variously pu-
bescent, margins entire; stipes 0.1-1.5
mm long.
53. Leaves densely dull white lanate-
floccose beneath, floccose to glabres-
cent above, narrowly elliptic to nar-
rowly ovate, 3.5-7 times as long as
wises styles: #6Gic.5....c aeceitbs..bs. Ac
S. bebbiana, p. 149
S. arbusculoides,
p. 169
S. sitchensis, p. 191
S. maccalliana,
p. 38
S. candida, p. 177
33
Salix L.
53. Leaves not pubescent as above, obo-
vate to broadly or narrowly elliptic,
2-4 times as long as wide; styles
yellow green.
54. Leaves with ferruginous trichomes
sparsely distributed on both sides,
especially on immature leaves ........ S. athabascensis,
p. 103
54. Leaves without ferruginous | tri-
chomes.
55. Petioles 3-15 mm long, yellowish;
stipes 0.5-2°mmalong 128... S. glauca, p. 95
55. Petioles 1-3 mm long, often red-
dish; stipes 0-0.5 mm long.
56. Pistillate aments subglobose,
densely flowered; styles 0.5-
OS2inmlong ates ees. S. brachycarpa ssp.
brachycarpa, p. 89
56. Pistillate aments cylindrical,
loosely flowered; styles 0.2-0.5
Wa ORG) S.ckao eee te S. brachycarpa ssp.
niphoclada, p. 91
34
Pentandrae (Borr.) Schneid.
Section 1. Pentandrae (Borr). Schneid.
1. SALIX PENTANDRA L.
S. pentandra L. Sp. Pl. 1016. 1753.
Description of species
Introduced shrubs or small trees up to 7 m tall; branchlets brown to reddish
brown, glabrous and glossy. Leaves broadly ovate to narrowly elliptic, the
largest mature leaves (3.5) 7-8.5 (11) cm long (excluding apex), (1.5) 2.5-3
(4.3) cm wide and 2.3-2.9 times as long as broad (excluding apex); apex
acuminate on later leaves, 7-12 mm long; base rounded; margins glandular
serrulate; immature leaves reddish and glabrous, mature leaves coriaceous,
the upper side dark green, the lower side green or pale, non-glaucous; peti-
oles 4-10 mm long, glandular at the distal end; stipules minute glandular
lobes or narrowly obovate to transversely ovate, 1.4-4 mm long, deciduous.
Aments coetaneous, on leafy, floriferous branchlets. Staminate aments 2-6
cm long, floriferous branchlets 1-2 cm long; stamens 5; filaments distinct,
pilose below middle; anthers 0.5-0.6 mm long. Pistillate aments 3.5-6 cm
long; floriferous branchlets 1.5-4.5 cm long; pistils ca. 2.5 mm long; capsules
5-6 mm long, glabrous; styles 0.5-1 mm long; stigmas 0.2-0.3 mm long; stipes
0.5-1 mm long, glabrous; nectaries 2, cup-like with lobes abaxially and adax-
ially, sometimes laterally, 0.4-0.6 mm long, about 0.5 times as long as stipe;
bracts narrowly elliptic, apex broadly acute to round, 2-3 mm long, pale
yellow, glabrate adaxially, pubescent at base abaxially, deciduous after
flowering.
Habitat Range
Cultivated. Petersburg, southeastern Alaska; Eur-
asia, from British Isles eastward to
Yenisei and Angara Rivers in central
U:S:5.A.
2. SALIX LASIANDRA Benth.
S. lasiandra Benth. Pl. Hartweg. 335. 1857.
S. lancifolia Anderss. Kg. Sv. Vet. Akad. Handl. 6: 34. 1867. S. /asiandra
var. /ancifolia (Anderss.) Bebb, in Watson, Bot. Calif. 2: 84. 1879.
S. lasiandra var. recomponens Raup, Sargentia 6: 149. 1947. (Type: Raup
& Soper 9076, A).
35
Plate I
Salix lasiandra Benth. Argus 6682
Scale: On all the plates, one square represents one centimetre.
36
Pentandrae (Borr.) Schneid.
Description of species
Shrubs or small trees 1-7 (11) m tall; branches brown, glabrescent or
sparsely lanate; branchlets tawny, brown or reddish, lanate to sparsely
pubescent with spreading trichomes or glabrescent, brittle. Leaves narrowly
to broadly ovate, the largest mature leaves 6.7-14.2 cm long, 1.3-3 cm wide
and 3.7-5.5 (8.7) times as long as wide; apex acuminate to caudate, the
apex or proximal leaves acute; base obtuse to round; margins glandular ser-
rate-crenate to serrulate; immature leaves often reddish and densely lanate
with white and ferruginous trichomes; the upper side of mature leaves gla-
brescent, green and glossy, the lower side sparsely pubescent, sometimes
with ferruginous trichomes, becoming glabrescent, thinly glaucous, pale in
about 20 per cent; petioles 1.3-3 cm long, glandular at distal end near base
of blade, lanate; stipules semi-ovate, 1.4-5.6 mm long, glandular dotted on
margins and at base. Aments coetaneous, on leafy, floriferous branchlets.
Staminate aments 2-3.5 cm long, floriferous branchlets 0.9-2 cm _ long;
stamens 4-5, filaments 3.5-4 mm long, pubescent at base; anthers 0.6-0.8
mm long; nectaries 2, adaxial and abaxial, small, up to 0.5 mm long,
globular. Pistillate aments 2.5-5 cm long, rachis pubescent, floriferous
branchlets 1.3-3.5 cm long; pistils 3-4.8 mm long, glabrous, capsules 5-6.5 mm
long; styles 0.4-0.8 mm long; stigmas 0.2-0.3 mm long, 2 horseshoe-shaped
lobes; stipes 0.9-1.2 mm long, glabrous; nectaries 1, adaxial, 0.2-0.4 mm
long, broader than long, about 0.25-0.5 times as long as stipe; bracts nar-
rowly oblong, apex acuminate, 1.7-3.2 mm long, tawny, pubescent on lower
half, margins distantly denticulate or entire, often revolute, deciduous after
flowering.
Habitat Range
Thickets on river banks and alluvial Boreal: Central Alaska; Yukon River
deposits; wet Calamagrostis-Carex and its tributaries; Matanuska River;
meadows. southeastern Alaska from Glacier Bay
to the head of Lynn Canal (Haines);
Liard River, Yukon Territory; east-
ward in the boreal forest to Saskatch-
ewan; southward in the codillera to
California and New Mexico (Map 4).
Discussion
Salix lasiandra is characterized by 4-5 stamens and 2 nectaries per staminate
flower, narrowly to broadly ovate leaves with long acuminate to caudate
apices and by immature leaves pubescent with ferruginous and white
trichomes.
This species is the western element of a North American complex including
the eastern S. /ucida Muhl. and the central S. serissima (Bailey) Fern. Boivin
(1967) has combined S. /asiandra and S. /ucida under the latter name, main-
taining that they are not distinguishable geographically or morphologically.
Further study may show that this view is tenable.
Several varieties of S. /Jasiandra have been based on variation in leaf glau-
37
Maccallianae Argus
NS
SS
SS
SS
AS oo
SC
7
Map 4 Salix lasiandra
cescence and in branchlet pubescence (Raup 1959). Following this view most
of the Alaska and Yukon material must be referred to var. /ancifolia which
has leaves glaucous or pale beneath and branchlets densely pubescent; others
with glabrous branchlets must be referred to var. /asiandra. S. lasiandra var.
recomponens, with leaves green beneath (non-glaucous) and branchlets
pubescent, described from the Mackenzie River basin, is unknown in our
flora. Branchlet pubescence varies widely in this species and within the
same population densely lanate to glabrescent branchlets may be _ recog-
nized. Furthermore, this variation does not seem to have any geographical
integrity and | am of the opinion that these taxa are minor variants not
deserving taxonomic recognition.
Section 2. Maccallianae Argus*
3. SALIX MACCALLIANA Rowlee
S. maccalliana Rowlee, Bull. Torr. Bot. Club 34: 158. 1907. (Type: McCalla
2252a, US).
*Maccallianae sect. nov.
Folia coriacea, concolora; pistilla sericea, 6-8 mm stamina 2; nectaria 4-6 loba, cupulata, circumdata
longa; styli 0.8-1.2 mm _ longi; bracteae fulvae; stamina. Species typica Salix maccalliana Rowlee.
38
Maccallianae Argus
Description of species
Shrubs 0.9-2.5-3.5 m tall; branches dark reddish brown, glabrous and glossy;
branchlets reddish brown or chestnut brown, glossy, puberulent with short,
curved trichomes becoming glabrescent or sometimes glabrous from the start.
Leaves coriaceous, narrowly elliptic to oblong, the largest mature leaves
5.2-7 cm long, 0.8-2 cm wide and 3.1-4.1 (5.7) times as long as wide;
apex acute to sometimes more or less acuminate; base acute to round;
margins entire and prominently glandular dotted or glandular serrulate to
glandular crenate; immature leaves reddish, sericeous on both sides with
white and ferruginous trichomes; the upper side of mature leaves glabres-
cent, the midrib sometimes remaining puberulent, glossy, the lower side
glabrescent and pale, non-glaucous; petioles 5-10 mm long, yellow brown,
puberulent adaxially; stipules small glandular lobes or sometimes narrowly
elliptic with glandular margins, 0.2-0.5 (2) mm long. Aments coetaneous, on
leafy, floriferous branchlets. Staminate aments 1.8-2.7 cm long, floriferous
branchlets 0.3-1.3 cm long; stamens 2, filaments 4.8-6.5 mm long, pubescent
near base, distinct; anthers 0.8-1.1 mm long; nectaries cup-like and surround-
ing the stamens, usually with 4-6 lobes. Pistillate aments 2-6 cm long,
densely flowered becoming loose in fruit, floriferous branchlets 1-2.8 cm
long; pistils 6-8 mm long, densely sericeous often with white and ferrugi-
nous trichomes, tawny or green, capsules about 8 mm long, tawny, sericeous-
villous; styles 0.8-1.2 mm long; stigmas 0.4-0.5 mm long, 2-lobed; stipes 0.8-2
mm long, densely sericeous; nectaries 1, adaxial, 0.4-0.6 mm long, about 0.5
times as long as stipe; bracts narrowly oblong, apex rounded, 1.6-3.6 mm
long, tawny or lemon green, sometimes brownish toward apex, villous to
glabrescent abaxially.
Habitat Range
Muskegs; wet Carex-Betula glandulo- Boreal, Aspen Parkland: Known in Yu-
sa fens; river margins. kon Territory only from Watson Lake,
southeastern Yukon; along the Liard
River in adjacent British Columbia;
southward in the Rocky Mountains to
Alberta; eastward to Saskatchewan,
southern Manitoba, Ontario, and Que-
’ bec (Map 5).
Discussion
Salix maccalliana is a very distinctive and relatively invariable species. Its
leaves are coriaceous, glossy and green on both sides; the immature leaves
are sericeous with white and ferruginous trichomes; the pistils are large
(6-8 mm long) and densely sericeous with long styles (0.8-1.2 mm _ long)
and are subtended by tawny or lemon green bracts; the stamens are Ssur-
rounded by a cup-like nectary with 4-6 lobes.
The taxonomic relationships of this species have been the subject of much
speculation. Because of the superficial resemblance of its aments to those of
39
Plate Il
Salix maccalliana Rowlee. Argus 4319
40
Subalbae Koidz.
170° 165° 160°
Map 5 Salix maccalliana
S. glauca it has often been aligned with that species (Rowlee 1907; Schneider
1920; Raup 1959). In his discussion Rowlee (1907) also suggested a rela-
tionship with S. /ucida of section Pentandrae on the basis of its leaves and
buds. This suggestion has not been followed by later authors, but it does seem
to have some merit and is further supported by the cup-like staminate nec-
taries and the large, long-beaked capsules. There are certain important points
of discrepancy, however, including the presence of 2 stamens instead of 4-5,
and sericeous rather than glabrous pistils; but | would suggest looking to
section Pentandrae rather than to S. glauca for at least one element of its
ancestry. The very high chromosome number (2n = 190-224, Suda and
Argus 1968) suggests that it is a very complex polyploid, probably with a
diverse ancestry.
Section 3. Subalbae Koidz.
4. SALIX BABYLONICA L.
S. babylonica L. Sp. Pl. 1017. 1753.
Plate Ill
Salix interior Rowlee. (A) Pistillate specimen, Cody 8273, (B) Staminate specimen, Cody 8157
42
Longifoliae Anderss.
Description of species
Introduced trees up to 12 m tall; branchlets slender, pendulous, yellowish to
brown, glabrous. Leaves narrowly ovate, the largest mature leaves 8-12 cm
long, 0.5-1.5 cm wide; apex long-acuminate; base acute; margins serrulate;
immature leaves sericeous, the upper side of mature leaves glabrate, yellow-
ish green, the lower side glaucous; petioles glandular at the distal end;
stipules 2-7 mm long, narrowly ovate, often absent. Aments coetaneous, on
leafy, floriferous branchlets. Staminate aments slender, up to 4 cm long,
floriferous branchlets 0.5-1.5 cm long; stamens 2, filaments distinct, pubes-
cent at base. Pistillate aments 2-3.5 cm long, slender, floriferous branchlets
present; capsules narrowly ovoid, 1-2 mm long, glabrous; styles about 0.5
mm long; stigmas short; stipes very short; nectaries 1, adaxial; bracts pale
yellow, pubescent and caducous.
Habitat Range
Cultivated. Petersburg and Wrangell, southeast-
ern Alaska; originally Asian but nat-
ural range obscured by widespread
cultivation throughout Asia, Europe,
and North America.
Section 4. Longifoliae Anderss.
5. SALIX INTERIOR Rowlee
S. interior Rowlee, Bull. Torr. Bot. Club 27: 253. 1900.
S. exigua ssp. interior Cronq. in Hitchcock, Cronquist, Ownbey and Thomp-
son, Vas. Pl. Pacific NW. 2: 51. 1964.
S. longifolia Muhl. in Muhl. and Willd. Neue Schr. Ges. Nat. Fr. Berlin 4: 238.
1803, non Lam. 1778.
S. longifolia var. pedicellata Anderss. Kg. Sv. Vet. Akad. Handl. 6: 55. 1867.
S. exigua ssp. interior var. pedicellata Cronq. in Hitchcock, Cronquist, Own-
bey and Thompson, Vas. Pl. Pacific NW. 2: 51. 1967.
Description of species
Shrubs 0.5-4 m tall, colonial, with shoots arising from roots; branches grey-
ish; branchlets reddish brown, sparsely sericeous becoming glabrescent.
Leaves narrowly oblong to linear, the largest mature leaves 4-12.8 cm long,
0.3-1 cm wide and 7-18.3 times as long as wide; apex acute; base narrowly
cuneate; margins distantly glandular denticulate to subentire or entire; the
upper side of mature leaves glabrous and green, the lower side sparsely seri-
ceous to glabrescent and pale green, non-glaucous; petioles 0.8-5 mm long;
stipules minute glandular lobes, rarely narrowly elliptic and up to 1.5 mm
43
Longifoliae Anderss.
130°
AS
Map 6 Salix interior
long. Aments coetaneous, on leafy, floriterous branchlets. Staminate aments
2-5 cm long, often branched and bearing lateral secondary aments, florif-
erous branchlets 0.5-5 cm long; stamens 2, filaments 2.4-2.8 mm _ long,
lower half pubescent, distinct; anthers 0.5-0.8 mm long, curved or coiled
after opening; nectaries 2, abaxial and adaxial. Pistillate aments 2.5-6 cm
long, unbranched, floriferous branchlets 1.3-5.5 cm long; pistils long-beaked,
2-4.5 mm long, glabrous, capsules 5-8 mm long; styles 0-0.1 mm long; stig-
mas 0.2-0.3 mm long, 4 broad lobes; stipes 0.6-0.8 mm long, pubescent or
glabrous; nectaries 2, abaxial and adaxial, 0.5-0.8 mm long, 2-3 times as
long as stipe; bracts oblong, apex acute to acuminate, 2.8-3.2 mm long,
tawny to yellow green, glabrous abaxially, tomentose adaxially, inflated,
deciduous after flowering.
Habitat Range
River banks and alluvial deposits Boreal: Yukon River and its trib-.
within the forested region. utaries, extending as far west as
Tanana; south to the Liard River,
Yukon Territory; in the cordillera to
California; eastward across Canada to
northeastern United States (Map 6).
ae
Chamaetia Dumort.
Discussion
Salix interior is a sand bar species spreading colonially, by shoots borne on
roots, over newly formed alluvial deposits. It is characterized by linear, usual-
ly glabrous and distantly glandular denticulate leaves, aments which are
often branched (Argus 1964) and bracts deciduous after flowering.
This species is part of the highly variable and taxonomically complex sec-
tion Longifoliae which includes S. fluviatilis, S. exigua, S. melanopsis, et al.
Revisions of this group have been proposed by Cronquist (1964) and by
Boivin (1967). The section was once thought to be endemic to America
(Schneider 19196) but the southern Asian species S. blakii Gorz, and possibly
others, clearly belongs here.
Section 5. Chamaetia Dumort.
6a. SALIX RETICULATA L. ssp. RETICULATA
S. reticulata L. Sp. Pl. 1018. 1753.
S. reticulata a (var.) glabra Trautv. in Ledeb. Fl. Alt. 291. 1833. Chamitea
reticulata Kerner, Verh. Zool. Bot. Ges. Wien. 10: 277. 1860.
S. reticulata (var.) subrotunda Ser. Essai Mon. Saules Suisse 29. 1815.
S. orbicularis Anderss. in DC. Prodr. 16 (2): 300. 1868. S. reticu/ata (ssp.?)
S. orbicularis Flod. Sv. Vet. Akad. Ark. Bot. 20A: 5. 1926. S. reticulata ssp.
orbicularis Flod. in Hult. Fl. Aleut. 162. 1937.
S. reticulata var. gigantifolia Ball, Proc. Nat. Acad. Sci. 21: 185. 1935.
S. reticulata f. villosa Kimura, in Tatewaki and Kobayashi, J. Fac. Agr., Hok-
kaido Imp. Univ. 36: 31. 1934.
Description of species
Dwarf shrubs, prostrate and rooting along stems; branches light brown with
short internodes; branchlets green to greenish brown, glabrous, rarely glau-
cous; buds glabrous or pubescent at tip, sometimes glaucous, bud scale
persistent at base of shoot. Leaves elliptic-circular to oblong, the largest
mature leaves 1.2-5-6.6 cm long, 0.8-5 cm wide and 1-1.5 times as long as
wide; apex round, obtuse or rarely retuse; base obtuse, rounded or cordate;
margins subentire to indistinctly glandular crenate, revolute; the upper side
of mature leaves dark green and glossy, glabrous or rarely villous with long
straight trichomes, the veins impressed, the lower side pale green and non-
glaucous or rarely glaucescent, sparsely pubescent with long, silky, caducous
trichomes, venation prominently reticulate, sometimes reddish; petioles (3)
10-25 (30-46) mm long, reddish or yellowish, glabrous; stipules minute
glandular lobes 0.2-0.4 mm long. Aments coetaneous on prominent, leafy,
45
Plate IV
.
8, oa. A,
SX
C!
%
SPOS
PRES
Salix reticulata L. (A, B, C) Subspecies reticulata, population sample, Argus 5621
(D) Subspecies glabellicarpa Argus, Argus 6645
46
Chamaetia Dumort.
floriferous branchlets. Staminate aments 1.1-5.2 cm long, floriferous branch-
lets 1.1-3.2 cm long; stamens 2, filaments about 2.8 mm long, pubescent
on lower half; anthers 0.3-0.4 mm long; nectaries 2-3, abaxial and adaxial,
sometimes more or less surrounding stamens. Pistillate aments 1-6 cm long,
floriferous branchlets 1.2-4.5 cm long, as long as vegetative branchlets,
glabrous to coarsely pubescent distally; pistils 2.5-3 mm long, densely
sericeous with white or mixture of white, red violet and ferruginous tri-
chomes, capsules about 4.5-5 mm long, sparsely sericeous; styles 0.2-0.3 mm
long; stigmas 0.2-0.4 mm long, 4-lobed; stipes 0-0.4-0.8 mm long, pubes-
cent; nectaries 2, adaxial, 0.5-0.8 mm long, equal to or 2 times as long as
stipe; bracts oblong to obovate, apex retuse or rounded, 0.8-1.8 mm long,
reddish, greenish, tawny or drying brownish, sometimes white puncticulate,
glabrous or glabrescent abaxially, densely pubescent adaxially.
Habitat Range
Arctic: tundra vegetation including Arctic, alpine: Throughout Alaska and
polygonal tundra, dry tussock tundra, Yukon Territory; southward in the
partly stabilized sand dunes, and Rocky Mountains to British Columbia;
Carex-Eriophorum meadows. Alpine: transcontinental across Arctic and
Dryas tundra; snow accumulation Subarctic Canada to Newfoundland
areas; stabilized talus slopes. In and Greenland; Eurasia (Map 7).
forested regions within mountains, it
may occur in moss both in Picea
glauca woods and in P. mariana
muskegs.
6b. SALIX RETICULATA ssp. GLABELLICARPA Argus
S. reticulata ssp. glabellicarpa Argus, Can. J. Bot. 43: 1021. 1965. (Type:
Calder & Taylor 36347, DAO).
Description of subspecies
Differs from ssp. reticulata in pistils glabrous or sparsely pubescent distally,
glaucous; stipes glabrous; styles 0.2-0.4 mm long.
Habitat Range
Alpine tundra. Alpine: Mount Gastineau near Ju-
neau; Queen Charlotte Islands, Brit-
ish Columbia (Map 8).
Discussion
Salix reticulata is a dwarf, trailing willow characterized by prominently retic-
ulate, elliptic-circular to oblong leaves and aments which are borne on long,
47
Chamaetia Dumort.
Map 7 Salix reticulata ssp. reticulata. Circles based on Skvortsov, 1966, as S. reticulata
leafy, floriferous branchlets. These branchlets are as long and bear as many
leaves as do the vegetative branchlets.
There are two variations within S. reticu/ata that deserve discussion. The
first is represented by S. reticulata ssp. glabellicarpa, a variant with glabrous
and glaucous pistils which was described from the mountains of the Queen
Charlotte Islands. Because of its localized distribution, the absence of gla-
brous pistils elsewhere within the species, and the possibility that it may have
survived the Pleistocene glaciation on a refugium on the Islands, it was as-
signed subspecific rank (Argus 19656). In 1967 it was found to occur also in
the alpine tundra on the slopes of Mount Gastineau near Juneau. In this re-
gion it is sympatric with ssp. reticulata, which is absent from the Queen
Charlotte Islands, and some signs of intergradation were observed. Some
specimens (Argus 6633 and 6644, SASK) were very sparsely pubescent,
whereas others were either glabrous (Argus 6620a, SASK) or had a few
trichomes at the apex (Argus 6632, SASK). This subspecies is to be expected
elsewhere in southeastern Alaska and coastal British Columbia where the
ranges of the two subspecies evidently overlap.
The second variant is represented by S. reticulata f. villosa which has vil-
lous trichomes on the underside of mature leaves. This type of pubescence
resembles that which occurs in S. vestita of the Rocky Mountains and eastern
North America, and would probably be interpreted as an indication of hybrid-
48
Retusae Kern.
170° 165° 160°
Map 8 Salix reticulata ssp. glabellicarpa
ization if the species were sympatric. | have seen specimens with this type
of pubescence throughout the range of ssp. reticu/ata in Alaska and the
Yukon, including Whitehorse, the Bering Sea Coast (Scammon Bay and Cape
Thompson), the Alaska Range (Rainbow Mountain, Donnelly Dome and Eure-
ka Roadhouse), and southeastern Alaska (Coronation Island), and specimens
with villous leaves have been reported from Attu Island and Unalaska (Kimu-
ra 1934). Whether or not this unusual pubescence is a remnant of an ancient
trans-Bering Straits distribution of S. vestita as suggested by Hultén (1967)
is difficult to say, but it does deserve further study.
Section 6. Retusae Kern.
7. SALIX POLARIS Wahl.
S. polaris Wahl. Fl. Lapp. 261. 1812.
S. pseudopolaris Flod. Sv. Vet. Akad. Ark. Bot. 20A (6): 8. 1926. S. polaris
ssp. pseudopolaris Hult. Fl. Alaska and Yukon 3: 510. 1943.
S. polaris var. selwynensis Raup. Contr. Arnold Arb. 6: 144. 1934.
49
Plate V
Salix polaris Wahl. Murray 1042 (left) and Viereck 8361 (right)
50
Retusae Kern.
Description of species
Dwarf shrubs often partly subterraneous, stems rooting, branches reddish
brown, glaucous; branchlets short and not conspicuously trailing, greenish
brown, glabrous; buds glabrous and glaucous, the scale often persistent at
base of shoot. Leaves obovate to narrowly elliptic, the largest mature leaves
1-2-2.8 cm long, 0.8-1-1.8 cm wide and 1.1-1.7-2.5 time as long as wide;
apex round, retuse or obtuse, usually conduplicate when pressed, base cune-
ate or round, inequilateral; margins entire, flat, often reddish, rarely ciliate;
the upper side of mature leaves glabrous or rarely ciliate toward margin,
glossy, the lower side glabrous or rarely pubescent with long, sparse, cadu-
cous trichomes, green and glossy, secondary veins prominently raised; pet-
ioles 2-5-10 mm long, yellow to reddish; stipules absent or minute glan-
dular lobes 0.1-0.4 mm long. Aments coetaneous, on leafy, floriferous
branchlets. Staminate aments 1.5-1.8 cm long, floriferous branchlets 0.5-
1.5 cm long; stamens 2, filaments 4-4.4 mm long, glabrous, distinct; anthers
about 0.6 mm long. Pistillate aments 1.5-3.5 mm long, floriferous branch-
lets 1.2-2 cm long; pistils about 2.5 mm long, reddish and glossy, entirely
pubescent or sparsely pubescent on distal half, capsules 4.8-6.4 mm long,
pale reddish brown, pubescent; styles 0.7-1.6 mm long, entire or bifurcate;
stigmas 0.2-0.6 mm long, 4 linear lobes; stipes 0.2-0.7 mm long, glabrous
or pubescent; nectaries 1, adaxial, 0.9-1 mm long, often 2-5 times as long as
stipe; bracts oblong to broadly obovate, apex rounded, uniformly brown or
dark brown, sometimes bicolour, sparsely pubescent to glabrescent abaxially,
sparsely pubescent adaxially.
Habitat Range
Tundra vegetation; late snowbed and Arctic, alpine: Bering Sea _ region;
snow flush areas; alpine scree slopes. eastern Brooks Range; central Alas-
ka; Alaska Range; southward to the
St. Elias Mountains, Yukon Territo-
ry; eastward to northern British Co-
lumbia; Eurasia (Map 9).
Discussion
Salix polaris is a dwarf species characterized by pubescent pistils, long styles
(0.7-1.6 mm long) and leaves non-glaucous beneath.
This species is commonly misunderstood in the North American literature,
and many specimens in herbaria are misidentified. Wiggins and Thomas
in A flora of the Alaskan Arctic Slope (1962) completely misunderstood the
species (as S. pseudopolaris), and many of the specimens they cited are S.
ovalifolia var. glacialis or S. ovalifolia var. ovalifolia. | have not seen any
authentic S. polaris from the Arctic slope of Alaska (reported by Hultén 1968,
Map 4), although it does occur in the eastern Brooks Range.
In my opinion the characteristics that have been used to distinguish S.
pseudopolaris from S. polaris, namely, light-coloured bracts with wavy hairs
(Hultén 1943) and a greater number of flowers in the aments (Skvortsov
51
Retusae Kern.
175° 170° 155° 135° 130°
5 WK
Map 9 Salix polaris. Circles based on Skvortsov, 1966
1966), and the characteristics that have been used to distinguish S. polaris
var. selwynensis, namely, longer aments and leaves ovate-elliptic or obovate
(Raup 1934), are inconsequential and scarcely extend the normal range of
variation of S. polaris. The taxonomic confusion results from a lack of spec-
imens, and from typological thinking that fails to recognize and understand
population variation. In his discussion of S. pseudopolaris Floderus (1926)
referred to the hybridogenous nature of many of the specimens he studied
and suggested that some characteristics were derived from crossing with S.
arctica, S. glauca or S. chamissonis. In my opinion Floderus’ presumption of
hybridity is, in most cases, conjectural and originates from a lack of appre-
ciation of character variation. However, as | studied S. polaris, | was repeat-
edly impressed with the apparent morphological links with S. rotundifolia,
S. arctica and S. ovalifolia and with the heterogenous composition of the
species. The material available to me is inadequate to fully understand this
species and extensive field and laboratory studies are indicated.
Salix polaris is apparently related to S. rotundifolia, S. phlebophylla and
S. stolonifera and is similar morphologically to some variants of S. arctica.
The comparison table (Table 2) will aid in distinguishing them.
52
Retusae Kern.
Table 2 Comparison table: Salix polaris, S. rotundifolia, S. stolonifera, S. phlebophylla and S.
arctica
Characteristics polaris rotundifolia stolonifera phlebophylla arctica
leaves non-glaucous non-glaucous glaucous non-glaucous glaucous
beneath beneath beneath beneath beneath
rarely ciliate sometimes non-ciliate + ciliate non-ciliate
margins ciliate margins margins margins margins but
with long
trichomes
beneath
non-persistent often persis- non-persistent persistent non-persistent
tent for several and skeleton-
years ized
pistillate more than 4-15-flowered more than more than more than
aments 15-flowered 25-flowered 25-flowered 25-flowered
pistils pubescent usually usually usually pubescent
glabrous glabrous pubescent
nectary 2-5 times 1-3 times 1.5-3 times less than 1.5-4 times
length stipe stipe stipe or equal to stipe
stipe
8a. SALIX ROTUNDIFOLIA Trautv. ssp. ROTUNDIFOLIA
S. rotundifolia Trautv. Nouv. Mem. Soc. Nat. Mosc. 2: 304. 1832.
S. polaris var. leiocarpa Cham. Linnaea 6: 542. 1831. S. leiocarpa Cov. Proc.
Wash. Acad. Sci. 3: 338. 1901.
S. behringica v. Seemen, Engler Bot. Jahrb. 21. 52: 6. 1895. (Type: Krause
85, US).
Description of species
Dwarf shrubs with slender branches arising from a taprooted caudex, form-
ing a highly branched, largely subterranean shrub about 2-3 cm tall; bran-
ches yellow brown, glabrous, glossy and with short internodes; branchlets
yellow brown, yellowish red or reddish green, sometimes glaucous, bearing
2-3 leaves; buds tawny to greenish or reddish, sometimes glaucous, the scale
persistent at base of shoot. Leaves circular, elliptic or sometimes narrowly
elliptic, the largest mature leaves 5-10-14 mm long, 4.2-6-9 (11.2) mm wide
and (0.9) 1.2-1.3 (2.2) times as long as wide; apex rounded to more or less
retuse; base rounded, narrowly cuneate in narrower leaves; margins entire,
usually reddish, sometimes ciliate, revolute; the upper side of mature leaves
glossy and glabrous, primary veins usually prominently raised, sometimes
impressed, the lower side glossy, glabrous and green, non-glaucous, the 3-4
secondary veins prominently raised and veinlets evident, marcescent leaves
often persistent for several years but not becoming skeletonized; petioles
1.4-2-3 mm long, tawny and glabrous; stipules usually minute glandular
lobes, 0.1-0.2 (0.4) mm long, yellowish or reddish. Aments coetaneous or
53
Plate VI
Salix rotundifolia Trautv. (A, B)
(Rydb.) Argus, Murray 788
54
Subspecies rotundifolia, Argus 5695, (C) Subspecies dodgeana
Retusae Kern.
serotinous, terminal on previous year’s shoot, and borne on floriferous
branchlet bearing 2 leaves. Staminate aments 0.3-1 cm long, 7-15-flowered,
floriferous branchlets 3-7 mm long; stamens 2, filaments about 1.8-4 mm
long, glabrous; anthers 0.4-0.6 mm long; nectaries long and narrow. Pistil-
late aments 0.7-2 cm long, rachis pubescent, 4-12 (15)-flowered, floriferous
branchlets 0.7-2.5 cm long, as long or longer than aments; pistils 1.5-2.5
mm long, glabrous or rarely sparsely pubescent at tip, glossy and reddish
brown, capsules 4-6-7.2 mm long, beak sometimes flattened; styles 0.5-1
mm long, entire or sometimes slightly bifid; stigmas 4 linear lobes, 0.4-0.5
mm long; stipes 0.4-0.8 mm long, glabrous or sometimes pubescent, nec-
taries 1, adaxial, 0.9-1.6 mm long, reddish, equal to or up to 3 times as long
as stipe; bracts broadly obovate, apex rounded or retuse, 1.6-2.8 mm long,
uniformly brown or bicolour and pale reddish brown at base, usually glabrous
or sparsely pubescent abaxially, sparsely pubescent adaxially with long
straggly trichomes appearing as cilia around margin.
Habitat Range
Tundra. Arctic, alpine: St. Lawrence Island;
Bering Sea islands; Seward Peninsula;
northern Alaska to Barter Island;
Aleutian Islands; Alaska Peninsula;
Kodiak Island; Talkeetna Range;
Alaska Range; Eagle Summit, central
Alaska; Ruby Range, southwestern
Yukon; to the Mackenzie Mountains,
Northwest Territories (Map 10).
8b. SALIX ROTUNDIFOLIA ssp. DODGEANA (Rydb.) Argus
S. rotundifolia ssp. dodgeana (Rydb.) Argus, Can. J. Bot. 47: 795. 1969.
S. dodgeana Rydb. Bull. N.Y. Bot. Gard. 1: 277. 1899. (Type: Rydberg &
Bessey s.n. NY).
Description of subspecies
Differs from ssp. rotundifolia in leaves smaller and narrower, 4-6 mm long,
1.7-3.6 mm wide and 1.7-2.7 times as long as wide, venation on upper side
of mature leaves less prominent; petioles shorter, 0.8-1.6 mm long. Pistillate
aments with fewer flowers, 2-4 (9)-flowered (Table 3).
Habitat Range
Tundra. Alpine, Arctic: Disjunct distribution
in North America; northwestern Wyo-
ming and adjacent Montana; Kluane
Lake region, Yukon Territory; Mac-
kenzie Mountains, Northwest Terri-
55
Retusae Kern.
Map 10 Salix rotundifolia ssp. rotundifolia. Circles based on Skvortsov, 1966
tories. It also occurs sporadically
throughout the range of ssp. rotun-
difolia in Alaska (Map 11).
Discussion
Salix rotundifolia is a dwarf, largely subterranean shrub characterized by
circular or elliptic leaves which have 3-4 prominent veins and are glossy and
green on both sides. The aments have few flowers (4-15) and the nectaries
usually exceed the stipes.
There is some variation in pistil indumentum from usually glabrous to
sometimes sparsely pubescent on the beak or, rarely, sparsely pubescent all
over. The pubescent-capsuled variants have been named S. rotundifolia f.
pilosiuscula Schneider (1919a), and Hultén (1943) regarded specimens with
pilose capsules as the hybrid S. phlebophylla x rotundifolia. To base a con-
clusion of hybridity on pistil indumentum alone is unsound, for it requires the
assumption that this characteristic is invariable. There are several species in
Alaska and the Yukon which have been shown to have variable pistil pubes-
cence, including S. hookeriana, S. reticulata, S. ovalifolia and S. phlebophylla,
and the possibility that the pilose-capsuled forms of S. rotundifolia are part of
the normal variation within this species must be seriously entertained.
56
Retusae Kern.
Map 11 Salix rotundifolia ssp. dodgeana. Siberian locality: C. Wright, 1853-1856,
Arakamechetchene Island, NY, US 26252
However, hybridization apparently does occur between S. phlebophylla
and S. rotundifolia and, within sympatric populations, intermediate specimens
can be detected on the basis of discordant combinations of characteristics
including leaf shape, leaf pubescence, skeletonization of leaves, nectary
length, and pistil indumentum. On the basis of these characteristics | have
determined a series of specimens from the Arctic slope and central Alaska
as S. rotundifolia > phlebophylla and S. phlebophylla > rotundifolia to in-
dicate the varying degrees of intermediacy. There are no evident signs of
infertility in the intermediate specimens and further study is necessary in
order to establish the origin of the intermediates and to prove the occurrence
of hybridization (Map 2).
Salix rotundifolia is closely related to S. phlebophylla and S. nummularia
(Table 4), and is similar to S. polaris and S. stolonifera (Table 2).
In 1967 Hultén noted that S. behringica had long been neglected by most
modern authors and that the type specimen was probably destroyed when the
Berlin Museum was bombed. There is a fragment of the type collection at the
U.S. National Herbarium which | have been able to examine (Krause, A. &
A. 85, Luetke Harbour, St. Lawrence Bay, Siberian Coast of Bering Sea,
1881, US 411218). The specimen consists of 3 leaves and 4 somewhat dam-
aged capsules. The leaves are elliptic, glabrous and non-glaucous; the cap-
57
Retusae Kern.
Table 3. Comparison table: Salix rotundifolia ssp. rotundifolia and ssp. dodgeana
Characteristics ssp. rotundifolia ssp. dodgeana
leaves 5-10-14 mm long 4-6 mm long
4.2-6-9 mm wide 1.7-3.6 mm wide
length/width (0.9) length/width
fe 1.3-t2.2) 1.7-2.7
apex acute and
+ conduplicate
when pressed
apex round to retuse,
rarely acute
veins faint above
0.8-1.6 mm
2-4 (9) -flowered
veins prominent above
1.4-2-3 mm
4-12-flowered
petiole length
pistillate aments
Table 4 Comparison table: Salix rotundifolia, S. phlebophylla and S. nummularia
Characteristics rotundifolia phlebophylla nummularia
persistent and not persistent
skeletonized
leaves often persistent,
not skeletonized
glossy highly glossy glossy
not arcuate, usually
pressing flat
arcuate, conduplicate
when pressed
not arcuate,
pressing flat
pistillate aments 4-12 (15) -flowered 25-flowered 4-5-flowered
pistils glabrous usually pubescent glabrous
nectary length 1-3 times stipe less than or about 2 times
equal to stipe stipe
capsule length 4-6-7.2 mm 2.9-4.88 mm ca. 3.5 mm
habitat wet tundra dry tundra dry tundra
sules and stipes are glabrous and the nectaries are slightly longer than the
stipes. | am of the opinion that this fragmentary material represents S.
rotundifolia.
Salix rotundifolia ssp. dodgeana is a diminutive race of the species which
is distinguishable only on the basis of quantitative characters (Table 3).
Salix dodgeana traditionally has been given a very narrow circumscription
(Raup 1959). However, when the variation, present even within the type
locality, has been taken into consideration it is evident that it intergrades
with S. rotundifolia (Argus 1969). It has been assigned subspecific rank be-
cause of its distinctive cordilleran distribution.
58
Retusae Kern.
9. SALIX NUMMULARIA Anderss.
S. nummularia Anderss. in DC. Prodr. 16 (2): 298. 1868.
S. tundricola Schljak. Bot. Mater. Herb. Bot. Inst. Akad. Nauk SSSR 16: 67.
1954. S. nummularia ssp. tundricola Love and L6ve, Bot. Not. 114: 51. 1961.
Description of species
Dwarf, trailing shrubs; branches slender, arising from a stout caudex, chest-
nut brown, glossy, glabrescent and sometimes glaucous; branchlets yellow
brown, sparsely pubescent with spreading trichomes. Leaves subcircular, the
largest mature leaves about 10-12 mm long, 8-10 mm wide and 1.2 times as
long as wide; apex round to retuse; base subcordate to round; margins entire
but with 3-4 pairs of glands or glandular teeth on lower half; mature leaves
glabrous and glossy on both sides, green and non-glaucous beneath, the
lowermost leaves with sparse, long trichomes beneath, venation prominently
reticulate and with 4-5 pairs of secondary veins, leaves may persist for 2-3
years but are more commonly deciduous each year; petioles 1.5-2 mm long,
reddish; stipules minute glandular lobes. Aments probably coetaneous or
serotinous, on short floriferous branchlets with 2-3 leaves. Staminate aments
unknown in the flora of North America. Pistillate aments about 3-5 mm long,
4-5-flowered, floriferous branchlets about 1.5 mm long; pistils about 2 mm
long, brownish and glabrous, capsules about 3.5 mm long; styles 0.2-0.4
mm long; stigmas 0.3-0.5 mm long, 4 linear lobes; stipes about 0.4 mm long,
glabrous or pubescent; nectaries 1, adaxial, about 0.8 mm long and about 2
times as long as stipe; bracts obovate, apex rounded to retuse, about 1.2 mm
long, pale brown, sparsely pubescent adaxially, more or less glabrous abaxially.
Habitat Range
Tundra. Arctic: Known in Alaska only from St.
Paul Island; widely distributed in
northern U.S.S.R. from the Kola
Peninsula to the Chukotsk Peninsula
(Skvortsov 1966) and in Transbaika-
lia (Map 12).
Discussion
The Eurasian Salix nummularia has not previously been known to occur in
North America. | have seen three specimens from St. Paul Island (Cole, July
8-9, 1941, WIS, ISC and Johnston, June 14, 1925, NA) which compare very
favourably with S. nummularia from the U.S.S.R. and seem to represent that
species. This species is characterized by prominently reticulate leaves which
have several glandular teeth on the lower half of the blade, aments produced
by lateral buds and often elongate and trailing vegetative shoots. These
characteristics contrast with S. rotundifolia which has non-reticulate, entire
59
Plate VII
Salix nummularia Anderss. Skvortsov 10589, 23 VII 1964, Northern Urals, U.S.S.R., GWA
60
Retusae Kern.
LA,
Map 12 Salix nummularia. Circles based on Skvortsov, 1966
leaves, aments produced by subterminal buds and short vegetative shoots
(Table 4).
| am accepting the opinion of Skvortsov (1966) in treating S. tundricola as
a synonym of S. nummularia. The Love and Léve (1961) combination of
these taxa which was accepted by Hultén (1968) was not accompanied by a
discussion and therefore cannot be evaluated.
10. SALIX PHLEBOPHYLLA Anderss.
S. phlebophylla Anderss. Ofvers. Vet. Akad. Forh. (Stockh.) 15: 131. 1858.
S. anglorum Cham. Linnaea 6: 541. 1831.
S. paleoneura Rydb. Bull. N.Y. Bot. Gard. 1: 267. 1899. (Type: Murdock s.n.
GH).
Description of species
Dwarf shrubs forming compact mats up to 2 m in diameter, stems thick and
arising from a taprooted caudex, stems partly subterranean and rooting,
mostly aerial and clothed with persistent, skeletonized leaves; branches red-
61
Plate VIII
Salix phlebophylla Anderss. Argus 5827
62
Retusae Kern.
dish brown, glossy and glabrous; branchlets glabrous, non-glaucous. Leaves
marcescent, narrowly obovate to broadly obovate or broadly elliptic, the
largest mature leaves 7-11-15 mm long, (2.5) 3-5-11 mm wide and 1.1-1.7-
2.3 (3.5) times as long as wide; apex obtuse, sometimes retuse, often con-
duplicate when pressed; base cuneate to broadly cuneate; margins entire,
often reddish and somewhat ciliate; the upper side of mature leaves glabrous
and glossy, the lower side sparsely pubescent with long, straight, caducous
trichomes, glossy and non-glaucous, the 3-4 or 5 secondary veins and some-
times the tertiary veins prominent on both sides; petioles 1.2-2.4-3.2 (4.8)
mm long, tawny and sparsely pubescent; stipules minute glandular lobes,
0.1-0.2-0.4 mm long. Aments coetaneous, borne on leafy, floriferous
branchlets. Staminate aments 1.3-2.5 cm long, floriferous branchlets 5-13
mm long; stamens 2, filaments 2.5-4 mm long, glabrous; anthers 0.3-0.5
mm long; nectaries 1, adaxial or 2, adaxial and abaxial, 0.4-0.6 mm long,
reddish or greenish. Pistillate aments 1.6-2.5 cm long, rachis pubescent and
bearing more than 25 flowers, floriferous branchlets 0.8-2.4 cm long; pistils
about 1.8 mm long, sericeous, with short, refractive trichomes, occasionally
pubescent only on beak or entirely glabrous, non-glaucous, capsules 2.9-4.8
mm long, reddish, sparsely sericeous or glabrescent; styles 0.3-1 mm long,
sometimes slightly bifid; stigmas 0.2-0.3 mm long, 2-lobed and broad; stipes
0.4-1.4 mm long, sparsely pubescent; nectaries 1, adaxial, 0.4-1 mm long,
shorter than or equal to stipe; bracts broadly oblong, apex rounded, 1-1.3
mm long, dark brown to black or bicolour, pubescent on both sides with long
straight or curly trichomes.
Habitat Range
Tundra. Arctic, alpine: St. Lawrence Island;
northern Alaska; northern Yukon;
Alaska Peninsula; mountains of cen-
tral Alaska; Alaska Range; to south-
eastern Yukon (Map 13).
Discussion
Salix phlebophylla is a dwarf, mat-forming shrub characterized by stems
bearing persistent, skeletonized leaves. Leaves may be persistent in other
species, such as S. rotundifolia and S. planifolia ssp. pulchra, but they do
not become skeletonized as in S. phlebophylla. The many-flowered aments,
pubescent pistils and nectaries which are shorter than the stipes also distin-
guish this species from S. rotundifolia. See S. rotunifolia for a discussion of
hybridization and Tables 2 and 4 for a comparison with related and similar
species.
63
Plate IX
Salix ovalifolia Trautv. (A) Variety arctolitoralis (Hult.) Argus, Argus 5945, (B) Variety ova-
lifolia, Argus 5788, (C) Variety glacialis (Anderss.) Argus, Argus 5981, (D) Variety cyclophylla
(Rydb.) Ball, Harms 5649
64
Retusae Kern.
Map 13 Salix phlebophylla. Circles based on Skvortsov, 1966
41a. SALIX OVALIFOLIA Trautv. var. OVALIFOLIA
S. ovalifolia Trautv. Nouv. Mem. Soc. Nat. Mosc. 2: 306. 1832.
S. ovalifolia var. camdensis Schneid. Bot. Gaz. 66: 139. 1918. (Type: Johan-
sen 116, CAN).
S. flagellaris Hult. Sv. Bot. Tidskr. 34: 376. 1940. (Type: Walpole 1672, US).
i Ee Fo eS ee
Description of species
Dwarf trailing shrubs; branches long, slender and trailing sometimes up to
45 cm long, arising from a stout caudex with a strong taproot, yellow or
greenish brown, glabrous or rarely sparsely pubescent, glossy; branchlets
long and trailing, yellow brown or greenish brown, glabrous or sparsely
pubescent toward distal end becoming glabrescent, non-glaucous; the distal
9-3 buds vegetative, the others reproductive, buds glabrous or sparsely
pubescent, bud scales persistent at base of shoot. Leaves obovate, elliptic
or broadly elliptic, the largest mature leaves 1.3-1.8-2.8 cm long, 0.7-1.2-
4.8 cm wide and 1.1-1.6-2.2 times as long as broad; apex obtuse, acute or
65
Retusae Kern.
round; base round, subcordate or acute, inequilaterial; margins entire, flat or
slightly revolute, reddish, often ciliate; immature leaves sparsely villous with
long straight trichomes, becoming glabrescent; the upper side of mature
leaves glabrescent and glossy, the venation prominently reticulate, the lower
side sparsely villous to glabrescent, glaucous, sometimes purplish, venation
reticulate; petioles 2-4-7 mm long, reddish or yellow, concave adaxially and
glabrous or sparsely ciliate; stipules minute glandular lobes, 0.2-0.6 (1.6)
mm long, the longer are narrowly elliptic prophylla sometimes with glandular
margins. Aments coetaneous, on leafy, floriferous branchlets. Staminate
aments 0.6-1.8 cm long, floriferous branchlets 0.3-1.2 cm long; stamens 2,
filaments 2-3.6 mm long, distinct and glabrous; anthers 0.3-0.5 (0.6) mm
long. Pistillate aments 0.9-1.5-2.8 cm long, floriferous branchlets 0.5-1.5-
2.5 cm long; pistils 2.5-4 mm long, dark purple, reddish or reddish brown,
glabrous and glaucous with about 11 per cent somewhat pubescent, cap-
sules 5.2-6.5 mm long, greenish or reddish brown, glabrous and glaucous;
styles 0.2-0.8 mm long; stigmas 0.3-0.6 mm long, 4-lobed; stipes 0.2-0.7-
1.4 mm long, glabrous or sometimes pubescent; nectaries 1, adaxial, rarely
2, adaxial and abaxial, 0.5-1-1.4 mm long, with two linear or several irregular
lobes, reddish, 1-3 times as long as stipe; bracts narrowly to broadly oblong,
apex rounded or rarely acute, 1.2-2 mm long, dark brown to blackish, some-
times bicolour and reddish at base, sparsely pubescent with straight or
curly trichomes.
Habitat
Usually at sea level on coastal beach
ridges; gravel spits; wet, Carex aqua-
tilis-Eriophorum meadows; _ saline
marshes; occasionally in upland
tundra.
Range
Arctic: Firth River, Yukon Territory;
westward along the coast to Alaska
Peninsula; Kodiak Island: Aleutian
Islands; Asia: Chukotsk Peninsula
(Map 14).
11b. SALIX OVALIFOLIA var. ARCTOLITORALIS (Hult.) Argus
S. ovalifolia var. arctolitoralis (Hult.) Argus, Can. J. Bot. 47: 795. 1969.
S. arctolitoralis Hult. Sv. Bot. Tidskr. 34: 373. 1940. (Type: Anderson 4705,
US).
Description of variety
Differs from var. ovalifolia in leaves sometimes narrowly elliptic, the largest
mature leaves 2.5-4.6 cm long, 1-2.2 cm wide and 1.6-3.4 times as long as
wide; petioles 4-16 mm long. Pistillate aments 2.2-5 cm long, floriferous
branchlets 1-4 cm long; pistils 5.2-9.6 mm long.
Habitat Range
Arctic: Northeastern Alaskan coast;
Unalakleet to Point Lay; northern
Yukon; Mackenzie Delta (Map 15).
Sea level; coastal beach ridges and
sand spits; tundra meadows; low
ridges near coast.
66
Retusae Kern.
160° 135° 130°
Map 14 Salix ovalifolia var. ovalifolia. Circles based on Skvortsov, 1968
11c. SALIX OVALIFOLIA var. CYCLOPHYLLA (Rydb.) Ball
S. ovalifolia var. cyclophylla (Rydb.) Ball, Proc. Nat. Acad. Sci. 21: 184.
1935.
S. cyclophylla Rydb. Bull. N.Y. Bot. Gard. 1: 274. 1899. (Type: Macoun
16645, NY).
S. rotundata Rydb. ex Macoun, Plants Pribilof Is., in D. Jordan, Fur Seals and
Fur-Seal Is. N. Pacific Ocean 3: 571. 1899, nom. nud., non Forbes 1829.
Description of variety
Differs from var. ovalifolia in leaves subcircular, apex round or retuse, the
largest mature leaves 1-1.5 times as long as wide, prominently reticulate
above; branches stout, not as conspicuously trailing.
Habitat Range
Sea level to ca. 100 ft on shores of Arctic: Bering Sea islands; Aleutian
lakes and lagoons; Empetrum tundra; Islands; from Rat Islands to Stepovak
67
Retusae Kern.
160° 155° 130° 125°
Map 15 Salix ovalifolia var. arctolitoralis
in moss on rocky pavement; mead- Bay, Alaska Peninsula (Map 16).
ows; beach ridges.
11d. SALIX OVALIFOLIA var. GLACIALIS (Anderss.) Argus
S. ovalifolia var. glacialis (Anderss.) Argus, Can. J. Bot. 47: 798. 1969.
S. glacialis Anderss. Ofvers. Vet. Akad. Forh. (Stockh.) 15: 131. 1858.
Description of variety
Differs from var. ovalifolia in leaves sometimes ovate, the largest mature
leaves 8.5-14 mm long, 4.5-7-9 mm wide; mature leaves sparsely pubescent
to glabrescent on both sides; margins remaining ciliate; petioles 1.1-3.2 mm
long. Staminate aments 4-9 mm long and globose, floriferous branchlets 2-3
mm long. Pistillate aments 0.7-1.3 cm long and globose, floriferous branch-
lets 2-8 mm long; pistils pubescent or rarely glabrous, capsules 4.3-5.2 mm
long, non-glaucous, 85 per cent pubescent or at least partly so, 15 per cent
glabrous; stipes 0.2-0.8 mm long.
68
Retusae Kern.
170° 140°
Map 16 Salix ovalifolia var. cyclophylla
Habitat Range
Coastal sandy-gravel spits. Arctic: Point Barrow; Meade River;
Topagoruk River; possibly Collinson
Point, Camden Bay (Map 17).
Discussion
Salix ovalifolia is characterized by a dwarf, trailing habit, glabrous and often
glaucous pistils, nectaries 1-3 times as long as the stipes and styles 0.2-0.8
mm long.
This species is closely related to S. sto/onifera and with that species consti-
tutes an arctic-alpine complex occurring in coastal Alaska and Yukon. There
is a zone of overlap between these species from the Kenai Peninsula to Ko-
diak Island and King Cove, Alaska Peninsula, within which intergradation
may occur. However, the material available to me is inadequate to determine
whether or not the taxa hybridize and to what degree they intergrade. Be-
cause of this lack of information and because each of the taxa occupy differ-
ent kinds of habitats, | am treating them as species.
Salix ovalifolia consists of four varieties: 1) var. ovalifolia, 2) var. arcto-
litoralis, 3) var. cyclophylla, and 4) var. glacialis. Salix ovalifolia var. ovali-
69
Retusae Kern.
Map 17 Salix ovalifolia var. glacialis
folia is a wide-ranging taxon occurring throughout the range of the other
varieties. It generally occupies habitats close to the ocean and is character-
ized by long, trailing branches and branchlets bearing small, obovate to
elliptic or sometimes subcircular leaves and glabrous and glaucous pistils.
Salix ovalifolia var. arctolitoralis has a disjunct distribution in northwestern
Alaska and the Mackenzie Delta region. It is characterized by large, some-
times narrowly elliptic leaves, large aments, large floriferous branchlets and
large pistils. It is essentially a robust form of the species and, as suggested
by its disjunct distribution, may be simply an environmental modification.
However, because of its characteristic appearance | have recognized it as a
variety until the necessary study has been completed (Argus 1969).
Salix ovalifolia var. cyclophylla is a Bering Sea variant of the species
characterized by cubcircular, prominently reticulate leaves and branchlets
often not as long and trailing as in the other varieties. Hultén (1967) notes
that this taxon seems to occupy upland rather than coastal habitats and for
that reason he would treat it as a species. In general his observation seems to
be supported. However, | have seen specimens of this variety from ‘‘sea
level” and the occasional specimen of var. ovalifolia from upland situations.
This taxon seems to be insufficiently distinct morphologically to be treated
as a species.
70
Retusae Kern.
Salix ovalifolia var. glacialis is an unusual variant of the species with a
very limited distribution on the Arctic Coast of Alaska. The type material,
which | have not seen, was collected by Captain Pullen, ‘between Cape
Barrow and Mackenzie River’ (Andersson 1858) and may have been col-
lected from the extensive Point Barrow population located on the sand spit
between the air strip and the old Eskimo village of Nuwuk. Andersson's
(1858) description depicts very well the distintive Point Barrow population
which is characterized by small, ovate leaves with ciliate margins, small
globose aments, short floriferous branchlets and usually pubescent pistils.
Point Barrow has been visited by many botanists and there are numerous
specimens of this taxon from the vicinity of Nuwuk. The overwhelming
number of specimens alone gives this taxon a disproportionate significance
and it may not even deserve varietal rank. However, it is a distintive local
variant which may prove to have some phytogeographic or evolutionary sig-
nificance and for that reason | have assigned it varietal status (Argus 1969).
Two of these varieties, the distinctive var. glacialis and the large var.
arctolitoralis, are quantitatively distinct from one another in reference to leaf
length, leaf width, leaf length/width, petiole length, floriferous branchlet
length and pistillate ament length (Fig. 2). However, their range of variation
overlaps that of the other varieties and the species as a whole presents a
completely intergrading variation pattern. Pistil indumentum also varies
significantly within the species. An analysis of pistil indumentum in popula-
tion samples of var. ovalifolia and var. glacialis and in herbarium specimens
of var. arctolitoralis and var. cyclophylla is shown on the graph in Fig. 3.
Within the species, excluding var. glacialis, 60 per cent of the pistils are
glabrous and glaucous, 29 per cent glabrous and 11 per cent variously
pubescent. In var. g/acialis pistil indumentum varies conversely. Within the
Point Barrow population 86 per cent of the pistils are pubescent, although
sometimes only sparsely so, and 14 per cent are glabrous.
Pistil indumentum in Salix has been assumed to be relatively invariable
and variation in pistil indumentum such as described here is often attributed
to hybridization. If we accept this assumption as an explanation of variation
in pistil pubescence in S. ovalifolia, an argument could be made for hybridi-
zation between: 1) the sympatric S. ovalifolia var. glacialis (pubescent
pistils) and S. rotundifolia (glabrous pistils), and 2) the glabrous-pistilled
varieties of S. ovalifolia and the pubescent-pistilled S. ph/ebophyl/a or S.
arctica. However, such a conclusion based on single character variation is
untenable unless it is supported by additional evidence of hybridization, such
as a reduction in fertility or a recombination of other characteristics. Only in
the case of several specimens of the putative hybrid S. arctica x ovalifolia
were other characteristics found to be recombined (see S. arctica and Table
6). It is therefore concluded that the variation in pistil indumentum described
here is the normal variational pattern for the species and not the result of
hybridization.
Hultén (1968) regards S. glacialis as the hybrid S. arctica x _ ovalifolia.
However, this assumption is unlikely since at Point Barrow, where glacialis
is most abundant, neither S. arctica nor ‘‘typical” S. ovalifolia occurs.
71
Retusae Kern.
others
ia aa RT RS 2,
A glac arcto
others
once terrae
5 10 20 50 mm
others
fe name oreremcmmmeace
0 ee ee 10 20 40mm
poets
Ee: glac parcto
ae aa aay a 7 een Femme Thea
others
F cyclo arcto
PR
1 2 = 4
Fig. 2. The variation of some morphological petiole length, (F) relative leaf shape
characteristics in Salix ovalifolia: (A) leaf (length/width). Code: glac = var. glacialis,
length, (B) pistillate ament length, (C) florif- arcto = var. arctolitoralis, cyclo = var. cyclo-
erous branchlet length, (D) leaf width, (E) phy/la. Logarithmic scales used in A-E.
12. SALIX STOLONIFERA Cov.
S. stolonifera Cov. Proc. Wash. Acad. Sci. 3: 333. 1901. (Type. Walpole 1075,
US).
Description of species
Dwarf shrubs arising from a stout caudex, the short branches may trail on the
surface or if buried may spread underground rhizomatously; branches dark
reddish brown, glabrous and glossy, sometimes glaucous; branchlets greenish
72
Retusae Kern.
60
a
var. glacialis Sa’
—a@ other vars. of
Percent of Population
glabrous glabrous artly pubescent
pubescent
&
glaucous
Pistil Indumentum
Fig. 3. The variation in pistil indumentum in Salix ovalifolia
or greenish brown, glabrous and glossy, sometimes glaucous; buds glabrous
and usually glaucous. Leaves broadly obovate or broadly elliptic, sometimes
elliptic or subcircular, the largest mature leaves 1.6-4.2 cm long, 1.2-3 (3.8)
cm wide and 1-1.5-1.9 times as long as wide; apex round, obtuse or retuse,
more or less conduplicate when pressed; base round, obtuse or acute, some-
times inequilateral and more or less conduplicate when pressed; margins
entire or with several irregularly spaced glands or glandular teeth on lower
half; the upper side of mature leaves glabrous or pubescent near margin,
glossy and plane or subreticulate, the lower side sparsely pubescent to
glabrescent, glaucous; petioles 3-9-20 mm long, tawny or reddish tawny,
concave adaxially and glabrous or ciliate; stipules usually small glandular
lobes 0.2-0.4-1.2 mm long, rarely absent or narrowly elliptic. Aments
coetaneous, on leafy, floriferous branchlets. Staminate aments 1.2-2 cm long,
floriferous branchlets 0.4-1 cm long; stamens 2, filaments about 4.5-4.8
mm long, glabrous, distinct; anthers 0.5-0.6 mm long. Pistillate aments
1.5-3-3.7 cm long in fruit, floriferous branchlets 0.8-2.5-6 cm long; pistils
4-5.6 mm long, greenish reddish brown or greenish yellow, glabrous or
the beak sparsely pubescent, glossy, sometimes glaucous; styles (0.6) 0.8-1.4-
1.6 mm long; stigmas 2-4-lobed, 0.2-0.6 mm long; stipes 0.2-0.6-0.8 mm long,
73
Plate X
Salix stolonifera Cov. Argus 6772
74
Retusae Kern.
glabrous, pubescent if pistil is so; nectaries 1, adaxial, 0.5-1.2 mm long, (0.8)
1.5-3 times stipe; bracts broadly oblong, apex rounded, 1.6-2 mm long, brown
to dark brown, sparsely pubescent to glabrescent, adaxially more or less
pubescent with long straight or curly trichomes usually 2 times as long as
bract.
Habitat Range
Tundra: slide rock; moraine; sandy Alpine: Southern coastal Alaska from
lake margins; to sea level. King Cove, Alaska Peninsula to Ko-
diak Island; Cape Peirce to the Kenai
Peninsula; southeastern Alaska and
adjacent British Columbia (Map 18).
Discussion
Salix stolonifera is characterized by long styles, 0.8-1.6 mm long, glaucous
branches and buds, and a tendency for buried branches to produce yellowish
underground shoots. Coville (1901) referred to these shoots as ‘“‘stolons”.
In the original description of S. sto/onifera, he contrasted it with S. ovalifolia
emphasizing its characteristic “production of slender, leafless, subterranean
branches or stolons’, and the absence of glaucescence on the capsules. The
presence of subterraneous branches is more or less characteristic of the
species but this condition seems to occur only in unstable habitats with an
aggrading ground surface. Many specimens of S. stolonifera from southern
coastal Alaska lack subterranean branches and have long branches trailing
on the surface similar to S. ovalifolia. The absence of capsule glaucescence is
superficially distinctive but | have seen some specimens of S. stolonifera
which have glaucous pistils, and in S. ovalifolia about 30 per cent of the
specimens have non-glaucous pistils (Fig. 3). Clearly these taxa are closely
related and | have seriously considered combining the two as suggested by
Raup (1959); however, because of their distinctive eco-geographical distribu-
tion and in the absence of evidence of intergradation in the narrow zone of
overlap (Alaska Peninsula and Kodiak Island), | have decided to treat them
as species.
Some of the variation within Salix stolonifera may be attributable to hy-
bridization. | have collected specimens from two possible hybrid swarms
involving S. sto/onifera, S. arctica and S. barclayi. The hybrid swarm in the
Glacier Bay area is located on a wet, unstable clay-gravel morainal slope
near Goose Cove, Muir Inlet. The open vegetation cover is dominated by
Dryas drummondiana and Salix with patches of A/nus incana and Salix
arctica. Many of the Salix, including S. sto/onifera, S. barclayi, S. arctica,
S. sitchensis, S. commutata and S. reticulata, are prostrate or only up to
3 dm tall. In this area | collected a series of putative hybrids including S.
arctica < stolonifera and S. barclayi x stolonifera. The hybrids S. arctica
x stolonifera are distinguished by their more or less pubescent pistils,
narrowly obovate leaves which are often cuneate at the base, pubescent
beneath, and by their prominent stipules. The hybrids S. barclayi x stolo-
nifera are characterized by small leaves that are irregularly glandular ser-
rulate, some young leaves reddish green, and a prostrate habit. A second
75
Retusae Kern.
65° K
Map 18 Salix stolonifera
hybrid swarm is located on Mount Glave in northwestern British Columbia
on the Haines Highway between Alaska and the Yukon. The habitat is a
partly stabilized scree slope and adjacent alpine slopes are vegetated with
Empetrum nigrum, Phyllodoce, Potentilla fruticosa, Salix stolonifera, S. retic-
ulata, S. arctica and scattered S. planifolia ssp. pulchra thickets. In this
area there are some very robust specimens which | have tentatively deter-
mined as S. arctica X stolonifera. Salix stolonifera requires study first, to
determine its relationship with S. ovalifolia and secondly, to assess its
capacity to hybridize with other species in the mountains of southeastern
Alaska.
In 1969 Hultén stated that the occurrence of Salix stolonifera in the Kodiak
Island refugium (NNW of Amara Lake) contributes to the tendency toward
endemism which occurs in that area, and that this species may have survived
the Wisconsin glaciation in that refugium. It is probable that S. sto/onifera
did survive the glaciation in the Pacific coastal region, but its present distri-
bution suggests that it may have survived in more than one refugium — at
least on the Kenai Peninsula (Heusser 1960) and in the region south of
Yakutat Bay. Its apparent absence between Yakutat Bay and the Kenai
Peninsula may be a collecting hiatus. However, if it is real, it would support
the hypothesis that this species survived the glaciation in more than one
refugium.
76
Retusae Kern.
The report of an isolated occurrence of S. sto/onifera in central Alaska
(Hultén 1968, Map 19) is doubtful, and Hultén’s material should be re-
examined.
13. SALIX SETCHELLIANA Ball
S. setchelliana Ball, Univ. Calif. Publ. Bot. 17: 410. 1934. (Type: Setchell
567, US):
S. aliena Flod. Sv. Vet. Akad. Ark. Bot. 27A: 1. 1935.
Description of species
Prostrate or semiprostrate shrubs up to 25 cm tall; branches grey brown
to reddish brown, bark loose and coriaceous, often grey lanate; branchlets
reddish, densely white lanate becoming grey lanate or glabrate, rarely red-
dish and glabrous from start. Leaves coriaceous, narrowly obovate, the largest
mature leaves 2.5-6.6 cm long, 1-2 cm wide and 2-3.9 times as long as
wide; apex round to obtuse or acute; base tapering into the short petiole or
sometimes acute; margins entire to glandular serrulate or irregular glandular
crenate; the upper side of mature leaves lemon green, glabrous, venation
impressed, the secondary veins tapering toward base, the lower side glabrous,
glaucescent or pale yellow green; petioles short 0-3 mm long, glabrous, red-
dish, sometimes glaucous, rarely inflated around buds, the branchlet below
the petiole glabrous and appearing to be a part of it; stipules absent or small
glandular lobes about 0.3 mm long. Aments coetaneous, on leafy, floriferous
branchlets. Staminate aments 1.3-2 cm long, floriferous branchlets 0.7-1.3
cm long; stamens 2, filaments 2.8-3.2 mm long, glabrous or sparsely villous
at base; anthers 0.6-0.8 mm long, slender; nectaries 2, abaxial and adaxial,
about 0.6 mm long; bracts 2-2.8 mm long, greenish to golden brown,
glabrous. Pistillate aments 4-20-flowered, 1.5-2.5 cm long, floriferous branch-
lets 1-2 cm long; pistils 3.5-4.8 mm long, brick red, glabrous, capsules 3.6-
10 mm long, grey brown to brick red; styles 0.3-0.4 mm long, bifid to base;
stigmas 0.4-0.5 mm long; stipes 0-0.6 mm long, glabrous; nectaries 1,
adaxial, 0.8-1 mm long, equal to or up to 1.33 times as long as stipe; bracts
broadly obovate, apex rounded and erose or entire, 2-3.6 mm long, papery,
golden tawny, glabrous.
Habitat Range
Pioneer on sandy beaches, sandy- Montane: Alaska Range; Kuskokwim
gravel margins of glacial rivers, and River; upper Tanana River; Matanus-
on glacial moraine. ka River; Chitina River; southwestern
Yukon and adjacent coastal Alaska
(Map 19).
77
Plate XI
Salix setchelliana Ball. Anderson 8431 (top) and Murray 365 (bottom)
78
Retusae Kern.
170° 165° 160°
Map 19 Salix setchelliana
Discussion
Salix setchelliana is a highly distinctive willow growing on unstable morainal
and alluvial surfaces associated with glacial streams. It is characterized by a
prostrate habit, branches with leathery, loose, reddish bark, usually lanate
branchlets, and coriaceous, lemon green leaves on very short petioles. The
pistils are brick red and almost completely surrounded by the golden tawny
bracts.
It is not closely related to any North American species but does seem to
have some affinities with the European S. retusa L. and is included here in
section Retusae.
The isolated locality for S. setchelliana from the north slope of the Brooks
Range (Hultén 1968, Map 3) was probably based on Spetzman’s (1959)
report of this species from the Itkillik River. | was unsuccessful in obtain-
ing documentation for this report from the University of Minnesota Herbarium
and it is possibly based on a misidentification.
79
Plate XIl
Salix arctica Pall. (A) Arctic coastal form, Argus 5914, (B) Pacific coastal form, Argus 6484
80
Glaucae Pax
Section 7. Glaucae Pax
14. SALIX ARCTICA Pall.
S. arctica Pall. Fl. Ross. 1: 86. 1788.
S. arctica R. Br. Bot. Ross Voy. App. 143. 1819, nom. nud. S. arctica R.
Br. ex Richards. Bot. App. in Frankl. J. 752. 1823.
S. anglorum Cham. Linnaea 6: 541. 1831, attempt to validate S. arctica R.
Br., but type represents S. phlebophylla.
S. torulosa Trautv. Nouv. Mem. Soc. Nat. Mosc. 2: 309. 1832. S. arctica var.
torulosa Raup, Contr. Gray Herb. 185: 49. 1959. S. arctica ssp. torulosa
Hult. Ark. Bot. Il. 7: 38. 1967.
S. crassijulis Trautv. Nouv. Mem. Soc. Nat. Mosc. 2: 308. 1832. S. arctica
ssp. crassijulis A. Skvortsov, in Tolmatchev, Fl. Arct. URSS 5: 59. 1966. S.
arctica ssp. crassijulis Hult. Ark. Bot. Il. 7: 38. 1967.
S. anglorum var. kophophylla Schneid. Bot. Gaz. 66: 130. 1918. S. arctica
var. kophophylla Polunin, J. Bot. 77: 271. 1931.
S. anglorum var. araioclada Schneid. Bot. Gaz. 66: 133. 1918. S. arctica var.
araioclada Raup, Sargentia 4: 100. 1943.
S. anglorum var. antiplasta Schneid. Bot. Gaz. 66: 134. 1918. S. arctica var.
antiplasta Fern. Rhodora 48: 44. 1946.
Description of species
Dwarf shrubs, usually prostrate or trailing but sometimes up to 3-5 dm tell in
protected habitats; branches stout or long, slender and appressed, sometimes
rooting, chestnut brown to dark brown, glabrescent and sometimes glaucous;
branchlets yellow green to chestnut brown, glossy, glabrous or sparsely
pubescent with straggly trichomes, sometimes densely villous at first but
soon becoming glabrescent, sometimes glaucous. Leaves narrowly obovate to
elliptic (Arctic Ocean populations), subcircular to broadly elliptic or narrowly
obovate (Bering Sea populations), obovate to narrowly obovate or broadly
elliptic (Pacific Coast and interior populations), the largest mature leaves
1.9-4.5-7.6 cm long (Arctic Ocean and interior populations), 2.5-5-8.5 cm long
(Bering Sea and Pacific Coast populations), 0.7-1.8-3.4 cm wide (Arctic
Ocean and interior populations), 1.9-3-6 cm wide (Bering Sea and Pacific
Coast populations) and 1.4-2.4-3.5 times as long as wide (Arctic Ocean,
interior and Pacific Coast populations) or 1.1-1.5-2.3 times as long as wide
(Bering Sea populations); apex obtuse, round or acute in narrow leaves,
obtuse and conduplicate to round or retuse in broad leaves; base narrowly to
broadly cuneate or round in the broadest leaves; margins entire, slightly
revolute, often glandular dotted on the lower quarter of the blade; the upper
side of immature leaves glabrous or sparsely pubescent, the lower side
81
Glaucae Pax
pubescent with long straight trichomes oriented toward the apex; the upper
side of mature leaves usually glabrescent, indumentum may persist along
the margins and on the lower half of the midrib, glossy, the lower side
glabrescent, some long straight trichomes may persist near the apex pro-
ducing a ‘‘bearded” apex, glaucous; petioles (3) 9-15-35 mm long, the length
more or less correlated with leaf length, tawny or reddish (especially in some
Arctic Ocean and interior populations) , adaxially usually concave and glabrous
to sparsely pubescent; stipules absent, minute glands or linear and up to 7-10
mm long. Aments coetaneous, on leafy, floriferous branchlets. Staminate
aments 2.2-5 cm long, floriferous branchlets 0.9-5.5 cm long; stamens 2, fila-
ments 4-5 mm long, glabrous and distinct; anthers 0.4-0.6-0.9 mm long (the
longest in the Aleutian Islands and central Pacific Coast populations); nec-
taries 1, adaxial, sometimes also with a second, smaller abaxial nectary.
Pistillate aments 4-8-12 cm long, 1.5-5 cm long in Arctic Ocean populations,
floriferous branchlets (1) 3-8 (12) cm long; pistils reddish or tawny, densely
white pubescent, capsules 5.6-9 mm long, reddish or tawny, sometimes
streaked with red, sparsely pubescent or sometimes remaining densely pu-
bescent or becoming glabrescent, glossy; styles 0.6-2.2 mm long, entire or
bifid up to one third the length of style, red in life, drying purplish; stigmas
0.2-0.8 (1) mm long, 4 linear or 2 broad lobes; stipes 0.2-1.6 mm long, elon-
gating in age, pubescent; nectaries 1, adaxial, 0.4-1.8 mm long, usually 1.5-4
times as long as stipe, rarely equal to stipe; bracts broadly oblong, apex
rounded to obtuse or rarely acute, 1.6-2 mm long, uniformly brown or bicolour,
apex dark brown to black (light brown bracts common in Pacific Coast popu-
lations), sparsely pubescent on both sides with long straight trichomes 2-3
times as long as bract.
Habitat
Occurring in a wide variety of tundra
situations: Arctic: Carex aquatilis
meadows; Carex-Alopecurus mead-
ows; sandy Kobresia tundra; E/lymus-
Deschampsia vegetation on _ beach
ridges; Empetrum-lichen heath; snow-
bed vegetation; polygonal tundra. Al-
pine: glacial moraine; talus slopes;
Range
Arctic, alpine and subalpine: Through-
out arctic and alpine Alaska and
Yukon Territory; absent from the
forests in the Yukon-Tanana River
region and coastal Alaska; extending
eastward to Greenland; southward
in the cordillera to New Mexico; cir-
cumpolar (Map 20).
Salix-Phyllodoce tundra; Dryas tun-
dra; subalpine shrubby tundra.
Discussion
Salix arctica is characterized by a dwarf habit, elliptic to narrowly obovate
or subcircular leaves, usually sparsely pubescent beneath with long straight
trichomes producing a “beard” at the apex, coetaneous flowering, aments
on prominent floriferous branchlets, pubescent pistils, and nectaries longer
than the stipes.
This species is a taxonomically difficult circumpolar taxon—morphologi-
cally polymorphic, nomenclaturally confused and seriously in need of mono-
graphic study. It is not within the scope of this paper to conduct the critical
82
Glaucae Pax
165° 160° 155° 125°
sss zs iy fs Ue ie ie
WY CUYD Be
eEce
he
Map 20 Salix arctica. Circles based on Skvortsov, 1966
studies required to resolve the many problems; therefore, | will refer the
reader to the pertinent literature and restrict myself to a few brief comments
on variation and nomenclature. The synonymy cited here includes only the
more important names used in the literature dealing with Alaska and the
Yukon.
In Alaska and the Yukon Hultén recognizes three major geographical varia-
tions. In 1943 he treated them, tentatively, as species, S. arctica Pall., S. cras-
sijulis and S. torulosa, and in 1917 he reduced them to the rank of subspecies.
Every author who has commented on the taxa within Salix arctica has noted
that, although the extremes are distinctive, they appear to be confluent and
that, for the most part, only trends are evident. Therefore, even the recogni-
tion of these taxa as subspecies presents problems.
One of the most distinctive variants, S. arctica ssp. crassijulis, has its
centre of occurrence in the Bering Sea, the Aleutian Islands and the Pacific
coastal regions. It is characterized by a tendency toward large, subcircular
leaves, although they may vary from broadly elliptic to narrowly obovate,
long petioles and large fruiting pistillate aments. In short, it is a robust
variant of the species with leaves tending to be somewhat broader. | have had
an opportunity to make and to study population collections of this form in
southeastern Alaska. In the Glacier Bay region a population collection was
83
Glaucae Pax
made on the moraine of the Casement Glacier. While collecting specimens
along transect lines, | noted that the robust plants were all associated
with A/nus thickets whereas the small plants occurred on the open
moraine. It has been shown by Crocker and Major (1955) that A/nus crispa
has the ability to fix nitrogen in the soil, probably through the association of
nitrogen-fixing bacteria with its roots, and that nitrogen accumulates under
alder at the rate of 62 kg/ha/per annum (55 lb./acre). Also, they showed
that the development of the soil nitrogen profile depends upon plant distribu-
tion, and it is possible to assume that there are higher nitrogen levels asso-
ciated with A/nus shrubs and lower levels in the open moraine. If this is so,
then it is possible that the more robust specimens of S. arctica are either
modified directly by the increased nitrogen levels or that such specimens are
able to grow only in nitrogen-rich soils. Several authors have suggested that
the smaller forms of the species are the product of more rigorous environ-
ments, but no one has proposed that the more vigorous forms may be the
product of more favourable environments. The possibility that the robust plants
may be the result of direct environmental modification certainly cannot be
ignored. Furthermore, the geographical integrity of this taxon is not as good
as it may first appear. Robust plants are most evident in western and southern
coastal Alaska but similar individuals occur in central Alaska. Porsild in 1951
reported this taxon from the Canol Road, Yukon, and Raup (1959) maintained
that var. arioclada in eastern North America could not be distinguished from
it. This preliminary evidence suggests that the taxon “‘crassijulis’’ may be
better recognized as a form rather than as a subspecies, but no new combi-
nation is proposed here.
The taxon S. torulosa, which is supposed to be the characteristic form of S.
arctica in central Alaska and the Yukon, is a very elusive one. It is character-
ized, according to Hultén (1943 and 1968), by light brown young twigs,
leaves narrower, acute, light green and more or less glabrescent in age,
styles 1-1.5 mm long and light brown, oblong or pointed bracts. In my ex-
perience it is virtually impossible to distinguish this taxon on the basis of
these characteristics, all of which occur outside central Alaska and the
Yukon in a wide variety of recombinations. For example, light brown bracts
are more common in Pacific coastal populations than in the central Alaska
populations and the variation in style length in central Alaska populations,
1-2 mm long, is exactly the same as occurs anywhere else. It is possible that
S. torulosa occurs in central Siberia and Mongolia (Skvortsov 1966) but |
doubt that it occurs in North America. It is my impression from examining a
large series of specimens that the central Alaska population of S. arctica,
rather than forming a distinct geographical entity, recombines most of the
characteristics of the arctic Alaska, Bering Coast and Pacific coastal popu-
lations.
In 1959 Raup recognized five ‘‘poorly defined” varieties of Salix arctica
in western America, var. arctica, var. kophophylla, var. araioclada, var.
antiplasta and var. torulosa. For a key to these taxa which, “will do little
more than indicate trends’ see Raup 1959, p. 50. This key is difficult to
use, reflecting the great and confluent variability within S. arctica.
84
Glaucae Pax
Those who wish to pursue this subject further will find the following ref-
erences required reading: Schneider 1918a; Hultén 1943, 1968; Ball 1950:
Raup 1959; Skvortsov 1966. There are a number of other, less important,
papers which are referred to by these authors. It is important to be aware
that none of these papers is monographic in its approach. In attempting to
solve problems as an adjunct to other, usually floristic, considerations,
the authors have often added as much to the confusion as to the elucida-
tion of the problems at hand.
A problem almost as difficult as recognizing the variation patterns within
S. arctica as taxa is to distinguish this species from its ‘‘relatives’’. In North
America and Eurasia S. arctica is related to S. glauca and S. sphenophylla
and in North America to S. brachycarpa. Salix arctophila, which many
American authors have related to S. arctica (see that species and Table 9),
is in reality related to S. chamissonis and only superficially resembles
S. arctica. The characteristics which can be used to distinguish S. arctica
from S. glauca and S. brachycarpa are few and often somewhat trivial
and variable. Nevertheless, these taxa are distinct species belonging to
the same section; Table 5 will aid in their identification. For a comparison
with S. sphenophylla see Table 6.
Hybridization involving S. arctica seems to occur very infrequently, if at all,
in Alaska and the Yukon. Skvortsov (1966) also comments that in arctic
U.S.S.R. hybrids involving this species are rare. In Flora of Alaska and
Yukon, Hultén (1943) recognized seven hybrid combinations involving S.
arctica s.l., most of which are very difficult to substantiate, and in 1968 he
cited ten taxa with which this species hybridizes. In my studies | have been
able to recognize three putative hybrids in our flora: 1) S. arctica x stoloni-
fera in the Glacier Bay and Haines regions of southeastern Alaska and ad-
jacent British Columbia, 2) S. arctica x ovalifolia in the Bering Sea regions,
and 3) S. arctica x glauca in northern Yukon (Map 1). All of these hybrids
are putative, and until the variation within the species is better understood,
it is very difficult to recognize hybridization.
The somatic chromosome number data for Salix arctica seem to suggest
the presence of two chromosomal races (Table 1): tetraploid in the eastern
American arctic and hexaploid in Asia and the western American arctic
(Suda and Argus 1969). The pattern is disturbed by the report of a tetraploid
count from Ogotoruk Creek, Alaska (Johnson and Packer 1968), but further
cytological studies may reveal a differentiation pattern that could aid in the
solution of the taxonomic problems in this very difficult species.
15. SALIX SPHENOPHYLLA Skvortsov
S. sphenophylla A. Skvortsov, in Tolmatchev, Fl. Arct. URSS 5: 62. 1966.
S. cuneata Turcz. in Ledeb. Fl. Ross. 3: 623. 1850, non Nutt. 1842.
S. sphenophylla ssp. pseudotorulosa A. Skvortsov, in Tolmatchev, FI. Arct.
URSS' 5: 63. 1966.
85
Plate XIill
Salix sphenophylla Skvortsov, Argus 5912
86
Glaucae Pax
Table 5 Comparison table: Salix arctica, S. glauca and S. brachycarpa
Characteristics arctica glauca brachycarpa
chromosome number 2n 76, 114 2n 76,95, 114 2n 38
habit dwarf shrubs up to erect shrubs 0.3-0.9 erect shrubs 0.3-0.9
3-5 dm tall, pros- (4.5) m tall (2) m tall, sometimes
trate or trailing prostrate
style length 0.6-2.2 mm 0.5-1 mm 0.2-0.5 or 0.5-0.8
(1.5) mm
style colour red in life greenish in life greenish in life
leaf pubescence lower side pubescent lower surface villous lower surface dense-
with long, straight tri- to villous-sericeous ly matted greyish
chomes or glabrescent, or glabrescent with white trichomes to
a‘‘beard” remaining at — short trichomes glabrescent or
apex sparsely lanate or
villous-lanate
bract colour usually black, some- light brown to tawny light brown to dark
times brownish to light brown or blackish
brown
bract pubescence long, straight trichomes, short, wavy trichomes _ short, wavy or
2-3 times length of bract straight trichomes
bract shape broadly oblong, apex ovate to narrowly elliptic to broadly or
rounded to obtuse, elliptic, apex acute to narrowly so, apex
rarely acute obtuse acute to obtuse
Table 6 Comparison table: Salix sphenophylla, S. arctica and S. ovalifolia
Characteristics sphenophylla arctica ovalifolia
branches trailing and rooting trailing and rooting trailing but not
or erect rooting
leaf base mostly cuneate cuneate to rounded round to subcordate
to acute or acute
leaf surface dull above usually glossy above glossy above
petiole length 4-14 (15) mm (3) 9-15-35 mm 2-4-7 mm
pistillate floriferous 2.5-4.5 (5) cm 3-8 (12) cm 0.5-1.5-2.5 cm
branchlet length
pistillate ament 2-4.5 (6) cm 4-8-12 cm 0.9-1.5-2.8 cm
length
pistil pubescence glabrous or sparsely densely white mostly glabrous
pubescent on beak pubescent and glaucous
style length 0.6-1.8 mm 0.6-2.2 mm 0.2-0.8 mm
Description of species
Dwarf shrubs, branches brownish, trailing and rooting, glaucous; branchlets
more or less ascending, greenish brown, glabrous, bud scale often persistent
at base. Leaves elliptic to narrowly or broadly so, the largest mature leaves
1.9-4.8 (6) cm long, 1-2 cm wide and 1.9-3 times as long as wide; apex
87
Glaucae Pax
acute to obtuse or rounded; base cuneate to acute (rarely rounded); margins
entire, sometimes ciliate; the upper side of mature leaves light green,
glabrous and dull, the lower side glabrous or sparsely pubescent with long,
caducous trichomes, glaucous, venation prominent; petioles 4-14 (15) mm
long, stipules minute lobes to 0.8 or sometimes 5.5 mm long. Aments
coetaneous, on leafy, floriferous branchlets. Staminate aments (unknown in
Alaska and the Yukon, description based on Siberian specimens) ca. 2.7-3
cm long, floriferous branchlets ca. 2.2-2.3 cm long, stamens 2, filaments ca.
45 mm long, glabrous, distinct; anthers (0.4) 0.5-0.6 mm long. Pistillate
aments 2-4.5 (6) cm long, floriferous branchlets 2.5-4.5 (5) cm long, sim-
ilar to vegetative shoots; pistils ca. 4 mm long, greenish, suffused with red,
glabrous or sparsely pubescent on the beak, capsules ca. 5-6 mm _ long,
glabrous or sparsely pubescent; styles 0.6-1.8 mm long, entire or partly
bifurcate; stigmas 0.4-0.6 mm long, four linear lobes, stipes 0.5-1.4 mm
long, glabrous or pubescent, nectaries 1, adaxial, 0.8-1.6 mm long, equal
to or up to 2 times stipe; bracts narrowly elliptic, dark brown to blackish,
sparsely pubescent on both sides, ca. 1.6-2 mm long.
Habitat Range
Polygonal tundra; Carex aquatilis Arctic: Little Diomede Island; Seward
fens; dry, rocky slopes covered with Peninsula; Barter Island; Nuvagapak
Dryas. Point; Cape Dalhousie, Northwest
Territories; Asia: Lake Baikal; Lena
River; eastward to Kamchatka and
the Chukotsk Peninsula (Map 21).
Discussion
Salix sphenophylla seems to be related to S. ovalifolia and S. arctica.
Skvortsov (1966) claims that it is not too closely related to S. arctica
but belongs to a group of Asian species including S. nakamurana Koidz. and
S. kurilensis Koidz. (S. longipetiolata Flod.). However, he does include it
in the same section as S. arctica and it is similar enough morphologically
so that he has described some specimens called S. torulosa by Hultén
(1943), included as part of S. arctica in this treatment, as S. sphenophylla
ssp. pseudotorulosa. According to Skvortsov, subspecies pseudotorulosa
occurs in eastern Siberia and Alaska and is characterized by very sparsely
pubescent pistils in contrast to the completely glabrous pistils of the species.
The Alaskan specimens that | have determined as S. sphenophylla vary in this
characteristic from specimens with completely glabrous pistils, to those with
sparsely pubescent pistils (Argus & Chunys 5912), to one in which most
pistils are glabrous but a few are very sparsely pubescent (Argus & Chunys
5848). In view of this almost continuous variation, | have decided not to
recognize the Alaska material as the subspecies. Salix sphenophylla can be
distinguished from S. ovalifolia by a large number of characteristics (Table
6), but from S. arctica it is distinguishable only on the basis of its glabrous
or sparsely pubescent pistils, and its dull leaves. Other characteristics which
Skvortsov uses in his key, such as light yellow branches, the cuneate leaf
88
Glaucae Pax
==--~
165 155 : -
175 180
Map 21 Salix sphenophylla. Circles based on Skvortsov, 1966
base, the supposed absence of stipules and the shorter styles are too vari-
able within these taxa to be of diagnostic value. It is with some hesitation
that | recognize S. sphenophylla in the North American flora, and a case
could be made for considering it either as an extreme variant of S. arctica
or perhaps as the hybrid S. arctica X ovalifolia. However, it is relatively
distinctive and | include it here with the hope that it will stimulate further
consideration of this eastern Asian species in North America.
16a. SALIX BRACHYCARPA Nutt. ssp. BRACHYCARPA
S. brachycarpa Nutt. N. Am. Sylva 1: 69. 1842. (Type col.: Nuttal/ s.n. GH).
S. desertorum B (var.) stricta Anderss. in DC. Prod. 16 (2): 281. 1868.
(Type: Bourgeau s.n. photo and frag. A). S. stricta Rydb. Bull. N.Y. Bot.
‘Gard. 1: 273. 1899.
S. desertorum y (var.) fruticulosa Anderss. in DC. Prod. 16 (2): 281. 1868.
S. brachycarpa var. glabellicarpa Schneid. Bot. Gaz. 66: 338. 1918. (Type:
Macoun 95374, CAN).
89
Plate XIV
Salix brachycarpa Nutt. (A) Subspecies brachycarpa, Argus 6877; (B, C) Subspecies nipho-
clada (Rydb.) Argus: Argus 5276 (B), and Argus 5276 (C)
90
Glaucae Pax
S. brachycarpa var. sansonii Ball, Univ. Calif. Publ. Bot. 17: 414. 1934.
(Type: Sanson 119, Banff Nat. Pk.).
S. brachycarpa var. psammophila Raup, J. Arnold Arb. 17: 230. 1936.
(Type: Raup 6888, GH).
Description of species
Shrubs erect, commonly 0.3-0.9 m tall, some up to 2-3 m tall; branches
stout, reddish brown, pubescent with persistent, greyish indumentum, some-
times glabrescent; branchlets densely white or grey villous-lanate, sometimes
sparsely pubescent with coarse trichomes. Leaves obovate to broadly or
narrowly elliptic to narrowly obovate, the largest mature leaves (1.2) 2.3-3
(4) cm long, 6-9-16 mm wide, (1.5) 2.8-3 (4) times as long as wide;
apex acute to obtuse; base rounded; margins entire; mature leaves pubescent
on both sides with densely matted greyish white trichomes, sometimes only
sparsely pubescent, glaucous beneath; petioles (0.5) 1-3 (4) mm long,
usually shorter than bud, often reddish; stipules less than 0.5-1.5 mm long,
broad, often obscured by the dense branchlet pubescence. Aments coeta-
neous, on leafy, floriferous branchlets. Staminate aments about 6-15 mm long,
short-cylindrical to globose; floriferous branchlets rarely more than 10 mm
long; stamens 2; filaments distinct or sometimes united at base, glabrous;
anthers about 0.4-0.5 mm long; nectaries 2, abaxial and adaxial. Pistil-
late aments 15-20 mm long, short-cylindrical to nearly globose; floriferous
branchlets 3-20 mm long; pistils densely white lanate, short-beaked, capsules
less pubescent, light brown; styles 0.5-0.8 (1.5) mm long; stigmas 2, 0.2-
0.3 mm long, each two-lobed; stipes usually absent or up to 0.25-0.5 mm
long, pubescent; nectaries 1, adaxial, often 0.5 times as long as pistil; bracts
elliptic to broadly so, light brown, often greenish early in development,
pubescent on both sides with straight or curly trichomes.
Habitat Range
Alpine slopes; unstable limestone Montane, boreal: Southeastern Yu-
scree; Betula glandulosa-Carex-Salix kon; adjacent British Columbia and
fens; margins of alkaline or marly Northwest Territories; southward in
ponds and fens; unstable gravel mar- the cordillera to Utah and Colorado;
gins of streams and lakes. eastward in Subarctic Canada to Hud-
son Bay, James Bay, Ungava, and the
Gaspé Peninsula; across southern AIl-
berta and Saskatchewan (Map 22).
16b. SALIX BRACHYCARPA ssp. NIPHOCLADA (Rydb.) Argus
S. brachycarpa ssp. niphoclada Argus, Contr. Gray Herb. 196: 119. 1965.
S. niphoclada Rydb. Bull. N.Y. Bot. Gard. 1: 272. 1899. (Type: Taylor
91
Glaucae Pax
135° 130°
Map 22 Salix brachycarpa ssp. brachycarpa
60, CAN). S. glauca ssp. niphoclada Wiggins, in Wiggins and Thomas,
Fl. Alaskan Arct. Slope 144. 1962.
?S. lingulata Anderss. in DC. Prodr. 16 (2): 281. 1868. (Type col.:
Kastalsky s.n. NY).
S. brachycarpa ssp. mexiae Ball, Univ. Calif. Publ. Bot. 17: 412. 1934.
(Type: Mexia 2131, A). S. niphoclada var. mexiae Hult. Ark. Bot. iy ie Eo
1967.
S. muriei Hult. Fl. Alaska and Yukon 3: 531. 1943. (Type: Murie s.n. A).
S. niphoclada var. muriei Raup, Contr. Gray Herb. 185: 60. 1959.
Description of subspecies
Differs from ssp. brachycarpa in low shrubs, erect to prostrate, often
spreading, 0.3-1 (2) m tall: branches thin and flexible, reddish brown,
greyish or yellowish brown, sometimes with persistent pubescence. The
largest mature leaves (2.1) 2.5-3.2 (4.5) cm long, 7-11-15 mm wide and
(2.1) 2.9-3.7 (5.5) times as long as wide; apex acute-attenuate: base
cuneate to rounded; margins entire or distantly glandular: immature leaves
sericeous with appressed trichomes, the upper side of mature leaves sparsely
92
Glaucae Pax
pubescent to glabrescent, the lower side sparsely lanate or villous-lanate,
glaucous, the margin sometimes ciliate; petioles reddish to yellowish; stipules
2 mm or less up to 4 mm long, sometimes prominent, narrowly ovate,
glandular on margin, pubescent. Staminate aments 1.6-3.7 cm long, narrowly
cylindrical; anthers about 0.3-0.5 mm long. Pistillate aments 2.3.5-5 cm
long; slender, loosely flowered, capsules sparsely pubescent, light brown
sometimes greenish; styles 0.2-0.5 mm long, entire or bifurcate; nectaries
0.5-1.5-2 mm long, sometimes with lobes laterally as well as adaxially,
often longer than pistil; bracts elliptic to narrowly elliptic or oval, apex
rounded or emarginate, 1.5-2.8 mm long, tawny, greenish tawny or some-
times dark brown to blackish, pubescent on both sides with short trichomes
or glabrescent abaxially.
Habitat
Arctic: tundra; Salix thickets on
stream margins and in sandy blow-
outs. Montane: dry alpine slopes; un-
stable limestone talus; glacial out-
wash; subalpine shrubby — tundra.
Boreal: early successional stages on
Range
Arctic, montane, boreal: Central
Alaska; Alaska Range; arctic Alaska;
Alaska Peninsula; throughout Yukon
Territory to adjacent British Colum-
bia; Northwest Territories to Hudson
Bay (Map 23).
alluvial deposits and on margins of
semi-saline prairies.
Discussion
Salix brachycarpa is characterized by a low shrubby habit, very short
petioles, 0.5-1-3 (4) mm long, subglobose (ssp. brachycarpa) or narrowly
cylindrical (ssp. niphoclada) aments, densely white lanate, almost sessile,
pistils, the stipes 0-0.5 mm long, and small anthers, 0.3-0.5 mm long.
This species is closely related to S. glauca and S. arctica from which it
may be distinguished by the characteristics in Tables 4 and 6. It is also re-
lated to the Eurasian S. reptans Rupr. and may eventually prove to be con-
specific with it. | have seen specimens of the latter species from eastern
Siberia (Gyda River and Yakutskaya) which have the short, subglobose
aments of typical ssp. brachycarpa and others with the slender aments of
ssp. niphoclada. However, there is wide variation within the specimens that
| have seen and | am unable at the present time to formulate a clear
concept of S. reptans. It is possible that the broad-leaved specimen that | have
named ssp. niphoclada, such as: Johnson, et al. 215 (Argus 1965a, Figs. 54
and 55), 673 & 215A, from Ogotoruk Creek; Harbo 42, from Cold Bay;
Schofield 2274, from Naknek; and Scott 2023, from the Wrangell Moun-
tains, may be referred to that species. For discussion of the broad-leaved
variant of ssp. niphoclada, see Argus 1965a.
In 1965 | discussed the historical development of the concepts of the
relationships of niphoclada and presented reasons for combining it with S.
brachycarpa rather than with S. glauca as has been proposed by Hultén
(1943), Raup (1959) and Wiggins (Wiggins and Thomas 1962). | pointed out
that niphoclada differed from S. glauca in a number of important char-
acteristics including stipe length, style length, petiole length, anther length,
93
Glaucae Pax
Map 23 Salix brachycarpa ssp. niphoclada
the shape of the proximal leaves on vegetative shoots and probably also in
chromosome number (based on stomata size). Chromosome counts from
Cape Thompson, Alaska, indicate that ssp. niphoclada is diploid (2n = ca. 38,
Johnson and Packer 1968), supporting the indirect stomatal length data
(Argus 1965a). It was also noted that it was very similar to S. brachycarpa
in all of these characteristics and that the two taxa seem to intergrade in
southern Yukon and northern British Columbia. Since that time | have been
able to collect additional specimens along the Alaska Highway in northern
British Columbia and | am convinced that there is a continuous intergrada-
tion between these taxa in that area. Ssp. brachycarpa differs from ssp. ni-
phoclada in ament shape, longer styles (Table 7) and in its coarsely
pubescent leaves.
The apparent intergradation that seems to exist between S. glauca and
S. brachycarpa ssp. niphoclada may be due to hybridization, which was
suspected to be relatively common in arctic Alaska and in the Alaska Range
(Map 1) as discussed by Argus (1965a), and to environmental modification,
which may produce robust forms of ssp. niphoclada resembling S. glauca.
Hybridization between these taxa needs to be verified.
94
Glaucae Pax
Table 7 Comparison table: Salix brachycarpa ssp. brachycarpa, ssp. niphoclada and S. glauca
Characteristics brachycarpa brachycarpa
ssp. brachycarpa ssp. niphoclada glauca
chromosome number 2n=38 2n=38 2n= (76, 95), 114
petiole length (0.5) 1-3 (4) mm 1-3-4 mm 2.5-5-15 mm
stipe length absent (0.25-0.5) mm (absent) 0.25-0.5 mm 0.25-1-2 mm
anther length ca. 0.4-0.5 mm 0.3-0.5 mm 0.5-0.8 mm
proximal leaves strap-shaped, apex strap-shaped, apex obovate to oblong,
on vegetative rounded to obtuse rounded to obtuse apex acute to
shoots obtuse
ament shape short-cylindrical to sub- narrowly cylindrical, cylindrical,
globose, densely flowered loosely flowered densely flowered
style length 0.5-0.8 (1.5) mm 0.2-0.5 mm 0.5-1 mm
17. SALIX GLAUCA L.*
S. glauca L. Sp. Pl. 1019. 1753.
S. desertorum Richards. Bot. App. in Frankl. J. 753. 1823. S. glauca (ssp.)
S. desertorum Anderss. Ofvers. Vet. Akad. Forh. (Stockh.) 15: 127. 1858.
S. glauca ssp. desertorum Hult. Fl. Alaska and Yukon 3: 527. 1943.
S. villosa Hook. Fl. Bor.-Am. 2: 144. 1838. S. glauca var. villosa Anderss.
Ofvers. Vet. Akad. Forh. (Stockh.) 15: 127. 1858. S. x glaucops a (var.)
villosa Anderss. in DC. Prodr. 16(2): 281. 1868.
S. villosa B (var.) acutifolia Hook. Fl. Bor.-Am. 2: 144. 1838. S. glauca
var. acutifolia Schneid. Bot. Gaz. 66: 327. 1918. S. glauca ssp. acutifolia
Hult. Ark. Bot. Il. 7: 40. 1967.
S. xX glaucops 8 (var). glabrescens Anderss. in DC. Prodr. 16 (2): 281.
1868. S. glauca var. glabrescens Schneid. Bot. Gaz. 66: 329. 1918.
S. glauca ssp. glabrescens Hult. Ark. Bot. Il. 7: 40. 1967.
S. glauca var. acutifolia f. poliophylla Schneid. Bot. Gaz. 67: 61. 1919.
S. glauca var. poliophylla Raup, J. Arnold Arb. 17: 233. 1936 (as poliophila).
S
. glauca var. alicea Ball, Univ. Calif. Publ. Bot. 17: 416. 1934.
S. glauca ssp. desertorum var. sericea Hult. Fl. Alaska and Yukon 3: 527.
1943.
S. glauca var. perstipula Raup, Sargentia 6: 154. 1947.
*See Argus 1965a for typification and further synonymy.
95
Plate XV
Salix glauca L. (A) Beringia phase (var. g/
AIX . glauca), Argus 6959, (B) Western phase (var. acu-
tifolia), Rouse 52, (C) Rocky Mountain phase (var. villosa), Argus 5276 : ;
96
Glaucae Pax
Description of species
Erect shrubs 0.3-0.9 (4.5) m tall, sometimes prostrate; branches reddish
brown to greyish, epidermis often exfoliating, glabrate or variously pubescent,
sometimes glaucous; branchlets pubescent to white villous, sometimes
glaucous. Leaves obovate, narrowly obovate or elliptic, the largest mature
leaves 2.4-6-10 cm long, 0.7-1.8-3.2 cm wide, 2-3-5.2 times as long as wide;
apex acute, acuminate or obtuse; base acute to acuminate or obtuse; margins
entire often glandular on lower portion of blade, sometimes revolute; the
upper side of mature leaves dark green, glossy, glabrescent or sometimes
villous, the lower side villous with short appressed trichomes to villous-
lanate or glabrescent; petioles 2.5-5-15 mm long, yellowish, pubescent or
glabrate; stipules narrowly ovate, apex attenuate to acute, minute to 1-7-17
mm long, margins glandular. Aments coetaneous, on leafy, floriferous branch-
lets. Staminate aments 1.6-2.5 cm long, floriferous branchlets 0.5-1.5 cm
long, persistent after aments fall; stamens 2; filaments distinct or united at
base, glabrous or pubescent near base; anthers 0.5-0.6-0.8 mm long, reddish
in preanthesis, becoming brown; nectaries 2, abaxial and adaxial. Pistillate
aments 2.2-4-7 cm long, cylindrical, floriferous branchlets 1-3.5 cm long;
pistils densely white lanate, capsules sparsely pubescent to glabrescent, light
brown, long-beaked; styles 0.5-1 mm long, entire or bifurcate; stigmas with
4 linear lobes sometimes as long as style; stipes 0.25-1-2 mm long, pubes-
cent; nectaries 1, adaxial 0.6-1.2 mm long, 0.4-2 times as long as stipe;
bracts ovate to narrowly elliptic, light brown to tawny, pubescent both sides
with short wavy trichomes, sometimes glabrate abaxially.
Salix glauca s.|. ranges in Eurasia from arctic Scandinavia to the Chukotsk
and Kamchatka Peninsulas, U.S.S.R.
Beringia Phase (var. g/auca)
Habitat
Arctic tundra; commonly forming
thickets on the edges of creeks and
rivers, On sandy and gravel flood-
plains, and on old beaches.
Western Phase (var. acutifolia)
Habitat
Common in Picea mariana muskegs,
along drainage channels in P. glauca
woods, and on river floodplains; sub-
arctic Salix-Betula glandulosa thick-
ets; alpine tundra.
Range
Arctic, alpine: Western Alaska Penin-
sula; Kodiak Island; Lake Illiamna;
east Cook Inlet; northward to the
Kuskokwin River; along the Bering
Sea Coast to the Seward Peninsula;
eastward along the Arctic Coast and
the Brooks Range (Map 24).
Range
Boreal, montane: Central Alaska;
Alaska Range; Yukon Territory; east-
ward to Great Bear and Great Slave
Lakes; southward into northern Brit-
ish Columbia (Map 25).
97
Glaucae Pax
KF ck Ss
Ths Ses
J
S
< aio
< a
ASS
Map 24 Salix glauca, Beringia phase (var. glauca). Circles based on Skvortsov, 1966, as
S. glauca
Rocky Mountain Phase (var. vi//osa)
Habitat Range
Along rivers and creeks; openings in Montane, Boreal: Rocky Mountains
spruce woods; thickets on subalpine from northern British Columbia and
slopes. adjacent Yukon Territory to Utah and
New Mexico; eastward to Hudson Bay
(Map 26).
Discussion
Salix glauca is a highly variable circumpolar taxon which, although subject
to repeated study (Schneider 19186; Hultén 1943; Raup 1959; and Argus
1965a), is not completely understood. In 1965 | published a study of this
complex in which | concluded that it was advisable to treat the species in
a broad sense and to recognize four geographical variants in North America.
The decision to refer to these geographical variants by informal phase
names was based on three considerations. First, the taxonomic concept
of the geographical variants was based more on geography than on clearly
defined morphological differences; secondly, extensive intergradation oc-
98
Glaucae Pax
165°
170° 165° 160°
Map 25 Salix glauca, Western phase (var. acutifolia)
curred in the areas of overlap of the ranges of the variants; and thirdly, speci-
mens which morphologically resemble each of the variants occurred within the
range of the other variants. The geographical variation of each of the phases
was described and problems which required special study were noted. Since
that time | have been involved in additional study; chromosome numbers have
been determined for the species in Alberta and Alaska (Suda and Argus
1968, 1969) and a study of S. athabascensis in Saskatchewan and the Yukon
has resulted in its separation from S. glauca but in its retention within the
S. glauca-complex (see S. athabascensis for discussion). However, this infor-
mation has not yet substantially aided in the solution of the problems of
classification, and the variation within the species and its relationships
remain obscure.
Skvortsov (1966) has independently arrived at a similar conclusion concern-
ing this species in arctic U.S.S.R. He notes that there is some geographical
correlation of characteristics in populations in Altai, Sayan, and Khanga
which usually bear small leaves with sparse pubescence, lack stipules, and
have short aments and floriferous branchlets. Whereas, in parts of the
Lena, Indigira and Kolyma Rivers the leaves are larger, more strongly
pubescent and have well developed stipules and longer aments. However, he
does not attempt to recognize these variants nomenclaturally. Within the cir-
99
Glaucae Pax
IROD
Map 26 Salix glauca, Rocky Mountain phase (var. villosa)
cumpolar extent of the species he suggests that there are four ‘‘races’’, one
in eastern North America and Greenland, a second in Eurasia and western
North America (west of Hudson Bay), a third beginning to be differentiated
in southern Siberia and a fourth in the Alps. He ascribes formal nomenclatural
recognition only to the latter which he recognizes as the species S. glauco-
sericea Floderus.
In Alaska and the Yukon | recognize three geographical phases: the
Beringia phase, which is very similar to the Scandinavian S. glauca and may
be named S. glauca L. var. glauca; the Western phase, which may be named
S. glauca var. acutifolia (Anderss.) Schneider; and the Rocky Mountain phase,
which may be named S. glauca var. villosa (Hook.) Andersson. For those who
may wish to recognize the geographical variants as informal units or as formal
taxa | have provided a key to the phases and maps depicting their approx-
imate distribution. The synonyms listed here include only important names
used in reference to the flora of Alaska and the Yukon; for a more complete
synonymy see Argus 1965a.
Key to the major variants of Salix glauca in Alaska and the Yukon.
1a. Shrubs (0.3) 0.9-1.2 (3) m tall; leaves 2.4.4 (5) cm long, elliptic to
obovate, 1.6-3 times as long as wide; stipules minute to 2-6 mm long,
100
Glaucae Pax
generally inconspicuous; pistillate aments 2-4 cm long. Rocky Mountain
phase (var. villosa).
1b. Shrubs prostrate to 4.5 m tall; leaves 4-10 cm long, obovate to nar-
rowly obovate, 2.8-4 times as long as wide; stipules 4-10 (17) mm long,
prominent; pistillate aments 3.5-7 cm long.
2a. Shrubs 0.9-2.1 up to 4.5 m tall; leaves dark green and glabrescent
above, villous-lanate or sometimes glabrescent beneath; petioles 3-7-16
mm long; stipules very prominent; branchlets villous to pubescent;
pistillate aments stout, long-cylindrical; bracts light brown to tawny.
Western phase (var. acutifolia).
2b. Shrubs prostrate to 0.3-0.9 m tall; leaves light green above, pubescent
on both sides, becoming glabrescent above, never villous-lanate
beneath; petioles 2-5-10 mm long; stipules variable; branchlets dense-
ly villous, internodes usually short; pistillate aments shorter, narrowly
cylindrical; bracts often dark brown. Beringia phase (var. g/auca).
Salix glauca is characterized by pubescent branchlets and leaves, the usual
occurrence of prominent, narrowly elliptic stipules, densely white lanate
pistils borne on distinct stipes, and bracts usually tawny but sometimes
brownish. It is related to S. athabascensis, S. brachycarpa and S. arctica (see
those species for comments and Tables 5 and 7). It is also related to the
Eurasian S. reptans, and to the European S. glaucosericea Flod., S. pyrenaica
Gouan, and S. stipulifera Flod. ex Hayren.
The Beringia phase (var. glauca) is characterized by densely villous branch-
lets with short internodes, leaves villous to pubescent on both sides, short
petioles and stipules usually exceeding the petioles, densely lanate pistils on
short stipes and bracts tending to be brownish rather than tawny.
The Western phase (var. acutifolia) is characterized by large leaves often
with a distinctive villous-lanate pubescence beneath and glabrescent to
sparsely pubescent above, long petioles and prominent narrowly elliptic
stipules. The aments are large and the pistils are borne on prominent stipes.
There are some glabrescent forms which are included within this phase.
The Rocky Mountain phase (var. vi/losa) is characterized by a tendency
toward more glabrescent leaves, less prominent stipules and smaller leaves
and aments. It intergrades with the Western phase in southern Yukon and
northern British Columbia and individuals are often very difficult to assign
to one phase or the other on morphological grounds.
There are seven authentic chromosome counts for the North American
Salix glauca (Suda and Argus 1968, 1969; Johnson and Packer 1968). Three
of the counts based on specimens from Alberta representing the Rocky
Mountain phase are hexaploid (2n = 114); three counts based on specimens
from Umiat, Alaska, representing the Western phase, include tetraploid,
pentaploid and hexaploid levels (2n = 76, 95 and 114); and one count
representing the Beringia phase from Ogotoruk Creek, Alaska, is tetraploid.
This evidence suggests that the North American S. glauca is basically hexa-
ploid in contrast to the octoploid levels reported for European specimens
(Holmberg 1931; Wilkinson 1944; Léve and Love 1948). This evidence may
eventually provide a basis for segregating the North American and Euro-
pean members into two major taxa; but in view of the intrapopulation
101
Plate XVI
Salix athabascensis Raup. Population sample Argus 5054
102
Glaucae Pax
variation in chromosome number for the Western phase and the tetraploid
count for the Beringia phase, many more counts are needed before this
apparent difference can be evaluated.
Within Alaska and the Yukon Salix glauca is suspected to hybridize with
S. arctica and with S. brachycarpa ssp. niphoclada. |In arctic Alaska the latter
cross is reportedly common (Argus 1965a), but this conclusion was based on
a subjective analysis of morphological intermediacy and character recombina-
tion, and therefore requires corroboration from other kinds of data. It could
be speculated that the three ploidal levels reported for the Umiat material
are the result of hybridization and backcrossing with S. brachycarpa ssp. ni-
phoclada. |If the basic number of S. glauca is 2n = 114 then hybridization
with ssp. niphoclada, which is thought to be diploid (2n = 38), could yield F;
tetraploids and backcross pentaploids. However, the tetraploid and pentaploid
specimens (Suda 271-66, 274-66 and 277-66, SASK) show no morphological
signs of hybridization and do not support such a hypothesis.
18. SALIX ATHABASCENSIS Raup
S. athabascensis Raup, Rhodora 32: 111. 1930. (Type: Raup 8129, CAN).
S. pedicellaris Pursh var. athabascensis Boivin, Phytologia 15: 411. 1967.
S. glauca L. (ferruginous form) Argus, Contr. Gray Herb. 196: 85. 1965.
S. fallax Raup, Contr. Arnold Arb. 6: 149. 1934. (Type: Raup 4312, A).
Description of species
Shrubs 0.6-0.9-1.3 m tall; branches grey brown, epidermis exfoliating; branch-
lets reddish brown, glossy, densely or sparsely pubescent with short tri-
chomes curved toward the stem, becoming glabrescent. Leaves narrowly
elliptic to elliptic or narrowly obovate, the largest mature leaves 1.7-3.7-5
cm long, 0.8-1.3-1.8 cm wide, 2-3.2 times as long as wide; apex acute or
more or less acuminate; base round or acute, often inequilateral; margins
entire, often glandular on lower portion of blade or indistinctly and distantly
glandular serrulate, flat or slightly revolute; the upper side of immature
leaves usually villous or sericeous with mixed white and ferruginous tri-
chomes, often rugulose; the upper side of mature leaves glabrescent, ferru-
ginous tichomes sometimes persistent on midrib, glossy and reticulate, the
lower side glabrescent but with some appressed white and ferruginous tri-
chomes persistent for some time, glaucous; petioles 3-5-10 mm long, tawny
or reddish, villous or puberulent adaxially; stipules glandular lobes, minute
to 0.2-0.5 mm long. Aments coetaneous, on leafy, floriferous branchlets.
Staminate aments 5-9 mm long, floriferous branchlets 2-3 mm long, per-
sistent after aments fall; stamens 2, filaments distinct, pubescent on lower
half; anthers 0.4-0.6 mm long, nectaries 2, abaxial and adaxial. Pistillate
aments 1.2-3.7 cm long, loosely flowered, especially in fruit, floriferous
103
Glaucae Pax
165° 155° 130°
Map 27 Salix athabascensis
branchlets 0.2-1.5 cm long; pistils 1.8-2 mm long, greenish or tawny, densely
sericeous with white or sometimes with white and ferruginous trichomes,
capsules 5.6-7.2 mm long, tawny, reddish or greenish, sparsely sericeous
to glabrescent; styles 0.5-1 mm long, partly bifid; stigmas 0.2-0.5 mm long,
4 linear lobes; stipes 0.8-1.2 mm long, pubescent; nectaries 1, adaxial,
sometimes 2-lobed, 0.4-0.8 mm long, usually 0.5 times as long as stipe;
bracts ovate, apex rounded, 1-1.6 mm long, tawny, sometimes bicolour and
brown at apex, pubescent on both sides or glabrescent abaxially.
Habitat Range
Carex-Salix-Betula glandulosa fens; Boreal: Central Alaska along the
Picea mariana muskegs and bogs. Tanana River; southern Yukon and
adjacent British Columbia; Northwest
Territories to Hudson Bay (Map 27).
Discussion
Salix athabascensis is characterized by glabrescent, narrowly elliptic leaves
usually sparsely pubescent with appressed white and ferruginous trichomes,
loosely flowered aments, grey sericeous pistils sometimes with ferruginous
104
Glaucae Pax
Table 8 Comparison table: Salix athabascensis and S. pedicellaris
Characteristics athabascensis pedicellaris
pistils pubscent glabrous
style length 0.5-1 mm 0.1-0.2 mm
branchlet short, curved trichomes very minute, straight
pubescence trichomes
leaves sparsely sericeous with white glabrous
and furruginous trichomes
and white trichomes, long stipes (0.8-1.2 mm long) and short nectaries. It is
morphologically similar to S. pedicellaris from which it may be distinguished
by the characteristics in Table 8.
In 1965 | incorrectly treated this species as a ‘form’ of S. glauca (Argus
1965a) and attributed some sterile intermediates to hybridization between
S. glauca (or S. planifolia or S. discolor) and S. pedicellaris. My 1965
opinion concerning the relationship of S. athabascensis and S. glauca was
based on experience at Churchill, Manitoba. In this area S. glauca occurs in
the tundra on the outcrop ridge and in streamside Salix thickets, whereas
S. athabascensis occupies river floodplains, Picea mariana muskegs and
forests, and also occurs occasionally in the tundra and in Salix thickets.
The distinction between these taxa in central Canada is very tenuous and
is based on the sparser leaf, branchlet and pistil pubescence of S. athabas-
censis and the characteristic occurrence of ferruginous trichomes on its
leaves and sometimes on its pistils. Since that time new information has
been obtained leading to reconsideration of this taxon. | am now of the opin-
ion that it is best treated as a species related to S. glauca and hybridizing
with S. pedicellaris.
Hybridization between S. athabascensis and S. pedicellaris is suspected to
be relatively common in central Canada, and hybrid swarms have been studied
in Saskatchewan and southern Yukon (Map 2). Apparent F,’s, which are
usually infertile and have a high percentage of aborted and unfilled pol-
len grains, may be distinguished by their sparsely pubescent pistils (often
with patches or streaks of pubescence), undeveloped ovaries, short styles,
sparsely pubescent branchlets and subcoriaceous leaves. A preliminary com-
parative chromatographic analysis of leaf pigments (Argus, unpublished)
supports a hypothesis of hybridization between these species, and further
study is in progress.
Chromosome counts indicate that S. athabascensis and S. pedicellaris
are both tetraploid, 2n = 76 (Suda and Argus 1969; Love and Ritchie 1966)
and that hybrids between these taxa, although sterile, are also tetraploid
(Suda and Argus 1968). The three chromosome numbers reported for
S. glauca from Churchill, Manitoba (Argus 1965a) were based on speci-
mens of S. athabascensis growing on the floodplain of the Churchill River,
and should be referred to that species. The three specimens counted exhib-
ited three ploidal levels, 2n = ca. 76 (Argus 15S); 2n = 95 (Argus 517-58)
and 2n = ca. 114 (Argus 26S). The tetraploid level is now thought to repre-
sent the general ploidal level for this species; the pentaploid and hexaploid
105
Plate XVII
eee chamissonis Anderss. Suvorova s.n., 12 VIII 1962, Leninogorsk, Eastern Altai, U.S.S.R.,
106
Myrtosalix Kern.
levels may be the product of hybridization and backcrossing with the hexa-
ploid S. glauca, or of autoploidy within S. athabascensis.
Section 8. Myrtosalix Kern.
19. SALIX CHAMISSONIS Anderss.
S. chamissonis Anderss. in DC. Prod. 16 (2): 290. 1868.
S. rectijulis Ledeb. ex Trautv. Nouv. Mem. Soc. Nat. Mosc. 2: 313. 1832.
p.m.p. (quoad pl. ex. Ins. Sti. Laurentii).
Description of species
Prostrate, usually trailing shrubs; branches long, trailing on ground, aments
and some vegetative branchlets arising at right angles to branch, reddish
brown; branchlets yellow green, glabrous or sparsely pubescent with curly
trichomes, becoming glabrescent. Leaves obovate to elliptic-obovate, the
largest mature leaves 3-5 cm long, 1.7-3 cm wide and (1.1) 1.6-1.9 (2.1)
times as long as wide; apex obtuse to round or retuse, rarely acute, some-
times conduplicate when pressed; base cuneate; margins prominently and
uniformly glandular serrulate; the upper side of mature leaves glabrous and
semi-glossy, sometimes drying black, the lower side glabrous or with cadu-
cous trichomes, thinly glaucous; petioles 5-13 mm long; stipules narrowly
elliptic, 2.8-9 mm long, glandular serrulate. Aments coetaneous, on leafy,
floriferous branchlets. Staminate aments 2.2-3.2 cm long, floriferous branch-
lets 1.3-2.2 cm long; stamens 2, filaments about 7 mm long, glabrous; an-
thers 0.5-0.6 mm long, reddish; nectaries 1, adaxial, about 0.8 mm long. Pis-
tillate aments 3-6 cm long, rachis sparsely pubescent, branchlets 1.5-4 cm
long; pistils 2.4-4.8 mm long, greenish red with reddish sutures, pubescent
all over, rarely glabrous, pubescent in strips or in patches at the base and
apex, indumentum of flat, crinkled, refractive trichomes, capsules 5-7 mm
long, bronze or greenish bronze with red sutures, generally glabrescent but
beak remaining sparsely pubescent; styles 0.8-1.2 mm long, reddish, drying
dark purple, sometimes bifid; stigmas about 0.4 mm long; stipes 0.2-0.4 mm
long, pubescent; nectaries 1, adaxial, 0.3-0.6 mm long, dark red, equal to or
slightly exceeding stipe; bracts ovate, apex obtuse to rounded, 1.2-2.8 mm
long, black to dark brown, pubescent on both sides, trichomes 2 times length
of bract.
Habitat Range
Tundra. Arctic, alpine: Attu Island; St. Law-
rence Island; Seward Peninsula; Arc-
tic slope; Brooks Range; southward
to Wiseman and Twelvemile Summit;
eastward to Richardson Mountains,
northern Yukon, and adjacent North-
107
Plate XVIII
Salix arctophila Cock. ex Heller. Argus 142-58
108
Myrtosalix Kern.
west Territories; Asia: eastern Sibe-
ria along the northern shores of the
Sea of Okhotsk; Kamchatka Penin-
sula; Commanders Islands; Chukotsk
Peninsula (Map 28).
Discussion
Salix chamissonis is wide ranging but infrequent in arctic and alpine tundra.
It forms relatively large disjunct populations on Twelvemile Summit and
Eagle Summit and is only known to occur elsewhere in central Alaska in the
Kokrines Range.
This species is closely related to the European S. myrsinites L., the Eura-
sian S. saxatilis Turcz. (S. fumosa Turcz.) and the eastern American S. arc-
tophila Cock. All four species have the characteristic pistil pubescence con-
sisting of crinkled, refractive trichomes and more or less glandular serrulate
leaf margins. S. chamissonis differs from S. myrsinites in having obovate
leaves that are glaucous beneath, and shorter petioles. It differs from S.
arctophila in its prominently and uniformly glandular serrulate leaf and stip-
ule margins.
20. SALIX ARCTOPHILA Cock. ex Heller
S. arctophila Cock. ex Heller, Cat. N. Amer. Pl. (ed. 3) 89. 1910.
Description of species
Dwarf shrubs, decumbent and often trailing; branches long, slender and
chestnut brown or greenish brown; branchlets long, slender, usually yellow
green and glabrous, rarely slightly glaucous. Leaves broadly elliptic or obo-
vate, sometimes narrowly elliptic, the largest mature leaves 1.7-3.7 cm long,
0.7-1.6 cm wide, 1.3-1.8 (4.3) times as long as wide; apex obtuse, round or
sometimes acute; base obtuse, acute or cuneate; margins entire with minute
glands or minutely glandular serrulate, rarely eglandular; immature leaves
glabrous or very sparsely pubescent beneath; the upper side of mature leaves
glabrous, glossy and yellow green, the lower side glabrous, plane and glau-
cous; petioles 3-13 mm long, yellow green; stipules absent or 0.4-4 mm
long, glabrous, margins glandular. Aments coetaneous, on erect, leafy, flori-
ferous branchlets. Staminate aments about 2-5 cm long, floriferous branch-
lets 0.8-2 cm long; stamens 2, filaments 6-7 mm long, glabrous and purplish;
anthers 0.5-0.6 mm long. Pistillate aments 2.1-6 cm long, floriferous branch-
lets erect, 1.5-5.3 cm long; pistils 3-4 mm long, sparsely and uniformly
pubescent with short, flat, crinkled, refractive trichomes, reddish purple,
capsules 5-6 mm long, glossy; styles 0.6-1 mm long, usually entire but some-
times bifid, red in life, drying purple; stigmas 0.2-0.4 mm long, each bifid;
stipes 0.8-1.2 mm long, pubescent; nectaries 1, adaxial, 0.5-0.9 mm long,
109
Myrtosalix Kern.
Map 28 Salix chamissonis. Circles based on Skvortsov, 1966
0.5-0.66 times as long as stipe; bracts broadly oblong, apex rounded, 0.8-
1.6 mm long, distinct, usually purplish red or black, pubescent with long
straight trichomes.
Habitat Range
Wet tundra. Arctic: Arctic Ocean Coast from Bul-
len, Alaska, to the Mackenzie River
Delta; eastern Brooks Range; Yukon
Territory; Canadian Arctic; Green-
land; northern Saskatchewan; Chur-
chill, Manitoba (Map 29).
Discussion
Salix arctophila is an eastern Canadian Arctic species which has been infre-
quently collected in Alaska and the Yukon. It enters eastern Alaska and
extends westward to Bullen on the Arctic Coast and to Jago Lake in the
Brooks Range. The citation by Hultén (1943) of a specimen from Nome
(Jones 9042) is incorrect and was based on a misidentification. Most North
American authors (Raup 1959; Argus 1966) have related this species to S.
arctica, and Drury (1962) implied that they are conspecific. However, Skvort-
110
Myrtosalix Kern.
165° 160° ¥ 130° 125°
165° 160°
Map 29 Salix arctophila
Table 9 Comparison table: Salix arctophila and S. arctica
Characteristics arctophila arctica
branchlets slender, trailing, yellow usually stout and not
green and glabrous conspicuously trailing,
brownish, sparsely pubescent
leaf margins glandular serrulate entire or glandular
on the lower half dotted on the lower quarter
leaf pubescence glabrous pubescent, at least beneath,
with long, straight trichomes
pistil trichomes flat, crinkled and cylindrical, straight or
refractive curly, non-refractive
nectary length 0.5-0.66 times stipe 1.5-4 times stipe
sov's view (1966) that its relationship is with S. chamissonis and S. myrsi-
nites, not with S. arctica, is the most tenable and S. arctophila could con-
ceivably be treated as a geographical race of S. chamissonis. Salix arctophila
is morphologically similar to S. arctica, and vegetative specimens are partic-
ularly difficult to distinguish (Table 9).
111
Plate XIX
Salix fuscescens Anderss. Argus 5871 (left) and Welsh 8178 (right)
112
Myrtilloides Koehne
Section 9. Myrtilloides Koehne
21. SALIX FUSCESCENS Anderss.
S. fuscescens Anderss. Kg. Sv. Vet. Akad. Handl. 6: 97. 1867.
S. arbutifolia auct. non Pall. 1788.
S. fuscescens var. reducta Ball, Proc. Nat. Acad. Sci. 21: 183. 1935. (Type:
Coville & Kearney 20879, US).
Description of species
Low, trailing shrubs spreading from a central caudex; branches yellow brown
and rooting, aments arising at right angles; branchlets reddish brown, green-
ish brown or yellowish brown early in season, glabrous. Leaves obovate to
elliptic, the largest mature leaves (1.4) 1.7-2.7 (3.7) cm long, 0.7-2.1 cm
wide and (1.4) 1.5-2.3 (2.5) times as long as wide; apex round or obtuse,
rarely retuse; base obtuse to round, acute or rarely cuneate; margins entire,
glandular crenate or glandular serrulate on lower half, occasionally glan-
dular crenate all around, usually revolute; immature leaves glabrous, the
proximal leaves rarely sparsely pubescent with caducous, ferruginous tri-
chomes; the upper side of mature leaves glabrous, glossy and bright green,
the veins raised and prominent or sometimes pale green, the lower side
glabrous, glaucous or pale green; petioles 2-5.6 (6.4) mm long, pale reddish
yellow, stipules absent or minute. Aments coetaneous, on long, leafy, florif-
erous branchlets. Staminate aments 0.8-1.3 cm long, rachis pubescent, florif-
erous branchlets 0.3-2.2 cm long; stamens 2, filaments 3.5-4 mm _ long,
glabrous; anthers 0.3-0.4 mm long; nectaries 1, adaxial, 0.6-0.7 mm long,
yellowish. Pistillate aments 1.5-6 cm long, sparsely flowered at proximal end,
floriferous branchlets 1.0-4.5 cm long; pistils 4.4-5 mm long, long-beaked,
dark reddish brown, usually sparsely pubescent with short ferruginous tri-
chomes or densely white sericeous, rarely glabrescent, capsules 5.5-8 mm
long, tawny to reddish or greenish and streaked with red, often sparsely
pubescent with appressed, ferruginous trichomes, sometimes glabrescent;
styles 0.1-0.4 mm long, sometimes bifid; stigmas 0.2-0.4 mm long, 4-lobed;
stipes 0.8-2.5 mm long, pubescent or glabrescent; nectaries 1, adaxial, 0.4-
0.6 mm long, 0.5-0.66 times as long as stipe; bracts oblong, apex rounded,
0.8-1.6 mm long, bicolour, apex dark brown and base tawny becoming uni-
formly brown, rarely completely tawny, pubescent on both sides with tri-
chomes longer than bract.
Habitat Range
Trailing in moss in wet tundra includ- Arctic, boreal: Throughout Alaska,
ing Eriophorum-Arctagrostis tussock except the Aleutian Islands and the
tundra, Eriophorum-Carex meadows, southeastern coast of Alaska; north-
and polygonal tundra; also occurring ern half of Yukon Territory; North-
in dry tundra and in Picea mariana west Territories from the Mackenzie
muskegs. Delta; southward to northern Mani-
113
Plate XX
Salix pedicellaris Pursh. Argus 5054C
114
Myrtilloides Koehne
toba; Asia: eastern Asia from the
Lena River to Chukotsk Peninsula;
Kamchatka; the region around
Okhotsk (Skvortsov 1966, Map 22)
(Map 30).
Discussion
Salix fuscescens is characterized by glabrous, obovate leaves, usually nar-
rowed toward the base and with several glandular serrulations on the lower
half, glossy above and glaucous beneath; pistillate aments loosely florewed,
pistils dark reddish brown and sparsely pubescent with short ferruginous
trichomes. This species is related to S. pedicellaris and S. myrtilloides from
which it can be distinguished by its pubescent pistils and broader leaves.
The type of the name S. arbutifolia has been shown by Skvortsov (1957)
to be the genus Chosenia, not Salix, and the new combination Chosenia
arbutifolia (Pall.) Skvortsov was made. Although the type specimen is only
vegetative, Skvortsov maintains that the identification of it as Chosenia is
unequivocal and | have accepted his opinion. This, however, necessitates
replacing the name S. arbutifolia, which has been widely used in the North
American literature, with S. fuscescens.
22. SALIX PEDICELLARIS Pursh
S. pedicellaris Pursh, Fl. Am. Sept. 2: 611. 1814.
S. myrtilloides L. (ssp.) S. pedicellaris Anderss. Kg. Sv. Vet. Akad. Handl. 6:
96. 1867. S. myrtilloides B (var). pedicellaris Anderss. in DC. Prodr. 16 (2):
230. 1868.
S. pedicellaris var. hypoglauca Fern. Rhodora 11: 161. 1909. (Type: Forbes
s.n, GH).
S. pedicellaris var. tenuescens Fern. Rhodora 11: 162. 1909. (Type: Fernald
s.n. GH).
S. hebecarpa Fern. Rhodora 26: 123. 1924. S. fuscescens var. hebecarpa
Fern. Rhodora 9: 224. 1907. (Type: Fernald & Collins 207, GH).
Description of species
Shrubs 2-15 dm tall; branches grey brown, glabrous or glabrescent, epi-
dermis grey and exfoliating; branchlets reddish brown or reddish yellow,
minutely puberulent with short erect trichomes, sometimes glabrous; buds
glabrous. Leaves coriaceous, narrowly elliptic, narrowly rectangular or nar-
rowly obovate, the largest mature leaves 2.3-5.3 cm long, 0.7-1.3 cm wide
and 2.6-4.4 times as long as wide; apex acute to round; base acute to round,
sometimes inequilateral; margins entire, often glandular dotted or glandular
serrulate on lower quarter of blade, sometimes revolute; immature leaves
115
Myrtilloides Koehne
Map 30 Salix fuscescens. Circles based on Skvortsov, 1966
glabrous, usually green or reddish and translucent; the upper side of mature
leaves glabrous, dull, usually glaucescent, and with raised, reticulate vena-
tion, the lower side glabrous and glaucous; petioles 3-4-8 mm long, reddish
or reddish yellow, glabrous or sometimes puberulent adaxially; stipules are
minute glands or glandular lobes about 0.1-1.5 mm long. Aments coetaneous,
borne on short, leafy, floriferous branchlets. Staminate aments 0.9-1.5 cm
long, floriferous branchlets 3-8 mm long; stamens 2, filaments 3.2-3.6 mm
long, glabrous, distinct or connate at base; anthers 0.4-0.6 mm long. Pistil-
late aments 1.3-3 cm long, often as broad as long, loosely flowered, florif-
erous branchlets 1.5-3 cm long; pistils 2.4-4.8 mm long, reddish or tawny,
glabrous and often glaucous, capsules 5.6-6.4 mm long, tawny; styles 0.1-
0.2 mm long; stigmas about 0.2 mm long, 2-lobed; stipes 2.1-3.2 mm long,
glabrous; nectaries 1, adaxial, 0.2-1 mm long, broad, 0.2-0.5 times as long
as stipe; bracts ovate or narrowly oblong, apex rounded, 0.8-1.6 mm long,
tawny, sometimes with a reddish tinge, glabrous abaxially, sparsely villous
adaxially.
Habitat Range
Wet Carex-Betula glandulosa-Salix Boreal: Watson Lake, southeastern
fens; muskegs. Yukon; adjacent British Columbia;
116
Myrtilloides Koehne
170° 135° 130° 125°
yy LEE, 4
Map 31 Salix pedicellaris
southern Northwest _ Territories;
southward in the Rocky Mountains to
Washington; eastward in the boreal
forest to Newfoundland; northern U.S.
from Minnesota to New _ England
(Map 31).
Discussion
Salix pedicellaris is a characteristic low willow of wet fens with erect
branches and coriaceous, glabrous leaves which are glaucous on both sides
and reticulate above. The aments are loosely flowered; the pistils are usually
reddish and glabrous and are borne on long stipes, 2.1-3.2 mm long. It is
one of two Salix species known to have leaves with a waxy glaucescence on
the upper leaf surface, the other being S. purpurea L.
In the Yukon it is known only from Watson Lake where it is locally abun-
dant. There is no evidence that this species occurs in the Pelly River Valley
as reported by Porsild (1951). The specimen (Porsi/d & Breitung 9776) on
which Porsild’s report was based has been re-identified as S. athabascensis.
This species hybridizes with S. athabascensis, which should be seen for dis-
cussion. A series of suspected intermediate specimens from Watson Lake
117
Plate XXI
Salix hastata L. Johnson 135 (left) and Viereck 7869 (right)
118
Hastatae Kern.
Table 10 Comparison table: Salix pedicellaris and S. myrtilloides
Characteristics pedicellaris myrtilloides
habit erect shrub, 2-15 dm tall subterranean, creeping shrub,
0.3-0.5 dm tall
leaf shape narrower: narrowly elliptic broader: subcircular to ovate
to narrowly rectangular or or narrowly elliptic
narrowly obovate
immature leaves glabrous sparsely pubescent
have been determined as S. pedicellaris > athabascensis. These specimens
are characterized by immature leaves which are sparsely sericeous with
white and ferruginous trichomes, ovaries sometimes undeveloped but not
always so, stipes pubescent and pistils with patchy pubescence. In all other
respects they are ‘‘typical’’ S. pedice//aris and are assumed to be backcrosses
between S. athabascensis x pedicellaris and S. pedicellaris (Map 2).
Salix pedicellaris is closely related to the Eurasian S. myrtilloides L. from
which it can be distinguished by the characteristics listed in Table 10.
Section 10. Hastatae Kern.
23. SALIX HASTATA.L.
o. Mastatia. k. So. Pl. 10T7. 1753.
S. farrae Ball, Contr. U.S. Nat. Herb. 22: 321. 1921. (Type: Farr 558, US).
S. hastata var. farrae Hult. Ark. Bot. Il. 7: 42. 1967.
S. farrae var. walpolei Cov. and Ball, Bot. Gaz. 71: 435. 1921. (Type: Wal-
pole 1624, US). S. walpolei Ball, Madrofo 6: 231. 1942. S. farrae ssp.
walpolei att. to Ball by Hult. Fl. Alaska and Yukon 3: 532. 1943.
S. hastata ssp. S. psiloides Flod. Sv. Vet. Akad. Ark. Bot. 20A: 54. 1926.
S. farrae var. microserrulata Ball, Univ. Calif. Publ. Bot. 17: 410. 1934.
Description of species
Shrubs 2-7 (10) dm tall in Arctic, 1-3 m tall in interior; branches brown,
usually glabrescent but indumentum may persist for 2-3 years; branchlets
reddish brown, commonly white villous with short curved trichomes, some-
times sparsely pubescent to glabrescent. Leaves elliptic to obovate, the
largest mature leaves 2.5-4-6.8 cm long, 0.1-2-3.2 cm wide and 1.6-2.3-2.6 (3.4)
times as long as wide; apex acute to more or less attenuate; base acute or
round; margins entire or indistinctly and irregularly glandular serrulate,
revolute; the lower side of immature leaves sparsely pubescent with white or
white and ferruginous trichomes, pale; the upper side of mature leaves gla-
brescent and glossy, ferruginous trichomes often persistent on the midrib, the
418
Hastatae Kern.
lower side glabrous or glabrescent, thinly glaucous or pale and non-glaucous;
petioles 1.5-4-6 (9) mm long, reddish, pubescent adaxially; stipules 1-4 (6)
mm long, ovate to elliptic, margins glandular dotted. Aments coetaneous, on
short, leafy, floriferous branchlets. Staminate aments 1.3-2.5 cm long, florif-
erous branchlets 0.1-3 mm long with small bract-like leaves; stamens 2,
filaments 3-4.4 mm long, glabrous; anthers 0.4-0.6 mm long. Pistillate
aments 2.5-4-7 cm long, often loosely flowered, floriferous branchlets 5-12
mm long; pistils 2-4 mm long, green and more or less reddish on lower half,
glabrous, capsules 3.2-8 mm long, greenish to tawny and remaining reddish
at base; styles 0.2-0.4 mm long, occasionally bifid; stigmas small 0.2-0.3
mm long, 2-4-lobed; stipes 0.4-0.8 (1.2) mm long, sparsely pubescent,
nectaries 1, adaxial, 0.3-0.6 mm long, broad at base and tapering toward
apex, about 0.5 times as long as stipe; bracts narrowly oblong, apex acute,
1.2-1.8 mm long, light brown or bicolour, sparsely pubescent on both sides
with long straight trichomes.
Habitat Range
Salix thickets along rivers and Arctic, montane: Northern Alaska;
streams; sandy tundra; Carex-Erio- mountains of central Alaska; Alaska
phorum meadows. Range; Dawson, Yukon _ Territory;
northwestern Northwest Territories;
Eurasia (Map 32).
Discussion
Salix hastata is characterized by branches villous with short, curved tri-
chomes; leaves with ferruginous trichomes on upper surface, especially the
upper surface of the immature leaves, non-glaucous beneath becoming pale
or sparsely glaucous and with more or less entire to irregularly glandular
serrulate margins; and pistils reddish, especially at base, borne on very short
stipes.
It is closely related to S. barclayi, and reports of the latter species from
arctic Alaska are usually misidentifications of S. hastata. These species may
be distinguished as in Table 11. For a comparison with S. novae-angliae, S.
myrtillifolia and S. commutata see Table 13.
|! agree with Hultén (1967) that S. farrae and S. walpolei should be in-
cluded within the Eurasian S. hastata. Salix hastata is highly variable; three
subspecies are recognized by Rechinger (1964) in Flora Europea and one
variety (var. farrae) is recognized in North America (Hultén 1967). The
North American taxon S. hastata var. farrae has a disjunct distribution in
southeastern British Columbia, in Alberta, Idaho, Montana and Wyoming, and
is distinguishable from the northern populations by its sparsely pubescent
branchlets and leaves distinctly glaucous beneath. However, the other char-
acteristics cited by Ball (1934) and reiterated by Hultén (1967), such as
yellowish branchlets, smaller, narrower leaves, shorter aments, and so on,
are of doubtful reliability. | am not convinced the variety is taxonomically
useful. Salix hastata in North America requires critical field study.
120
Cordatae Barr. ex Hook.
170° 160°
Map 32 Salix hastata. Circle based on Skvortsov, 1966
Table 11 Comparison table: Salix hastata and S. barclayi
Characteristics hastata barclayi
leaf glaucescence non-glaucous or thinly glaucous beneath glaucous beneath
leaf pubescence ferruginous trichomes no ferruginous
on immature leaves trichomes
leaf colour not drying dark often drying dark
petioles reddish green
style length 0.2-0.4 mm 0.6-1.6 mm
pistils reddish, at least at base green
Section 11. Cordatae Barr. ex Hook.
24. SALIX RIGIDA Muhl.
S. rigida Muhl. in Muhl. and Willd. Neue Schr. Ges. Nat. Fr. Berlin 4: 236.
1803.
S. cordata (ssp.) rigida Anderss. Kg. Sv. Vet. Akad. Handl. 6: 158. 1867.
121
Plate XXIl
Salix rigida Muhl. Argus 6069
122
Cordatae Barr. ex Hook.
S. cordata Muhl. in Muhl. and Willd. Neue Schr. Ges. Nat. Fr. Berlin 4: 236.
1803, non Michx. 1803.
S. cordata y (var.) mackenzieana Hook. Fl. Bor.-Am. 2: 149. 1838. (Type:
Richardson s.n. GH). S. mackenzieana Barr. ex Hook. Fl. Bor.-Am. 2: 149.
1838, pro. syn. S. mackenzieana Barr. ex Anderss. Kg. Sv. Vet. Akad. Hand.
6: 160. 1867. (as mackenziana). S. * mackenzieana Anderss. in DC. Prodr.
16 (2): 252. 1868. (as S. cordata x rostrata). S. rigida var. mackenzieana
Crong. in Hitchcock, Cronquist, Ownbey and Thompson, Vas. Pl. Pacific NW.
2: 63. 1964.
S. mackenzieana var. macrogemma Ball, in Piper and Beattie, Fl. Northwest
Coast 116. 1915. S. rigida var. macrogemma Crong. in Hitchcock, Cronquist,
Ownbey and Thompson, Vas. Pl. Pacific NW. 65. 1964.
Description of species
Shrubs 0.3-3 m tall; branches reddish brown, glabrous and glossy; branch-
lets reddish brown to yellow green, glabrous or velutinous. Leaves narrowly
oblong-obovate, 5-10.5 cm long, 1.2-2 cm wide and 3.7-5-6 times as long as
wide; apex gradually to abruptly acuminate; base round to rarely sub-
cordate; margins serrulate; immature leaves reddish, translucent, densely to
sparsely pubescent; the upper side of mature leaves glabrescent, the midrib
often remaining velutinous, the lower side glaucous and glabrescent; petioles
8-17 mm long, reddish, velutinous adaxially; stipules narrowly elliptic to
ovate, 5-9 (20) mm long, margins serrulate; buds velutinous to glabrescent,
inner bud scale separating from the outer and often clinging to base of shoot.
Aments coetaneous or subprecocious, on short, floriferous branchlets. Stami-
nate aments 1.5-2.5 cm long; floriferous branchlets 2-5 mm long; stamens
2, filaments connate at base, glabrous; anthers 0.5-0.6 mm long. Pistillate
aments 3-5 cm long; floriferous branchlets 3-13 mm long; pistils slender,
reddish or greenish and glabrous, capsules 4-5 mm long, tawny; styles 0.5-
0.75 mm long; stigmas small; stipes 1-2 mm long, glabrous, nectaries 1,
adaxial, 0.2-0.5 mm long, ca. 0.25-0.5 times as long as the stipe; bracts
narrowly elliptic, apex acute and reflexed in fruit, 1-2 mm long, tawny to
dark brown, sparsely villous.
Habitat Range
Sand bars and mud flats along rivers. Boreal: Infrequent in southern Yukon
and adjacent British Columbia; south-
ward to Washington, California, and
Arizona; eastward to Newfoundland
and Virginia (Map 33).
Discussion
Salix rigida is a transcontinental North American species which just barely
enters southern Yukon. The northwestern North American populations are
sometimes treated as the species S. mackenzieana (Raup 1959) or as S.
123
Cordatae Barr. ex Hook.
175° 170° 165° 160° 155° 150° 145° 140° 135° 130° 125° 120° 115°
WSS
170 165
175° 180° 175°
Map 33 Salix rigida
rigida var. mackenzieana (Cronquist 1964). The taxon mackenzieana is sup-
posed to be distinguished from S. rigida by its longer stipes (3-4 mm long in
fruiting material) and immature leaves glabrous, or nearly so. | have not
found these characteristics to be consistent or useful in distinguishing be-
tween mackenzieana and rigida, and | prefer to recognize them both under
the latter name until the entire complex can be studied.
Salix rigida is closely related to S. monticola, and although it may be dis-
tinguished by its narrower leaves and shorter styles (Table 12), there are
intergradations in leaf shape in southern Yukon and the two may eventually
be combined on the varietal level.
25. SALIX MONTICOLA Bebb
S. monticola Bebb in Coult. Man. Bot. Rocky Mt. Reg. 336. 1885. (Type:
Greene s.n. F).
S. cordata var. monticola Kelso, Biol. Leafl. 34: 7. 1946.
S. padifolia Rydb. Bull. Torr. Bot. Club 28: 272. 1901. non Anderss. 1859.
124
Cordatae Barr. ex Hook.
Table 12 Comparison table: Salix rigida, S. monticola and S. barclayi
Characteristics rigida monticola barclayi
immature leaves reddish, translucent reddish, translucent green, opaque
leaf shape narrowly oblong-obovate _ elliptic or obovate elliptic or obovate
leaf I/w 3.7-5-6 1.4-2-2.8 1.6-2.1-2.8
petioles reddish reddish green
branchlet glabrous or glabrous or densely to sparsely
pubescence velutinous sparsely pubescent villous
flowering time coetaneous or precocious coetaneous
subprecocious
floriferous 0.3-1.3 cm 0-0.5 cm (0.5) 1-2(3) cm
branchlet length
style length 0.5-0.75 mm 0.8-1.2 mm 0.6-1.6 mm
(Type: Rydberg & Vreeland 6389, NY). S. padophylla Rydb. Bull. Torr. Bot.
Club 28: 499. 1901. S. pseudomonticola Ball var. padophylla Ball, J. Wash.
Acad. Sci. 28: 450. 1938. S. barclayi var. padophylla Kelso, Biol. Leafl. 34:
8. 1946.
S. pseudomonticola Ball in Standl. Contr. U.S. Nat. Herb. 22: 321. 1921.
(Lectotype: Sanson 233, US). S. barclayi var. pseudomonticola Kelso, Biol.
Leafl. 34: 8. 1946.
Description of species
Shrubs 1-2-4 m tall; branches dark reddish brown to yellow brown, glossy;
branchlets yellow green, glabrous or sparsely pubescent to glabrescent and
glossy, sometimes sparsely glaucous. Leaves narrowly elliptic, elliptic or
obovate, the largest mature leaves 4.1-6.9 (8) cm long, 2.0-3.3 (4) cm wide
and 1.4-2-2.8 times as long as wide; apex broad and abruptly acuminate to
acute; base obtuse, round to subcordate, often inequilateral; margins glan-
dular crenate-serrulate; immature leaves reddish, translucent and sparsely
pubescent, the midrib often remaining reddish; the upper side of mature
leaves green and glabrous or glabrescent, the lower side glabrescent and
glaucous; petioles 6-12 (20) mm long, usually reddish, glabrous or pubescent;
stipules ovate with round apex, 1-5-12 mm long, margins crenate-serrulate
or often indistinctly so, glabrous on both sides and glaucous beneath. Aments
precocious, usually sessile or on very short floriferous branchlets bearing
several bract-like leaves. Staminate aments 2.5-3 cm long, sessile; stamens
2, filaments 2-3.5 mm long, distinct or connate at base, glabrous; anthers
0.4-0.5 mm long, drying purplish. Pistillate aments 4-9 cm long, usually
sessile, floriferous branchlets less than 0.5 cm long; pistils about 4.5 mm
long, green and glabrous, capsules 5.5-6.5 mm long, tawny; styles 0.8-
1.2 mm long; stigmas 0.2-0.3 mm long, 2 slender lobes; stipes 0.9-1.5-2.2
mm long, glabrous; nectaries 1, adaxial, 0.3-0.8 mm long, 0.5-0.66 times as
long as stipe; bracts narrowly oblong, apex rounded or attenuate, 1.2-2.4
125
Plate XXIll
Salix monticola Bebb. Pistill
5121 (right) istillate specimen, Cody 8056 (left) and vegetative specimen, Argus
126
Cordatae Barr. ex Hook.
180° 175° 170° 165° 160° 155° 150° 145° 140° 135° 130° 125° 120° 5°
Map 34 Salix monticola
mm long, dark brown to blackish, sometimes bicolour, sparsely villous on
both sides with long straight trichomes.
Habitat Range
Salix-Carex fens in drainage ways in Boreal: Central Alaska; eastern Alas-
Picea glauca forests; P. mariana mus- ka Range; across the southern half of
kegs; Populus balsamifera forests. Yukon Territory; Northwest Territo-
ries; southward in the cordillera to
Colorado and New Mexico; eastward
to Saskatchewan and South Dakota
(Map 34).
Discussion
Salix monticola is characterized by precocious flowering, sessile aments,
reddish immature leaves, petioles and lower midribs, small rounded stipules
and sparse leaf and stem pubescence. It is a member of the complex includ-
ing S. rigida and S. barclayi. The species may be distinguished on the basis
of characteristics in Table 12. It is also related to S. myrtillifolia and S.
novae-angliae, which can be distinguished from it by their non-glaucous
127
Plate XXIV
Salix barclayi Anderss. Argus 6080 (left) and Argus 6680 (right)
128
Cordatae Barr. ex Hook.
leaves. It may sometimes be confused with S. /anata ssp. richardsonii from
which it may be distinguished by its small stipules and very sparse leaf and
branchlet pubescence.
Salix monticola is a western North American species reaching its north-
western distributional limit in central Alaska. It varies somewhat in the
Rocky Mountains, and two specific names have been based on differences in
leaf shape. | am following Cronquist (1964) in combining the broad-leaved
S. padophylla with the narrower-leaved S. monticola. The species in Alaska
and the Yukon is represented by the broad-leaved variant.
26. SALIX BARCLAY! Anderss.
S. barclayi Anderss. Ofvers. Vet. Akad. Forh. (Stockh.) 15: 125. 1858. (Type:
Barclay s.n. photo and frag. A).
S. barclayi f. rotundifolia Anderss. in DC. Prodr. 16(2): 254. 1868.
S. barclayi f. grandifolia Anderss. in DC. Prodr. 16(2): 254. 1868.
S. barclayi f. angustifolia Anderss. in DC. Prodr. 16(2): 255. 1868.
S. conjuncta Bebb, Bot. Gaz. 13: 111. 1888. (Type: Parry s.n. F).
Description of species
Shrubs usually 1-3 m tall, but varying from prostrate to 7 dm or up to 5 m
tall; branches dark reddish brown or sometimes chestnut brown, glabrous or
with indumentum persistent for two years; branchlets yellow green, glossy,
densely to sparsely villous with long straggly trichomes, becoming sparsely
pubescent or glabrescent, occasionally glabrous. Leaves elliptic or obovate,
rarely narrowly elliptic, the largest mature leaves 3.3-7 (9.9) cm long, 1.2-
3.5 (4.8) cm wide, 1.6-2.1-2.8 (4) times as long as wide; apex broad,
the tip acuminate or acute; base commonly rounded, sometimes subcordate
or acute and inequilateral; margins glandular serrulate, occasionally suben-
tire; the upper side of immature leaves sparsely villous, especially along
midrib, the lower side glabrous or very sparsely pubescent; the upper side of
mature leaves glabrescent but indumentum may persist along midrib, the
lower side glabrous and glaucous, leaves green in life, drying dark brown or
blackish especially along the venation; petioles 3-6-14 (20) mm long, tawny,
villous adaxially; stipules 1.5-5.13 (22) mm long, ovate or narrowly elliptic, the
apex round or attenuate, margins glandular dotted or glandular serrulate,
the lower half of upper side often glandular dotted, the lower side glabrous
and glaucous. Aments coetaneous, on leafy, floriferous branchlets. Staminate
aments 2.5-4 cm long, floriferous branchlets 5-11 mm long; stamens 2, fila-
ments 5.6-6.5 mm long, glabrous, distinct; anthers 0.6-0.7 mm long. Pis-
tillate aments (2.5) 4-7 (8) cm long, floriferous branchlets (0.5) 1-2 (3)
cm long; pistils about 3.5 mm long, green and glabrous, capsules 5-6.5 mm
long; styles 0.6-1.6 mm long; stigmas 0.4-0.6 mm long, 2-lobed; stipes 0.4-
129
Cordatae Barr. ex Hook.
Map 35 Salix barclayi
1.4 mm long, glabrous or pubescent; nectaries 1, adaxial, 0.4-0.8 mm long,
about 0.5 times as long as stipe; bracts narrowly oblong, apex acute to at-
tenuate, 1.6-2.8 mm long, light brown, dark brown or bicolour and apex
darker, pubescent on both sides with long straight trichomes.
Habitat
Large Salix thickets on glacial mo-
raine; lake and river shores; subal-
pine and alpine slopes; occasionally
in muskegs, fens, Picea glauca-Popu-
lus tremuloides, and Picea sitchensis
forests.
Discussion
Range
Boreal: Southern coastal Alaska from
the eastern Aleutian Islands to Hyder;
Alaska Range; Tanana River; south-
ern Yukon and adjacent Northwest
Territories; southward in the Rocky
Mountains to British Columbia,
Washington, and Alberta (Map 35).
Salix barclayi is characterized by elliptic to obovate leaves sparsely pubescent
above and glabrous and glaucous beneath, often drying brownish, margins
glandular serrulate, stipules prominently glandular dotted above;
pistils
glabrous and nectaries half as long as the stipes.
130
Cordatae Barr. ex Hook.
It is related to S. novae-angliae, S. myrtillifolia and S. commutata from
which it may be distinguished by its glaucous leaves (Table 12); to S. hook-
eriana from which it may be distinguished by a series of characteristics in-
cluding its prominent stipules and glabrous mature leaves (see S. hookeriana
for discussion); to S. monticola, S. rigida (Table 12) and to S. hastata (Table
11), each of which should be referred to for further discussion.
This species forms relatively constant populations in reference to branch-
let pubescence, leaf shape, pubescence and serration and to stipule size and
shape in southern coastal Alaska, from the Kenai Peninsula southward. At
the northwestern end of this range the branchlets tend to be more glabres-
cent, and further westward to the Aleutian Islands specimens with leaves
sparsely pubescent at maturity become more frequent. In the Alaska Range
there are specimens with large and more persistent stipules, some with ine-
quilateral, attenuate apices resembling S. /anata ssp. richardsonii. These
specimens can be distinguished from S. /anata ssp. richardsonii by their nar-
rower leaves, more distinctly serrulate leaf margins and by aments borne on
leafy, floriferous branchlets (see Table 18).
In northwestern British Columbia, between Alaska and the Yukon, there is
a possible introgressive population which deserves some comment. On the
south side of Chilkat Pass S. barclayi has the typical coastal stipule and
branchlet morphology. The stipules are ovate or narrowly elliptic, round or
attenuate at the apex, shorter than the petioles and not persistent beyond one
year; the branchlets are glabrous to sparsely villous. However, in the alpine
vegetation on Chilkat Pass (between the coastal and the interior drain-
ages on the Haines Highway) there is a population of S. barclayi which ap-
proaches S. Janata ssp. richardsonii in some morphological characteristics.
The stipules are broader at the base and the apex is inequilaterally attenuate;
the stipules are longer than the petioles (Fig. 4), and are often persistent
into the second year. The branchlets are more coarsely villous with more
persistent pubescence. The specimens can be identified as S. barclayi on the
basis of aments borne on distinct, leafy, floriferous branchlets, a character-
istic which does not vary in the intermediate population, but S. /anata ssp.
richardsonii also occurs in these alpine populations and hybridization and
introgression may be an explanation for this unusual variation within S.
barclayi.
See S. stolonifera for a discussion of hybridization with S. barclayi.
27. SALIX HOOKERIANA Barr.
S. hookeriana Barr. in Hook. Fl. Bor.-Am. 2: 145. 1838.
S. amplifolia Cov. Proc. Wash. Acad. Sci. 2: 282. 1900. (Type: Coville &
Kearney 1153, US).
Description of species
Trees or shrubs 0.6-1-5 (8) m tall, rarely prostrate, the largest stems about
131
Plate XXV
Salix hookeriana Barr. Argus 6126
132
Cordatae Barr. ex Hook.
13 cm in diameter; branches thick and very brittle, reddish brown, usually
with lanate or villous-lanate indumentum persistent for 2-3 years; branchlets
usually densely white villous-lanate and remaining so throughout the year,
sometimes becoming sparsely villous; buds densely white villous. Leaves
broadly elliptic to elliptic or broadly obovate, the largest mature leaves 3.6-7
(10.3) cm long, 1.9-3.9-6.3 cm wide and 1.5-1.9 (2.1) times as long as
wide; apex round, with acute tip or obtuse, often conduplicate when pressed;
base round; margins distantly and irregularly glandular crenate especially
near the base, often crisped in life, sometimes entire, often more or less
revolute; immature leaves densely white villous on both sides; the upper
side of mature leaves sparsely pubescent, yellow green and glossy, the
lower side sparsely villous to glabrescent except for the midrib which
retains long straight trichomes, thinly glaucous; petioles 4-12 mm long, dense-
ly white lanate to sparsely villous; stipules usually absent but sometimes
present on vigorous shoots, 0.5-1.8 (2.5) mm long. Aments coetaneous or
subprecocious, on leafy, floriferous branchlets. Staminate aments 2.5-5.5 cm
long, usually very thick, floriferous branchlets 0-0.5-2.3 cm long, densely vil-
lous, leaves usually small and bract-like; stamens 2, filaments about 9 mm
long, glabrous, distinct or slightly connate at base; anthers (0.5) 0.8-1 mm
long. Pistillate aments 2.2-8-14 cm long, ovaries at distal end sometimes
undeveloped, capsules at proximal end usually reflexed, floriferous branch-
lets 1-3.5 cm long with normal or bract-like leaves; pistils 3.6-6.4 mm long,
green, glabrous or partially or completely lanate, capsules up to 8-10 mm
long, tawny; styles 1.1-2.3 mm long, red in life; stigmas 0.5-0.6 mm long;
stipes 0.6-1.8 mm long, lanate; nectaries 1, adaxial, 0.6-0.8 mm long, 1-2-
lobed, about 0.5-0.66 times as long as stipe; bracts broadly oblong, apex
rounded, sometimes retuse or acuminate, 2.4-3.6 mm long, brown or black-
ish, villous on both sides.
Habitat Range
Stabilized sand dunes; wet Carex Pacific coastal: Yakutat Bay region;
meadows near coast; beach ridges; Childs Glacier; Middleton Island; Ko-
edges of Heracleum-Angelica mead- diak Island; southwestern British Co-
ows; Salix thickets on morainal flats. lumbia; Washington; Oregon; Cali-
fornia (Map 36).
Discussion
Salix hookeriana is a very distinctive coastal tree or shrub characterized by
brittle branches; densely villous-lanate branchlets with long, persistent villous
trichomes at the base; leaves pubescent on both sides, although sometimes
sparsely so; stipules apparently absent except on vigorous sprout shoots, or,
if present, very small; aments often very stout and long, pistils glabrous or
pubescent and styles red in life.
This species is related to S. barclayi, but it is not conspecific with it as
maintained by Hultén (1967). The two are amply distinct and show no inter-
gradation even within sympatric populations at Yakutat and Tanis Mesa in
southern coastal Alaska. Salix hookeriana can be distinguished from S.
133
Cordatae Barr. ex Hook.
stipule length (mm)
1 2 3 4 5 10 15
petiole length (mm)
Fig. 4 The variation in stipule and petiole acterized by having larger stipules than the
length in Salix barclayi. A population from coastal populations (dots). However, wide
the Three Guardsman Pass, Haines Highway, variation is revealed in both regions. The
British Columbia (circles) is generally char- scales are logarithmic.
barclayi by its brittle branches, villous-lanate branches and branchlets, leaves
pubescent beneath, stipules absent or very small and red styles.
Salix hookeriana was first described from Alaska as S. amplifolia and was,
until 1967, known only from the type material. In 1967 | revisited the type
region at Yakutat Bay and found this willow to be abundant as a tree on
stabilized coastal sand dunes. It was subsequently recognized and collected
near the town site of Yakutat and at Tanis Mesa. Positively identified spec-
imens, originally cited as S. barclayi and its hybrids (Thomas 1957), have
since been seen from Middleton Island. In addition, more or less equivocal
specimens have been seen from Childs Glacier near the Copper River and
from Kodiak. This species should be sought in suitable habitats elsewhere in
coastal Alaska.
The decision to combine S. amplifolia with S. hookeriana was reached after
careful consideration of ample material of both taxa. Each of these taxa is
134
Cordatae Barr. ex Hook.
180° 175° 170° 165° 160° 155° 150° 145° 140° 135° 130° 125° 120° 115°
A
Map 36 Salix hookeriana. Circle based on a specimen of doubtful identity
highly variable, and the differences which have been used to differentiate
between them, such as style length, leaf pubescence and size of the florif-
erous branchlets, are in themselves highly variable and not diagnostic.
Salix hookeriana has been reported for eastern Siberia by Floderus (1933)
who cited a collection from the Anadyr region, and by Nasarov (1936) from
Len-Kolymsk (Zhiganska). Skvortsov (1966) is inclined to doubt the accuracy
of these records.
28. SALIX COMMUTATA Bebb
S. commutata Bebb, Bot. Gaz. 13: 110. 1888. (Type: Cusick 826, F).
S. barclayi var. commutata Kelso, Biol. Leafl. 34: 8. 1946.
S. commutata var. denudata Bebb, Bot. Gaz. 13: 111. 1888. (Type: Cusick
1304c, -F)..
S. commutata var. puberula Bebb, Bot. Gaz. 13: 111. 1888.
S. commutata var. sericea Bebb, Bot. Gaz. 13: 111. 1888. (Type: Howell s.n.
F). S. commutata (ssp.) mixta Piper, Contr. U.S. Nat. Herb. 11: 216. 1906.
135
Plate XXVI
Salix commutata Bebb. Argus 6588
136
Cordatae Barr. ex Hook.
Description of species
Low shrubs 0.2-1 (2) m tall; branches dark brown, sometimes remaining
pubescent for two years; branchlets densely white lanate to sparsely villous.
Leaves elliptic to broadly or narrowly elliptic, the largest mature leaves (2.8)
3.5-5.5 (10) cm long, 1.3-3.4 (4.4) cm wide and (1.5) 1.7-2.6 (3.4) times
as long as wide; apex acute, acuminate on vigorous leaves; base round or
rarely subcordate; margins glandular serrulate on proximal half and entire
on distal half or completely entire or completely glandular serrulate; im-
mature leaves densely villous on both sides; the upper side of mature leaves
yellow green, villous with long straggly trichomes, the lower side villous-
lanate with mixture of long straight and short curly trichomes, green (non-
glaucous) and glossy, leaves sometimes becoming sparsely villous on both
sides; petioles short, 1.5-7 (10) mm long, villous; stipules half-ovate, (0.8)
1-6 (9) mm long, green, sparsely villous and glandular serrulate. Aments
coetaneous, on leafy, floriferous branchlets. Staminate aments 1.5-3.5 cm
long, floriferous branchlets 0.6-2 cm long; stamens 2, filaments 5-7 mm
long, glabrous, distinct or connate below; anthers 0.4-0.9 mm long. Pistillate
aments 3.5-7 cm long, floriferous branchlets 1.2-3.5 cm long; pistils pyri-
form, 2-4 mm long, usually reddish but sometimes greenish, glabrous, cap-
sules 4.4-6.4 mm long, reddish, greenish or tawny; styles 0.6-1.2 mm long;
stigmas about 2 mm long, 2-lobed; stipes 0.3-1.2 mm long, glabrous; nec-
taries 1, adaxial, 0.3-0.6 mm long, 0.5-0.66 time as long as stipe; bracts
narrowly oblong, apex acute, 1.1-2 mm long, tawny, dark brown or bicolour
and darker at apex, pubescent with short wavy or long straight trichomes.
Habitat Range
A pioneer species on glacial moraine Alpine, boreal: Southern coastal Alas-
and rocky slopes; alpine tundra (up ka from Unalaska; Aleutian Islands;
to 5,000 ft); gravel benches along Kodiak Island; Kenai Peninsula; cen-
rivers; Salix thickets in wet fens. tral Alaska Range; southward to the
northern end of southeastern Alaska;
southern Yukon Territory, especially
the mountains near the Northwest
Territories boundary; adjacent North-
west Territories and British Colum-
bia; southward in the cordillera to
California, Utah, Montana, and Wyo-
ming (Map 37).
Discussion
Salix commutata is characteristically a late-flowering species with non-glau-
cous leaves which have a distinctly straggly pubescence composed of a mix-
ture of curly and straight trichomes. The petioles are short (1.5-7 mm long)
and the stipules are often as long as the petioles. The pistils are usually red-
dish or reddish orange but may grade into yellow green. This species is re-
lated to S. novae-angliae, S. myrtillifolia and is similar to S. hastata. These
137
Plate XXVII
Salix novae-angliae Anderss. Argus 6935 (left) and Argus 5067 (right)
138
Cordatae Barr. ex Hook.
170° 165° 160° 155°
Map 37 Salix commutata
taxa may be distinguished as in Table 13. Since S. commutata is often sym-
patric with S. barclayi the two are sometimes confused, but the distinctive
non-glaucous leaves and persistent pubescence on both sides of the leaves,
as well as the reddish pistils, serve to distinguish S. commutata.
Salix commutata is variable in leaf shape and pubescence, margin denti-
tion and pistil colour. Two varieties have been based on variation in leaf in-
dumentum, var. denudata and var. sericea, but they seem to be well within
the normal variational limits for the species. Immature leaves are usually
densely villous, but as they unfold, the lower surface near the apex may be-
come glabrescent, and throughout leaf ontogeny the general pubescence
progressively diminishes so that old leaves may be almost glabrescent. The
leaf margin dentition varies considerably, from entire to glandular serrulate
on the lower half only, to entirely serrulate with all intermediate stages pos-
sible. Some specimens with serrulate leaf margins were recognized by Hultén
(1943) as the hybrid S. commutata x myrtillifolia. In my judgement, there is
no justification for using hybridization to explain this variation.
139
Cordatae Barr. ex Hook.
Table 13 Comparison table: Salix commutata, S. novae-angliae, S. myrtillifolia and S. hastata
Characteristics commutata novae-angliae myrtillifolia hastata
leaves beneath non-glaucous non-glaucous non-glaucous non-glaucous or
thinly glaucous
immature leaf densely villous sparsely villous glabrous pubescent with
pubescence with white with white or white or ferrugi-
trichomes rarely ferruginous nous trichomes
trichomes
mature leaf sparsely villous- glabrescent glabrous glabrescent
pubescence lanate
pistil colour usually reddish green green partly reddish
leaf margins serrulate to crenate crenate to cre- entire or irregu-
entire nate-serrulate larly serrulate
style length 0.6-1.2 mm 0.5-0.9 mm 0.3-0.5 mm 0.2-0.4 mm
29. SALIX NOVAE-ANGLIAE Anderss.
S. novae-angliae Anderss. Kg. Sv. Vet. Akad. Handl. 6: 160. 1867. (Type:
Drummond 665, NY).
S. myrsinites L. (ssp.) S. pseudomyrsinites Anderss. Ofvers. Vet. Akad. Forh.
(Stockh.) 15: 129. 1858. S. novae-angliae (ssp.) S. pseudomyrsinites Anderss.
Kg. Sv. Vet. Akad. Handl. 6: 160. 1867. S. novae-angliae a (var.) pseudo-
myrsinites Anderss. in DC. Prodr. 16 (2): 253. 1868. S. myrtillifolia Anderss.
var. pseudomyrsinites Ball ex. Hult. Fl. Alaska and Yukon 3: 538. 1943.
S. myrsinites L. (ssp.) S. curtiflora Anderss. Ofvers. Vet. Akad. Forh.
(Stockh.) 15: 130. 1858. S. novae-angliae (ssp.) S. pseudocordata Anderss.
Kg. Sv. Vet. Akad. Handl. 6: 161. 1867. S. novae-angliae B (var.) pseudo-
cordata Anderss. in DC. Prodr. 16 (2): 253. 1868. S. myrtillifolia (ssp.)
curtiflora Rose, Contr. U.S. Nat. Herb. 3: 573. 1896. S. pseudocordata Rydb.
Fl. Colo. 94. 1906.
Description of species
Erect shrubs 0.6-2-4 m tall; branches greyish brown or dark reddish brown,
indumentum may persist for 2-3 years or the branches soon become gla-
brate, glaucous; branchlets usually densely white villous with long straight
trichomes, becoming sparsely pubescent with straight and curved trichomes,
sometimes glaucescent. Leaves narrowly elliptic to narrowly obovate, the
largest mature leaves 3.2-4.5-6.8 cm long, 1-1.7-2.7 cm wide and 2-2.8-
4.8 times as long as wide; apex broadly acute to obtuse or round; base
round; margins glandular crenate or glandular crenate-serrulate; immature
leaves reddish and villous; the upper side of mature leaves glabrescent, but
midrib remaining pubescent with white or sometimes ferruginous trichomes,
glossy, the lower side usually glabrescent, rarely sparsely villous, glossy,
pale green, non-glaucous, the leaves becoming brownish on drying; petioles
140
Cordatae Barr. ex Hook.
180° 175° 170° 165° 160° 155° 150° 145° 140° 135° 130° 120° 115°
Map 38 Salix novae-angliae
2.5-3-7 mm long, yellow green, pubescent adaxially; stipules semi-ovate 0.5-
2-7 mm long, margins glandular and more or less crenate. Aments coetane-
ous, on leafy, floriferous branchlets. Staminate aments 1.5-2 cm _ long,
floriferous branchlets about 3 mm long; stamens 2, filaments about 2.5-4 mm
long, glabrous; anthers 0.4-0.7 mm long. Pistillate aments (1.5) 2.2-3-4.5 cm
long, floriferous branchlets 0.5-1-1.8 cm long; pistils 4-5.5 mm long, green,
glabrous, capsules 4.4-6.4 mm long; styles 0.5-0.9 mm long; stigmas 0.2-
0.4 mm long; stipes 0.8-1.4 mm long, glabrous; nectaries 1, adaxial, 0.2-
0.4 mm long, about 0.25 times as long as stipe; bracts oblong, apex rounded,
0.7-0.9 mm long, bicolour, apex dark brown and tawny below, or uniformly
brown, glabrescent or pubescent on both sides with long wavy trichomes.
Habitat Range
Salix thickets on shores of lakes, Boreal: Central Alaska; southern half
streams, and rivers with Betula of Yukon Territory and adjacent
glandulosa-Alnus or Populus balsami- Northwest Territories; southward in
fera; prairie margins; muskegs. the cordillera to California; eastward
to Saskatchewan (Map 38).
141
Plate XXVIII
Salix myrtillifolia Anderss. Argus 373-58
142
Cordatae Barr. ex Hook.
Discussion
Salix novae-angliae is characterized by erect shrubs 0.6-2-4 m tall; leaves
green on both sides, sometimes pale beneath but non-glaucous, often drying
brownish, margins glandular crenate; pistils glabrous, stipes about 1 mm long
and about 4 times as long as the nectary.
This species is closely related to S. myrtillifolia and has long been con-
fused with it in Alaska and the Yukon. However, it has been evident to stu-
dents of western Canadian flora and Alaskan flora that there are two related
elements in this area, one a low, spreading shrub (S. myrtillifolia) and the
other a tall shrub (S. novae-angliae). In 1943 Hultén recognized the tall
shrub as S. myrtillifolia var. pseudomyrsinites and contrasted it with S. pseu-
docordata with which it is probably synonymous. Hultén’s comments on this
taxon confused students of the group because he stated that the styles are 1
mm long or longer, whereas they are 0.5-0.9 mm long, and that they have
glaucous leaves, whereas they are non-glaucous. In 1968 Hultén stated that
var. pseudomyrsinites has “acute, long leaves and longer styles” and that
the variety seems to represent hybrid forms. | agree with his morphological
differences, but | know of no evidence that suggests that this taxon repre-
sents hybrid forms. Raup (1959) also commented on the binary nature of S.
myrtillifolia but noted that the taller plants were connected with the shorter
plants by a series of transitional forms. My studies have not led to the same
conclusion, and | am able to describe a combination of characteristics which
can be used to distinguish S. novae-angliae from S. myrtillifolia (Table 14).
The characteristics of plant height and leaf pubescence are usually diagnos-
tic and, when combined with style and stipule length, permit the identification
of all specimens. It is hoped that distinguishing these species will result in
eco-geographical information that will further improve our understanding of
their taxonomy.
There is a serious nomenclatural problem associated with S. novae-angliae
and S. myrtillifolia which can only be resolved by a study of the Drummond,
Bourgeau, Richardson and Burke material originally studied by Andersson
(1858), and by the typification of his taxa. | have seen one specimen, Drum-
mond 665, Rocky Mountains (NY), which was originally determined by An-
dersson as S. myrsinites L. and was probably a part of the original material
on which the inadmissible names S. myrsinites (ssp.) S. pseudomyrsinites
or S. myrsinites (ssp.) S. curtiflora were based. These names were later
changed to S. novae-angliae (ssp.) S. pseudomyrsinites and S. novae-angliae
(ssp.) S. pseudocordata, respectively (Andersson 1867). The Drummond
specimen is typical of what is usually called S. pseudocordata in western
Canada, a name which represents, at least in part, Andersson's S. novae-
angliae. Until further original material can be studied, it seems best to assume
that Andersson’s name, S. novae-angliae, circumscribes two recognizable
elements: one is S. novae-angliae (S. pseudocordata or S. myrtillifolia var.
pseudomyrsinites) and the other is S. myrti/lifolia. Until the nomenclatural
problems surrounding Andersson's intraspecific names can be resolved and
until his original specimens can be restudied, the recognition of S. novae-
angliae and S. myrtillifolia as species will emphasize their distinctness and
143
Plate XXIX
Salix pyrifolia Anderss. Pistillate specimen, Argus 7481, 9 VII 1969, Bi
4 : P , Big Country Pond, New-
foundland, GWA (left) and staminate specimen, Argus 6079 (right) :
144
Cordatae Barr. ex Hook.
Table 14 Comparison table: Salix novae-angliae and S. myrtillifolia
Characteristics novae-angliae myrtillifolia
habit tall shrubs, 1-4 m tall low, decumbent shrubs
less than 1 m tall
leaf pubescence juvenile and mature leaves glabrous
leaves pubescent at
least on upper midrib
leaf length 3.2-4.5-6.8 cm 2.5-5 cm
leaf colour paler beneath green to pale beneath
stipule length 1-5 mm minute to 1-2 mm
style length 0.5-0.9 mm 0.3-0.5 mm
will avoid some of the confusion associated with the names S. pseudocordata
and S. myrtillifolia var. pseudomyrsinites.
30. SALIX MYRTILLIFOLIA Anderss.
S. myrtillifolia Anderss. Ofvers. Vet. Akad. Forh. (Stockh.) 15: 132. 1858.
S. novae-angliae Anderss. (ssp.) S. myrti/lifolia Anderss. Kg. Sv. Vet. Akad.
Handl. 6: 162. 1867. S. novae-angliae Anderss. y (var.) myrti/lifolia Anderss.
in -DG.-Prodr. 16(2):-253. 1868.
Description of species
Low shrubs 1-9 dm tall, usually decumbent or trailing and rooting along
stem; branches greyish brown, glabrescent; branchlets greenish brown to
dark reddish brown, glossy, sparsely pubescent with short curved trichomes,
sometimes more or less glaucous; buds sparsely pubescent, sometimes glau-
cous. Leaves narrowly elliptic to narrowly obovate, the largest mature leaves
2-3.3-5.1 cm long, 0.9-1.5-2.2 cm wide and 1.7-2.2-3.6 times as long as
wide; apex broad, the sides forming an acute angle but the tip round; base
round or cuneate; margins glandular crenate to glandular crenate-serrulate;
immature leaves glabrous, sometimes reddish and translucent; the upper side
of mature leaves green, glossy and glabrous, the lower side pale green, glos-
sy and glabrous, the leaves sometimes blackening on drying; petioles 1.5-8
mm long, tawny and glabrous; stipules narrowly elliptic, 0.2-1.8 (3) mm
long, margins glandular, glabrous. Aments coetaneous, on short, leafy, florif-
erous branchlets. Staminate aments 1.3-3.5 cm long, floriferous branchlets
2-7 mm long; stamens 2, filaments about 2.5 mm long, glabrous; anthers
0.3-0.5-0.6 mm long. Pistillate aments 1.3-4.2 cm long, floriferous branch-
lets 3-10 mm long; pistils 3-3.5 mm long, glabrous and green, capsules 4-6
mm long, greenish or tawny; styles 0.3-0.5 mm long; stigmas about 0.2 mm
long, 2 horseshoe-shaped lobes; stipes 0.6-1.6 mm long, glabrous; nectaries
1, adaxial, 0.2-0.4 mm long, 0.1-0.5 times as long as stipe; bracts narrowly
145
Cordatae Barr. ex Hook.
165° 160°
Map 39 Salix myrtillifolia
oblong, apex rounded, 0.4-1.1 mm long, dark brown or bicolour, darker
brown to black at apex and tawny below, glabrous to sparsely pubescent
abaxially, sparely pubescent with long straight or curly trichomes adaxially.
Habitat Range
Muskegs; fens; wet lake margins; Boreal: Central Alaska; Mackenzie
wet river banks; subalpine Picea River Delta; southern half of Yukon
glauca-Betula papyrifera thickets. Territory; eastward across boreal
Canada to Newfoundland; south in
the Rocky Mountains to British Co-
lumbia and Alberta (Map 39).
Discussion
Salix myrtillifolia is characterized by low, decumbent shrubs trailing in moss;
leaves green on both sides, often drying brownish; very short styles (0.3-
0.5 mm long); and bracts sparsely pubescent and shorter than the capsule
bearing stipes. It is closely related to S. novae-angliae, which should be seen
for discussion (Table 14).
Hultén (1968) maps three localities, outside the range presented here
(Map 39), which | have not verified: Cape Beaufort, Okpilak River region and
146
Cordatae Barr. ex Hook.
Chignik. The first two records are highly suspect and are probably based on
misidentifications. The latter record from the Alaska Peninsula may be cor-
rect, but | have not seen any specimens of S. myrtillifolia from this region; if
the record is correct, it represents a significant range disjunction.
31. SALIX PYRIFOLIA Anderss.
S. pyrifolia Anderss. Kg. Sv. Vet. Akad. Handl. 6: 162. 1867.
S. balsamifera Barr. ex Anderss. Ofvers. Vet. Akad. Forh. (Stockh.) 5: 125.
1858.
Description of species
Shrubs 1-3 m tall; branchlets glabrous, shiny, dark reddish brown, rarely
greenish, drying black; buds and foliage with a balsam-like fragrance. Leaves
narrowly elliptic, narrowly ovate to ovate, the largest mature leaves 3-6
(8.5) cm long, 2-3.5 (4) cm wide, and 1.6-2.5 times as long as wide; apex
acute; base cordate to rounded; margins glandular serrulate on immature
leaves, becoming coarsely serrate or crenate in age; immature leaves mem-
branaceous and translucent, thinly pubescent or glabrescent, green on both
sides or thinly glaucous beneath; mature leaves subcoriaceous, opaque, the
lower surface reticulate veined and glaucous; petioles 7-15 mm long, pubes-
cent, sometimes glandular at the distal end; stipules small, caducous. Aments
coetaneous, on short, leafy, floriferous branchlets. Staminate aments 2-5 cm
long, floriferous branchlets 5-7 mm long; stamens 2, filaments glabrous or
pubescent at base. Pistillate aments loosely flowered, 2.5-6 (9) cm long,
floriferous branchlets 0.5-3 cm long; pistils glabrous, capsules 5-9 mm long,
spreading or reflexed; styles 0.5-1 mm long; stipes 2.5-4 mm long, nectary 1,
adaxial; bracts oblong, 1.5 mm long, tawny and pilose.
Habitat Range
Muskegs; fens; wet lake and slough Boreal: Known in Yukon Territory
margins. only from Palmer Lake on the west-
ern flanks of the Mackenzie Moun-
tains; western Northwest Territories;
northeastern British Columbia, east-
ward across boreal Canada to New-
foundland; south to northern New
York (Map 40).
Discussion
Salix pyrifolia is characterized by membranaceous and translucent imma-
ture leaves, subcoriaceous mature leaves with a reticulate lower surface and
often with a cordate base. The buds and foliage have a balsam-like fragrance.
This distinctive species is known in our area only from Palmer Lake in
northern Yukon. This record is a considerably northward extension of its
147
Plate XXX
>
Sa
%
nT
~*~ A
mud ©
mG
way
0 >
prt .
‘jz~®
we =.
ae
Salix bebbiana Sarg. Cody 8083 (left) and Argus 13-62 (right)
148
Vetrix Dumort.
known range. It should be sought elsewhere in the wet forests and fens
of the Yukon and northwestern Northwest Territories.
Section 12. Vetrix Dumort.
32. SALIX BEBBIANA Sarg.
S. bebbiana Sarg. Gard. and For. 8: 463. 1895. (Type: Richardson s.n. NY).
S. rostrata Richards. Bot. App. in Frankl. Journ. 753. 1823, non Thuill, 1799.
S. cinerascens (var.) occidentalis Anderss. Ofvers. Vet. Akad. Forh. (Stockh.)
15: 122. 1858. S. vagans (ssp.) cinerascens (var.) occidentalis Anderss. Proc.
Amer. Acad. Arts and Sci. 4: 16. 1858. S. vagans (ssp.) S. rostrata Anderss.
Kg. Sv. Vet. Akad. Handl. 6: 87. 1867. S. livida var. occidentalis Gray, Man.
Bot. 5th ed. 464. 1867. S. /ivida (var.) rostrata Dippel, Handb. Laubh. 2: 255.
1892. S. depressa L. ssp. rostrata Hiitonen, Mem. Soc. Faun. Fl. Fenn. 25:
82. 1950.
S. perrostrata Rydb. Bull. N.Y. Bot. Gard. 2: 163. 1901. (Type: Rydberg 1078,
NY). S. rostrata var. perrostrata Fern. Rhodora 16: 177. 1914. S. bebbiana
var. perrostrata Schneid. J. Arnold Arb. 2:71. 1920.
S. xerophila Flod. Bot. Not. 1930: 334. 1930.
S. bebbiana var. depilis Raup, Sargentia 6: 159. 1947. (Type: Raup & Soper
9306, A).
Description of species
Shrubs to small trees 0.5-10 m tall; branches divaricate, reddish brown,
pubescent to glabrescent; branchlets densely pubescent with straggly tri-
chomes. Leaves elliptic to obovate or narrowly ovate, the largest mature
leaves 2.6-6 (7.2) cm long, 1-2 (3) cm wide and (1.8) 2.2-3.5 (3.9) times
as long as wide; apex acute to obtuse, often conduplicate when pressed; base
round to obtuse, rarely acute; margins entire to crenate, slightly revolute; the
upper side of mature leaves pubescent with straight and wavy trichomes, or
glabrescent, the lower side sericeous-lanate with short wavy trichomes or
glabrescent, glaucous and rugose; petioles 2-9 (12) mm long, pubescent;
stipules deciduous, (0.4) 1-5.6 mm long. Aments coetaneous or subpreco-
cious, on short, leafy, floriferous branchlets. Staminate aments 0.6-1.5 cm
long, floriferous branchlets 1-3 mm long; stamens 2, filaments about 6 mm
long, glabrous and distinct; anthers 0.5-0.8 mm long. Pistillate aments
2.8-5 cm long, floriferous branchlets 3-15 mm long; pistils 5.5-7 mm long,
long-beaked, greenish, rarely reddish, sericeous, capsules 5.5-8.5 mm long,
glabrescent to sparsely pubescent; styles 0.1-0.4 mm long; stigmas 0.4-
0.7 mm long, 2.4-lobed; stipes 2.8-4.8 mm long, pubescent; nectaries 1,
adaxial, 0.4-0.5 mm long, about 0.1 times as long as stipe; bracts narrowly
oblong, apex acuminate, 1.2-3.2 mm long, tawny, rarely reddish at apex,
sparsely pubescent becoming glabrescent.
149
Vetrix Dumort.
Map 40 Salix pyrifolia
150°
Sg
145°
Habitat
Riverine and upland forests with Picea
glauca, Betula papyrifera, Populus
tremuloides; wet lowland thickets;
Picea mariana muskegs; prairie mar-
gins; dry south-facing slopes; dis-
turbed areas, such as roadsides and
burns.
Range
Boreal, temperate: Central Alaska;
eastern Alaska Peninsula; Kodiak
Island; Kenai Peninsula; absent in
Pacific coastal Alaska from Prince
William Sound to southeastern Alas-
ka; southern half of Yukon Territory
and adjacent Northwest Territories;
transcontinental in boreal and temper-
ate Canada; south in the cordillera
to Arizona and New Mexico; eastward
across northern United States; Eur-
asia: Kola Peninsula to Chukotsk Pen-
insula (Map 41).
Discussion
Salix bebbiana is one of the most common willows in the forested part of
Alaska and the Yukon. It is characterized by rugose leaves with entire to
crenate margins, deciduous stipules, pistils on very long stipes (2.8-4.8 mm
150
Vetrix Dumort.
° 120°
D
125
Map 41 Salix bebbiana. Circle based on Skvortsov, 1966, as S. xerophila
long), nectaries about 0.1 times as long as the stipes, and tawny bracts.
There is considerable variation in leaf and branchlet pubescence and in leaf
rugosity within S. bebbiana. This variation has provided the basis for the
description of a series of progressively more glabrescent taxa, from the pubes-
cent and rugose-leaved var. bebbiana to the glabrescent, plane-leaved var.
perrostrata to the almost glabrous var. depilis. In the material that | have
studied from Alaska and the Yukon, about 60 per cent are of the pubescent
form and 40 per cent of the glabrescent form, with numerous intermediates.
In the opinion of many authors, including Hultén (1943), Raup (1959), and
Argus (1964), these variants are not worthy of taxonomic recognition and are
considered here to be synonyms of S. bebbiana.
In 1950 Hiitonen decided, without presenting adequate documentation, that
S. bebbiana should be regarded as a subspecies of its European relative
and proposed the name S. depressa ssp. rostrata. In 1967 Hultén concurred
in this judgement, also without providing explanation or justification. The
proposal of this combination is unfortunate, for it further compounds a no-
menclatural tangle which must be resolved. Linnaeus described S. depressa
in 1753 and compared it with S. caprea; in the second edition of Species
Plantarum (1763) he modified this view and decided that it was a variety of
S. lanata (Skvortsov 1966). According to Enander (1907) the type material
151
Plate XXXI
Salix scouleriana Barr. Argus 6086 (left) and Argus 6840 (right)
152
Vetrix Dumort.
of Linnaeus’ S. depressa is a mixture of S. caprea and some hybrid form
(Schneider 1917). It was Fries (1832) who used the name S. depressa to
refer to the taxon under consideration. However, since he did not see the
Linnaean type material, Skvortsov (1966) argues that we have no more rea-
son for accepting Fries’ view than that Linnaeus. In 1806 S. starkeana was
described by Willdenow. This was followed by the description of two addi-
tional, but synonymous, names, S. /ivida Wahlenberg (1812) and S. vagans
Andersson (1858). The name S. starkeana was accepted for the European
taxon by Rechinger in Flora Europea (1964) and by Skvortsov in Flora
Arctica URSS (1966). In addition, they both recognized a second taxon with
a northern European-Asian distribution, S. xerophila, which differs from S.
starkeana in that it has more pubescent leaves and branchlets, longer petioles
(6-8 mm long) and pubescent filaments (Rechinger 1964). Skvortsov ob-
serves that these two taxa are connected by transitional forms, but prefers to
treat S. xerophila as a separate species. He notes further that S. xerophila
is very closely related, if not identical, to S. bebbiana — an observation with
which | am in complete agreement. | have examined material of S. xerophila
from Asia (in CAN and US) and | cannot distinguish it from the North Amer-
ican S. bebbiana. | therefore suggest that these names be treated as syn-
onyms. However, this suggestion would not solve the problem of the correct
application of S. depressa or of the taxonomic relationship between S.
bebbiana and S. starkeana. | am inclined to recognize S. starkeana and S.
bebbiana as distinct species in conformity with some recent taxonomic
treatments (Rechinger 1964; Skvortsov 1966, 1968).
33. SALIX SCOULERIANA Barr.
S. scouleriana Barr. in Hook. Fl. Bor.-Am. 2: 145. 1838. (Type: Scouler s.n.
photo and frag. A).
S. brachystachys var. scouleriana Anderss. in DC. Prodr. 16 (2): 224. 1868.
S. flavescens var. scouleriana Bebb, Bot. Gaz. 7: 129. 1882.
S. flavescens Nutt. N. Am. Sylva 1: 65. 1843, non Host, 1828. S. nuttallii
Sarg. Gard. and For. 8: 463. 1895. S. scouleriana var. flavescens Henry, FI.
30. B:G; 98: 1915.
S. stagnalis Nutt. N. Am. Sylva 1: 66: 1842.
S. brachystachys Benth. Pl. Hartw. 336. 1857. S. scouleriana var. brachy-
stachys M.E. Jones, Willow Fam. 15. 1908.
S. capreoides Anderss. Ofvers. Vet. Akad. Forh. (Stockh.) 15: 120. 1858. S.
flavescens var. capreoides Bebb, Gard. and For. 8: 373. 1895. S. nuttallii
var. capreoides Sarg. Gard. and For. 8: 463. 1895.
S. scouleriana var. poikila Schneid. J. Arnold Arb. 2: 12. 1920. S. scouleriana
f. poikila Schneid. J. Arnold Arb. 3: 76. 1922.
153
Vetrix Dumort.
S. scouleriana var. coetana Ball, J. Wash. Acad. Sci. 24: 73. 1934. (Type:
Thompson 9297, A).
S. scouleriana var. thompsoni Ball, J. Wash. Acad. Sci. 24: 75. 1934.
Description of species
Shrubs 2-7 m tall or trees 10-20 m tall and up to 7.2 dm in diameter at base;
branches dark reddish brown to yellow brown, glossy, usually remaining
pubescent for 2-3 years; branchlets greenish brown to brown, velutinous to
villous-lanate. Leaves obovate, elliptic to narrowly elliptic, the largest mature
leaves 5-6.5-8 cm long, (1.3) 2-3 cm wide and (2.1) 2.4-3.3 (4.9) times as
long as wide; apex acute to round with acute tip, acuminate in narrower
leaves; base cuneate; margins usually entire and distantly glandular, to ir-
regularly glandular serrulate or subcrenate, revolute; immature leaves velu-
tinous to villous-lanate; the upper side of mature leaves pubescent, becoming
glabrate, dark green and glossy, the lower side sericeous, sometimes sparse-
ly so, with appressed white and/or ferruginous trichomes, sometimes sparsely
to densely white villous, glaucous; petioles 5-10 mm long, 8-19 mm long on
vigorous leaves, tawny and velutinous becoming reddish brown and sparsely
pubescent; stipules minute to 0.8-3.5 mm long, 3.2-10 mm long on vigorous
shoots, half ovate, glandular serrulate, dark green and glossy above, pubes-
cent beneath. Aments precocious, often on short, leafy or bracteate, florif-
erous branchlets. Staminate aments 1.5-4 cm long, floriferous branchlets
0-3-7 mm long; stamens 2, filaments 5-6 mm long, glabrous to very sparsely
pubescent at base; anthers 0.5-1 mm long. Pistillate aments 1.5-5 cm long,
floriferous branchlets 0.2-2.5-15 mm long; pistils 4-8.5 mm long, grey green,
densely sericeous, capsules 4.5-11 mm long, beak long and slender, tawny,
sericeous, coma surrounding seeds often rusty coloured; styles 0.2-0.5 mm
long; stigmas 0.6-1.2 mm long, 2-lobed, about 2 times as long as style; stipes
0.8-2 mm long, sericeous; nectaries 1, adaxial, 0.2-0.7 mm long, 0.33-0.5 times
as long as stipe; bracts narrowly elliptic, apex acuminate, 2-4.5 mm long,
dark brown to black, sometimes bicolour, villous on both sides with long
straight trichomes about 2 times as long as bract.
Habitat
Dry Pinus contorta and Picea mariana
forests; muskegs; mature woods at
edges of rivers and _ lakes; Salix
thickets; meadows; disturbed areas.
Discussion
Range
Boreal: Central Alaska; Kenai Penin-
sula; southeastern Alaska; Yukon Ter-
ritory; southward to British Columbia,
California, and New Mexico; eastward
in the boreal forest to Manitoba and
South Dakota (Map 42).
Salix scouleriana is a precocious flowering species characterized by veluti-
nous branchlets, usually obovate leaves with appressed, white and ferrugi-
nous trichomes or densely white villous beneath; pistils long-beaked and
stigmas about 2 times as long as the styles.
154
Vetrix Dumort.
165° 160° 155° 150° 120°
possi ts 7 P= GLME
Map 42 Salix scouleriana
This species is part of a complex including the eastern North American S.
discolor Muhl. and S. humilis Marsh., the Eurasian S. caprea L. and the Eu-
ropean S. coaetanea Flod. It is possible that, as Cronquist (1964) states,
‘““Monographic study may show these [S. discolor and S. caprea] to be geo-
graphic phases of a single species. . . .'’ However, | would urge the requisite
‘“monographic study” be made before any new combinations are proposed. A
disjunction exists in western Alaska and eastern Siberia between the ranges
of S. caprea and S. scouleriana, so that evidence of intergradation will not be
readily available and the taxa have different chromosome numbers; S. caprea
is 2n = 38 (Blackburn and Heslop-Harrison 1924) and S. scouleriana is
2n = 76 (Suda and Argus 1968) or 2n = ca. 114 (Taylor and Mulligan 1968).
The pubescence on the underside of mature leaves is of two types. The
most common type consists of glossy, sericeous pubescence composed of ap-
pressed white and ferruginous trichomes. The ferruginous trichomes are very
conspicuous and give the leaves their characteristic ‘‘rusty’’ appearance. It
has been observed that some ferruginous trichomes are glandular and that a
reddish drop may form at their base, or that several trichomes may be matted
together. Not all ferruginous trichomes are glandular and this observation
requires further study. The second type consists of densely villous pubes-
cence composed of curly, white trichomes and is the basis of the name var.
155
Plate XXXIl
Salix planifolia Pursh ssp. planifolia. Flowering pistillate specimen, Calder 24404 (left) and
fruiting pistillate specimen, Argus 5052 (right)
156
Arbuscella Ser. ex Duby
Table 15 Comparison table: Salix scouleriana and S. sitchensis
Characteristics scouleriana sitchensis
leaves glaucous beneath non-glaucous beneath
branchlets tenacious and more brittle and densely
sparsely pubescent sericeous
stipule length 0.8-3.5(10) mm minute to 0.4-1.5 mm
stamen number 2 1
style length 0.2-0.5 mm 0.5-0.8 mm
stigma length 0.6-1.2 mm 0.1-0.2 mm
poikila. This type of pubescence commonly occurs on leaves of sprout shoots
or on shoots borne on this year’s branchlets, and both pubescence types may
occur on the same plant. Leaves with the densely white villous pubescence
resemble S. sitchensis, and confusion may occur if only vegetative material
is available. The two species may be distinguished as in Table 15.
Section 13. Arbuscella Ser. ex Duby
34a. SALIX PLANIFOLIA Pursh ssp. PLANIFOLIA
S. planifolia Pursh, Fl. Am. Sept. 2: 611. 1814.
S. phylicifolia L. ssp. planifolia Hiitonen, Mem. Soc. Faun. Fl. Fenn. 25: 82.
1950.
S. chlorophylla Anderss. Kg. Sv. Vet. Akad. Handl. 6: 138. 1867. (possibly
S. discolor, at least in part).
S. monica Bebb, in S. Wats. Bot. Calif. 2: 90. 1879. (Type: Bolander s.n. F).
S. planifolia var. monica Jeps. Man. Fl. Pl. Calif. 265. 1923. S. chlorophylla
var. monica Flod. Sv. Vet. Akad. Ark. Bot. 29A: 33. 1939.
S. nelsonii Ball, Bot. Gaz. 40: 379. 1905. (as nelsoni). (Type: Nelson 7580,
NY). S. chlorophylla var. nelsonii Flod. Sv. Vet. Akad. Ark. Bot. 29A: 34.
1939. S. planifolia var. nelsonii Ball ex E. C. Smith, Amer. Midl. Nat. 27:
246. 1942.
S. pennata Ball, Bot. Gaz. 60: 45. 1915. (Type: Suksdorf 15, US). S. phylici-
folia var. pennata Cronquist, in Hitchcock, Cronquist, Ownbey and Thompson,
Vas. Pl. Pacific NW. 2: 63. 1964.
Description of species
Shrubs 1-4 m tall; branches dark brown to reddish brown, glabrescent, some-
times glaucous, epidermis exfoliating; branchlets brownish to greenish brown,
glossy, glabrous or densely villous to sparsely pubescent, sometimes glauces-
cent. Leaves elliptic to narrowly elliptic, the largest mature leaves 3.5-4.9
157
Arbuscella Ser. ex Duby
(6.5) cm long, 0.9-1.5 (2.2) cm wide and 3-4.7 times as long as wide; apex
acute; base acute; margins subentire to glandular crenate, revolute especially
near base; immature leaves reddish, sparsely pubescent or densely sericeous
with white or white and ferruginous trichomes, non-glaucous at first but
becoming progressively glaucous beneath from the apex; the upper side of
mature leaves glabrescent, highly glossy and with impressed venation, the
lower side sparsely pubescent with white or white and ferruginous trichomes,
glaucous; petioles 3-7 (13) mm long, reddish or yellow; stipules narrowly
elliptic, 0.8-2.8 mm long, margins glandular, glabrous. Aments precocious,
sessile on branches or with 2-3 brown or greenish bracts at base. Staminate
aments 1.5-3 (5) cm long; stamens 2, filaments about 7-8 mm long, gla-
brous; anthers 0.5-0.6 mm long. Pistillate aments 1.5-6 cm long; pistils
2-2.8 mm long, tawny, densely sericeous with white and/or ferruginous
trichomes, capsules (3.2) 5.5-6 mm long, tawny or greenish, sparsely seri-
ceous; styles 0.6-1.8 mm long; stigmas 0.4-0.5 (0.8) mm long, 2-4 linear
lobes; stipes 0.5-0.6 mm long, pubescent; nectaries 1, adaxial, 0.5-0.6 mm
long, equal to or up to 3 times as long as stipe; bracts oblong, apex acute
to rounded, 1.2-2 mm long, black to dark brown, pubescent on both sides
with long straight trichomes about 2 times the length of the bract or rarely
with shorter, curly trichomes.
Habitat Range
Common in Salix thickets and in Boreal, Arctic: Southern Yukon Ter-
Carex fens at edges of streams and ritory, adjacent Northwest Terri-
lakes; Picea mariana muskegs; open- tories and British Columbia; east-
ings in P. glauca forests; arctic tun- ward through the boreal forest and
dra in eastern Canada. the southern Canadian Arctic to
Newfoundland; southward in eastern
British Columbia to Alberta, Califor-
nia, and New Mexico; eastward to
northeastern United States (Map 43).
Discussion
Hultén’s report (1968, Map 47) of Salix planifolia ssp. planifolia (as S.
phylicifolia ssp. planifolia) from Alaska and the Mackenzie Delta region
conflicts with the maps presented here. The Alaska records may be based on
misidentified material of ssp. pu/chra. However, it is possible that ssp. p/ani-
folia occurs in southeastern Alaska, between the Chugach Mountains and the
Alaska Range, and should be sought there. | have seen extensive collections
of S. planifolia from the Mackenzie Delta region and they are all referable
to ssp. pulchra. °
158
Arbuscella Ser. ex Duby
170°
120°
Map 43 Salix planifolia ssp. planifolia
34b. SALIX PLANIFOLIA ssp. PULCHRA (Cham.) Argus var.
PULCHRA
. planifolia ssp. pulchra (Cham.) Argus, Can. J. Bot. 47: 798. 1969.
. pulchra Cham. Linnaea 6: 543. 1831.
. phylicoides Anderss. Ofvers. Vet. Akad. Forh. (Stockh.) 15: 123. 1858.
S
S
S
S. fulcrata Anderss. 8 subglauca Anderss. in DC. Prod. 16(2): 244. 1868.
S. phylicifolia var. subglauca Boivin, Natur. can. 93: 437. 1966.
S. pulchra var. looffiae Ball, Madrofio 6: 228. 1942. (Type: Looff 1197, US).
S. pulchra var. palmeri Ball, Madrofio 6: 229. 1942. (Type: Palmer 121, US).
S. parallelinervis Flod. Sv. Vet. Akad. Ark. Bot. 20A: 35. 1939. S. pulchra
ssp. parallelinervis A. Skvortsov Bull. Mosk. Obshch. Isp. Pri. Biol. 66: 31.
1961.
S. pulchra ssp. kalarica A. Skvortsov Bull. Mosk. Obshch. Isp. Pri. Biol. 66: 31.
1961. (probably = ssp. pulchra).
159
Plate XXXIll
Salix planifolia Pursh ssp. pulchra (Cham.) Argus var. pulchra. Pegau 94-68 (left) and
Suda 248-66 (right)
160
Arbuscella Ser. ex Duby
Description of subspecies
Low shrubs prostrate to 0.1-1.8-3 m tall; branches glossy; branchlets rarely
glaucescent; buds sometimes large and beaked. Leaves sometimes marces-
cent, commonly rhombic but often narrowly elliptic, elliptic or sometimes
obovate, the largest mature leaves (2.2) 3.2-6 (7.5) cm long, (0.8) 1-2.3
(2.6) cm wide and (1.7) 2-3.6 (4.7) times as long as wide; apex acute to
obtuse, sometimes acuminate; base acute to cuneate; margins entire or sub-
entire sometimes glandular or glandular crenate; immature leaves glabrous or
sparsely pubescent with white or ferruginous trichomes especially on midrib
and toward margins; the upper side of mature leaves dark green and gla-
brous, the lower side glabrous; petioles 2.8-8-10 (15) mm long, glabrous or
sparsely pubescent adaxially; stipules linear (2.4) 3.6-14 (32) mm long,
equalling or exceeding petiole, distantly glandular serrulate, glabrous, glau-
cous abaxially, often persistent for 2-4 years. Anthers 0.4-0.6-0.8 mm long.
Pistillate aments 1.2-2.5 cm long in flower, 2.5-8 cm long in fruit, florif-
erous branchlets absent but rachis sometimes elongated and bracteate with
several dark brown or, rarely, green bracts; stipes 0.2-0.8 mm long; necta-
ries 0.4-1.2 mm long, yellow or reddish, 2-3 times as long as stipe.
Habitat Range
Common throughout arctic and alpine Arctic, subarctic, alpine: General
tundra; open Picea mariana-lichen throughout Alaska; uncommon in
woodlands; Salix-Betula thickets on Yukon Territory and the Tanana River
stream and lake margins; open Picea lowlands; absent from Aleutian Is-
glauca-Betula glandulosa woodlands. lands and southern coastal Alaska
from Prince William Sound eastward;
Asia: widespread from Novaya Zemlya
to Chukotsk Peninsula (Map 44).
34c. SALIX PLANIFOLIA ssp. PULCHRA var. YUKONENSIS
(Schneid.) Argus
S. planifolia ssp. pulchra var. yukonensis (Schneid.) Argus, Can. J. Bot. 47:
800. 1969.
S. pulchra var. yukonensis Schneid. J. Arnold Arb. 1: 72. 1919. (Type: East-
wood 373, A).
S. anadyrensis Flod. Sv. Vet. Akad. Ark. Bot. 25A: 9. 1933. S. pulchra var.
anadyrensis A. Skvortsov, Bull. Mosk. Obshch. Isp. Pri. Biol. 66: 30. 1961.
Description of variety
Differs from var. pulchra in shrubs 0.6-1.8-3 (4.5) m tall; branches gla-
brescent or with patches of villous indumentum, sometimes with persistent
pubescence on 3-year-old stems; branchlets light brown, densely white grey
161
Arbuscella Ser. ex Duby
115°
65°
oe sian este ~ - - ce
Map 44
pulchra
Salix planifolia ssp. pulchra var. pulchra. Circles based on Skvortsov, 1966, as S.
villous to sparsely pubescent, indumentum of white or sometimes white and
ferruginous trichomes. Mature leaves usually pubescent on midrib with white
or ferruginous trichomes; petioles persistently villous-tomentose with white
and ferruginous trichomes; stipules glabrous or pilose.
Habitat
Salix thickets at edges of lakes and
streams; Picea mariana muskegs;
margins of Carex fens; Betula papyri-
fera-Alnus crispa woods; shrub-heath
alpine tundra.
Range
Boreal, alpine: Yukon, Kuskokwim and
Tanana Rivers and their tributaries;
Umiat; northern Yukon to Anderson
River, Northwest Territories; southern
Yukon and adjacent British Columbia;
Asia: (as var. anadyrensis) middle
Indigirka River; middle and lower
Kolmia, Anadyr, and Penzhina Rivers
(Skvortsov 1961) (Map 45).
Discussion
In North America Salix planifolia consists of two races, a transcontinental
ssp. planifolia and a northwestern ssp. pulchra. The races overlap and inter-
162
Arbuscella Ser. ex Duby
Map 45 Salix planifolia ssp. pulchra var. yukonensis
grade in southern Yukon and western Northwest Territories. These races are
related to the European S. phylicifolia and to S. bicolor, which is restricted to
the mountains of Europe from central Germany to northern Spain and Bul-
garia, but is absent from the Alps (Rechinger 1964). The areas of ssp. pulchra
and S. phylicifolia overlap in northern Russian between the Yenisei and
Pechora Rivers (Skvortsov 1966, Maps 30 and 31). Within the Asian S. pulchra
Skvortsov (1961) recognized two subspecies, ssp. paral/l/elinervis and ssp.
kalarica and one variety, var. anadyrensis. | have seen specimens of the
variety that were determined by Skvortsov and | consider them to be equi-
valent to var. yukonensis. The ssp. parallelinervis, which is characterized by
smaller stipules, the presence of a short floriferous branchlet and lateral veins
of leaves paralleling the margin, seems to me to be confluent with ssp. pu/chra
and, at the most, deserving of varietal status; however, no new combination
is proposed here. Similarly, ssp. ka/arica, which occurs between the Stanov-
highlands and Dshugdshuria and is characterized by large leaves which are
pilose beneath, small stipules and yellow shoots, could be assigned varietal
status. | have not seen specimens of this taxon and therefore no new combi-
nation is proposed.
The relationship between the North American S. p/anifolia and the Europe-
an S. phylicifolia requires detailed consideration. There are three facets to
163
Arbuscella Ser. ex Duby
the question: the first is the uncritical proposal of new taxa; the second is
the failure to recognize the relationship between S. planifolia and S. pulchra;
the third is a consideration of differences between S. pl/anifolia and S. phyli-
cifolia.
In 1950 Hiitonen proposed the new combination S. phylicifolia ssp. plani-
folia, giving as an explanation Heribert-Nilsson’s observation that S. phylici-
folia consisted of a number of diverse forms, some of which diverge more
from S. phylicifolia than does the North American S. planifolia, and then con-
cluding that the North American taxon must therefore be a subspecies. Un-
fortunately, this decision to form a new combination was not based on a
thorough study of the problem, and once a new name is in the literature, it
assumes an importance of its own and may be accepted by later authors who
assume that the combination was the result of careful study. Cronquist
(1964) and Hultén (1967 and 1968) have accepted the Hiitonen name, and
others, including Boivin (1967) and Breitung (1957), came to this conclusion
independently. However, no one has presented evidence in support of his
opinion. The willingness to accept these taxa as conspecific was probably
aided by Raup’'s citation (1959) of two specimens reputedly representing S.
phylicifolia, one from Pine Creek, Yukon, and a second from Aniak, Alaska.
| have restudied the first specimen (Raup & Raup 11785, ALA 20026) and it
is simply S. glauca; the second specimen was not available at the Gray Her-
barium and | cannot comment on it. As far as | can determine, there are no
authentic specimens which support the occurrence of S. phylicifolia in North
America.
In the European literature S. phylicifolia and S. pulchra are generally con-
sidered to be distinct species, distinguishable mainly on the basis of the very
prominent, linear stipules of S. pu/chra. Rechinger (1964) separated the two
on the basis of habit and habitat with S. pu/chra included with, “low shrubs
of Arctic or high mountains; catkins terminal or sub-terminal’’ and S. phyli-
cifolia included with, “trees and shrubs usually of lower latitudes or lower
altitudes; catkins lateral’. Furthermore, he includes S. pu/chra with S. arctica
in subgenus Salix and S. phylicifolia in subgenus Caprisalix, a separation
which is clearly incorrect and which has tended to obscure the relationship
which evidently does exist. Skvortsov (1961) does comment that within the
area of overlap between S. phylicifolia and S. pulchra intermediate types do
occur, but he was able to distinguish the two on the basis of stipule size and
did not see fit to combine the two taxa. However, he did include them in the
same section (Arbuscella Duby).
In North America the morphological extremes of ssp. p/anifolia and ssp.
pulchra may be distinguished on the basis of characteristics included in
Table 16. In southern Yukon and western Northwest Territories there are
intermediates which cannot be assigned to one taxon or the other, and when
stipule length is compared with petiole length in a large number of speci-
mens, the overlap and intergradation are graphically evident (Fig. 5). While
the generalization holds that the absolute and relative stipule lengths are diag-
nostic, there are also many exceptions. On the Haines Highway in northwest-
ern British Columbia, in the region between the Three Guardsmen Pass and
the Chilkat Pass, there is a series of intergrading populations of ssp. plani-
folia and ssp. pulchra in which considerable variation in stipule size and
164
Arbuscella Ser. ex Duby
15 ss
OO
O © ©
Sue €, ®
10 ONO <6 © @ eo
oo @ Oe Q e Oe © 8
OO gi @aHdO5ex ed e ee
= 2 08 PO. ee
= oe ®. ®. ? ® & te ©
= © as r
eae, OOo © OMeyoli an, meas aM
= e °@ e e e@
a4 O ee 60 @ e
53 i) ‘4 @ & : & .
g
| 2 3 4 5 10 15
Stipule length (mm)
Fig. 5 A comparison of stipule and petiole Generally, ssp. planifolia and ssp. pulchra var.
length variation in Salix planifolia ssp. plani- pulchra can be distinguished on the basis of
folia (circles), ssp. pulchra var. pulchra_ stipule length, but there is some overlap
(dots) and ssp. pulchra var. yukonensis which is further accentuated by var. yukonen-
(diamonds). In these taxa there is no correla- sis, which may have stipules 2-3 mm long.
tion between stipule length and petiole length. The scales are logarithmic.
presence is manifest. There are ‘typical’ ssp. pu/chra with large, linear
stipules (Argus & Chunys 6786, 6761), others with one large, linear stipule
per leaf or none (6879), others without stipules in 1967 but with old, per-
sistent stipules from 1966 (6792), and yet others that are exstipulate (6790,
6771). The latter specimens are indistinguishable from ssp. planifolia except
that the immature leaves are not as prominently pubescent as is usually the
case in this taxon. Such diversity strongly suggests a hybrid swarm, which is
further evidence of the closeness of the relationship between these taxa.
The patterns produced by a paper chromatographic analysis of crude leaf
extracts (largely phenolic compounds) are very similar, and both taxa have
165
Arbuscella Ser. ex Duby
q)
5 6 7
Fig. 6 A representative chromatographic pat- (2) moderately absorbing: yellow, (3) inten-
tern produced by paper chromatography of ely absorbing: intensely absorbing, (4)
crude methanolic-HCI leaf extracts in Salix faintly absorbing: pink, (5) faintly absorbing:
planifolia ssp. planifolia and ssp. pulchra. moderately absorbing, (6) pink: pink, (7)
Colour code — (Ultraviolet light: Ultraviolet moderately absorbing: absent.
light after fuming with NH.) — (1) blue: blue,
the very prominent absorbing spot near the centre (Fig. 6). Also, the chromo-
some number is the same for both taxa, 2n = 76 (Suda and Argus 1968,
1969; Love and Lé6ve 1964). The report by Love (1954) of 2n = 152 for
Salix planifolia ssp. planifolia is not included here because it conflicts with
later reports (including numerous unpublished counts by Suda) and because
the voucher specimen is unavailable for verification of the identification.
Thus far | have demonstrated that ssp. p/anifolia and ssp. pulchra are so
closely related that they intergrade within the area of overlap and may be
regarded as races of the same species. The next question is, what is the
relationship of these taxa to S. phylicifolia. In the first place it is significant
that European taxonomists have long regarded these two taxa as distinct and
that some would even place them into different subgenera (Rechinger 1964).
166
Arbuscella Ser. ex Duby
Table 16 Comparison table: Salix planifolia ssp. planifolia and ssp. pulchra
Characteristics ssp. planifolia ssp. pulchra
stipules narrowly elliptic, linear,
0.8-2.8 mm long, (2.4) 3-6-14 (32) mm
shorter than long, generally
the petioles equalling or exceeding
the petioles
immature leaves sparsely to densely sparsely pubescent or
sericeous glabrous
Table 17 Comparison table: Salix planifolia ssp. planifolia, ssp. pulchra and S. phylicifolia
Characteristics ssp. planifolia ssp. pulchra phylicifolia
floriferous absent or the rachis absent or the rachis 6-20 mm long,
branchlets may be elongated may be elongated leafy
and bare or with and bare or with
2-3 green bracts 2-3 green bracts
leaf subentire to entire to subentire, glandular serrulate
margins glandular crenate rarely glandular or
glandular crenate
stipule length 0.8-2.8 mm (2.4) 3.6-14 (32) mm absent to minute,
. up to 3 mm long
petiole length 3.7 (t3))mm 2.8-8-10 (15) mm 6-15 mm
Secondly, the chromosome number of S. phylicifolia is 2n = 114 (Federova-
Sarkissova 1946; Hakansson 1933; Heribert-Nilsson 1935) and 2n = 88
(Blackburn and Heslop-Harrison 1924; Wilkinson 1944). This is in con-
trast to 2n = 76 for ssp. planifolia and ssp. pulchra. Finally, morphologically
these taxa may be distinguished on the basis of a number of characteristics
(Table 17). Admittedly, these characteristics are variable and exceptions are
not unknown. However, since there are some characteristics which can serve
to distinguish these taxa, and since there is a difference in chromosome num-
ber, there does seem to be ample reason for considering them as two species,
at least until the group can be studied monographically.
Salix planifolia ssp. pulchra var. yukonensis can be distinguished from var.
pulchra on the basis of a number of characteristics which were previously
outlined, the most important of which is the densely white grey villous to
sparsely pubescent branchlets. Similar variation in branchlet pubescence
occurs in ssp. planifolia, but it lacks the geographical integrity of var. yuko-
nensis and has therefore not been recognized taxonomically. To my knowledge
there is no evidence that var. yukonensis is a hybrid or is influenced by hy-
bridization, as suggested by Hultén (1968).
There are several varieties which have been based on differences in leaf
shape, including var. /ooffiae and var. palmeri. However, in the light of the
wide variation in leaf shape, often the product of the environment, these
varieties do not deserve taxonomic recognition.
167
Plate XXXIV
Salix arbusculoides Anderss. (A) Pistillate specimen, Ar i
, Argus 4753, (B) Vegetative specimen,
Viereck 8054, (C) Staminate specimen, Viereck 7892, (D) Leaf, Viereck 8054
168
Arbuscella Ser. ex Duby
35. SALIX ARBUSCULOIDES Anderss.
S. arbusculoides Anderss. Kg. Sv. Vet. Akad. Handl. 6: 147. 1867.
S. arbusculoides var. glabra Anderss. Kg. Sv. Vet. Akad. Handl. 6: 148. 1867.
S. humillima Anderss. in DC. Prodr. 16 (2): 248. 1868.
Description of species
Shrubs 1-4 m tall or trees 5-6 m tall, basal diameter up to 2.8 dm; branches
slender, reddish brown, glossy, sometimes thinly glaucous, epidermis exfoliat-
ing; branchlets sparsely velutinous, indumentum of short erect or sometimes
curly trichomes. Leaves narrowly ovate to very narrowly ovate or elliptic, the
largest mature leaves (3.8) 6-7 (7.8) cm long, (0.7) 1-1.5 (1.8) cm wide
and 3-6.5 times as long as wide; apex acute to narrowly acute or obtuse;
base acute; margins glandular serrulate, sometimes distantly so; the upper
side of mature leaves glossy and glabrous, the lower side sericeous with short,
white or rarely ferruginous trichomes appressed and oriented toward apex,
rarely glabrescent; petioles (3) 5-8 (11) mm long, reddish or yellowish; stip-
ules 0.9-2.4 mm long, margins revolute, glandular, a white secretion often
accumulating and drying at the base. Aments coetaneous or subprecocious,
subsessile or on short, floriferous branchlets. Staminate aments 1.8-2.5 cm
long, floriferous branchlets about 2 mm long and bearing several bract-like
leaves; stamens 2, filaments about 3-4.8 mm long, glabrous, distinct; anthers
about 0.5 mm long. Pistillate aments 2-7.5 cm long, loosely flowered with
gaps between the “whorls” or densely flowered, floriferous branchlets 0-5
mm long; pistils 3-4.5 mm long, sericeous, capsules 4-5 mm long, sparsely
sericeous with white and ferruginous trichomes; styles 0.3-0.5 mm _ long;
stigmas 0.2-0.4 mm long, 4-lobed; stipes 0.6-0.9 mm long, velutinous;
nectaries 1, adaxial, 0.6-0.8 mm long, rod-like, equal to or slightly exceed-
ing stipe; bracts oblong, apex obtuse or rounded, 0.8-1.2 mm long, pubes-
cent on both sides with curly or long straight trichomes.
Habitat Range
Stream banks; openings in Picea
glauca-Betula forests; muskegs; Salix
thickets at the edge of arctic and
alpine tundra.
Boreal: Common throughout central
Alaska, Yukon Territory and adjacent
Northwest Territories; not reaching
the Pacific Coast; extending east-
ward throughout the boreal forest
to Hudson Bay; southward in the
eastern Rocky Mountains to north-
eastern British Columbia and Alberta
(Map 46).
169
Plate XXXV
Salix lanata L. ssp. richardsonii (Hook.) A. Skvortsov. (A) Flowering specimen, Little 18248,
(B) Argus 5192, (C) Argus 616
170
Lanatae Koehne
Discussion
Salix arbusculoides is a common forest species characterized by narrowly
elliptic leaves, distinctly glandular serrulate on the margins, glabrous above
and sericeous beneath with short, straight, white or ferruginous trichomes
oriented toward the apex. Occasional specimens have glabrescent or glabrous
leaves and have been named var. glabra Anderss., but the taxonomic signif-
icance of this variation is doubtful. The styles are very short and the necta-
ries equal or slightly exceed the short stipes.
The Siberian Salix boganidensis Trautvetter (1847) seems to be closely
related to S. arbusculoides, and further study may show that they are con-
specific.
Section 14. Lanatae Koehne
36. SALIX LANATA L. ssp. RICHARDSONII (Hook.) A. Skv.
S. lanata L. ssp. richardsonii (Hook.) A. Skvortsov, in Tolm. Fl. Arct. URSS
5: - 103..1966.
S. richardsonii Hook. Fl. Bor.-Am. 2: 147. 1838.
Description of subspecies
Shrubs 0.6-3 (7) m tall; branches reddish brown, glossy, pubescent with
persistent, greyish, coarse, spreading or matted indumentum; branchlets
densely white lanate. Leaves elliptic, narrowly ovate to broadly obovate, the
largest mature leaves (3) 3.8-6.8 (8) cm long, (1.1) 1.5-4 (7.3) cm wide
and 1.1-2-2.8 times as long as wide; apex acute; base obtuse or acute to
rounded, rarely inequilateral; margins entire or subentire to glandular ser-
rulate or irregular crenate; immature leaves sometimes pubescent with fer-
ruginous trichomes; the upper side of mature leaves glabrous or sparsely
villous, the lower side glabrous or sparsely villous, glaucous, sometimes pale
green; petioles 5-18 mm long, more or less lanate, yellow, sometimes pur-
plish; stipules 6-17 (25) mm long, linear to ovate, base broad and sometimes
irregularly lobed, margins glandular serrulate or prominently and irregularly
toothed, apex inequilaterally attenuate, usually persistent for several years.
Aments precocious, sessile on branches of previous year. Staminate aments
2.5-4.5 cm long; stamens 2, filaments 8-8.5 mm long, glabrous; anthers
0.6-0.7 mm long; nectaries 1, adaxial, 1-1.2 mm long. Pistillate aments
4.5-9.5 cm long, rachis densely white lanate at base; pistils 2.5-3.6 mm
long, green and glabrous, capsules 4.5-7.2 mm long; styles 1.2-1.6 (3.2) mm
long; stigmas 2-4-lobed, 0.4-1.2 mm long; stipes 0.2-0.5 mm long, pubescent;
nectaries 1, adaxial, 0.4-0.8 mm long, 2-3 times as long as stipe; bracts
narrowly obovate, apex acute or obtuse, 2-2.6 mm long, dark brown or
rarely blackish, pubescent both sides with long, wavy, white or rarely yellow
trichomes 2 times as long as bract.
171
Lanatae Koehne
S
Ly
S ™
\ . Sas
SSS s
S S
Map 46 Salix arbusculoides
Habitat
Salix thickets on alluvial gravels
or sands; thickets including Picea
mariana and Betula glandulosa on
mountain slopes; wet Carex aquatilis
meadows.
Discussion
Range
Arctic, alpine: Throughout Alaska and
Yukon Territory; absent from the Aleu-
tian Islands and the Pacific coastal
region; rare near the Arctic Coast of
Alaska; southward to northern British
Columbia; eastward across the North-
west Territories to Hudson Bay and
Baffin Island (Map 47).
Salix lanata ssp. richardsonii is characterized by persistent stipules which are
broad at the base and have an inequilateral, attenuate apex, precocious
aments, glabrous pistils and nectaries 2-3 times as long as the stipes. This
taxon is similar morphologically to S. barclayi and S. planifolia ssp. pulchra
from which it may be distinguished by the characteristics in Table 18.
The possibility of hybridization with S. barclayi has been discussed under
that species. Hybridization with S. alaxensis var. alaxensis has been suggest-
ed as an explanation for the densely pubescent capsules on a specimen from
Cape Thompson (Belson 87).
172
Lanatae Koehne
Map 47 Salix lanata ssp. richardsonii. Circles based on Skvortsov, 1966
Salix lanata ssp. richardsonii is the western North American-eastern Asian
race of the Eurasian S. /anata. It is distinguished from ssp. /anata by its
glabrous to glabrescent, smaller leaves and generally by aments with a
whitish rather than yellowish pubescence. There is some variation in these
characteristics even within North American material, and plants with distinctly
pubescent mature leaves have been seen from Meade River (Argus &
Chunys 5265), Kotzebue (Argus & Chunys 5972) and the lower Yukon River
(Rouse 37). Specimens with yellowish bract pubescence have been seen from
the Sheenjek River (Kessel S4), Kiana (Clark, 4 June 1937), Kennecott
(Argus B3), Eureka Roadhouse (Anderson 8448) and Mile 132, Canol Road,
Yukon Territory (Porsild & Beitung 9460). Skvortsov (1966) regards these
taxa as races because they intergrade in the region between the Lena and
Enisey Rivers. They are evidently closely related, and it does seem to be
appropriate to treat them as races of a single species. The eastern Canadian
Arctic S. calcicola had also been reduced to a subspecies of S. Janata (Hultén
1967), thus organizing this circumpolar complex into three races. Hultén
(1967), in making the latter combination, simply states by way of an explana-
tion, “S. Janata, S. richardsonii and S. calcicola together form a circumpolar
area of closely related taxa best regarded as races of one species.” | am in
agreement with these decisions, for they coincide with my impressions of
these taxa.
173
Plate XXXVI
Salix barrattiana Hook. Argus 6920 (branch) and Argus 644 (ament)
174
Lanatae Koehne
Table 18 Comparison table: Salix /anata ssp. richardsonii, S. barclayi and S. planifolia ssp.
pulchra
Characteristics
stipules
stipule shape
flowering time
floriferous
branchlets
bracts
immature leaf
pubescence
nectary length
stipe length
leaf shape
pistils
bract
pubescence
leaf base
lanata ssp. richardsonii
persistent for
several years
linear to ovate,
broad at base,
apex inequilateral
precocious
none
dark brown to
blackish
white or ferruginous
trichomes
2-3 times stipe
0.2-0.5 mm
elliptic, narrowly
ovate or broadly
obovate
glabrous
white or yellow
trichomes
obtuse, acute to
rounded
Description of species
barclayi
non-persistent
ovate to narrowly
elliptic
coetaneous
present
light brown to dark
brown or bicolour
only white
trichomes
0.5 times stipe
0.4-1.4 mm
elliptic, obovate or
rarely narrowly
elliptic
glabrous
white trichomes
rounded, rarely sub-
cordate or acute
37. SALIX BARRATTIANA Hook.
albertana Rowlee, Bull. Torr. Bot. Club 34: 157. 1907.
planifolia ssp. pulchra
often persistent for
2-4 years
linear, apex
equilateral
precocious
none
dark brown to
blackish
white or ferrugi-
nous trichomes
2-3 times stipe
0.2-0.8 mm
rhombic to narrowly
elliptic, elliptic
or obovate
densely
white sericeous
white trichomes
acute to cuneate
barrattiana Hook. Fl. Bor.-Am. 2: 146. 1838. (Type: Drummond 650, NY).
barrattiana « (var.) /atifolia Anderss. in DC. Prod. 16 (2): 274. 1868.
barrattiana B (var.) angustifolia Anderss. in DC. Prod. 16 (2): 274. 1868.
barrattiana var. marcescens Raup, Sargentia 6: 157. 1947.
Low, often depressed, alpine shrubs, 0.3-1 m tall, forming mats up to 3-4 m
in diameter; branches gnarled, reddish brown, glossy and pubescent; branch-
lets with short internodes, coarsely villous, buds oily. Leaves elliptic to obo-
vate or narrowly obovate, the largest mature leaves 3.7-7.5-9.5 cm long,
1-1.6-2.9 cm wide, 2.2-4.2 (5) times as long as wide; apex acute; base
acute or obtuse; margins entire or glandular to very finely glandular serrulate;
175
Plate XXXVII
Salix candida Fliigge ex Willd. Argus 4868
176
Villosae Rouy
the upper side of mature leaves sparsely pubescent and glossy, the lower side
white or grey sericeous-lanate, the leaves crowded on the short branchlets;
petioles 4-10-15 mm long; stipules broadly ovate, about 5 mm long, margins
glandular and oily. Aments precocious, sessile on branches of previous year.
Staminate aments 3-5 cm long; stamens 2, filaments 5-6 mm long, glabrous;
anthers 0.4-0.6 mm long. Pistillate aments 4.5-11 cm long; pistils 2.5-3
mm long, densely grey white sericeous, capsules 4.5-6 mm long, sparsely
sericeous; styles 0.6-1.6 mm long; stigmas 0.2-0.3 mm long, 4-lobed; stipes
0.2-0.6 mm long, sparsely pubescent to glabrous; nectaries 1, adaxial, 0.5-1.2
mm long, longer than stipe; bracts narrowly oblong, apex acute to rounded,
2.8-3 mm long, black to dark brown, pubescent with long straight trichomes
which exceed bract by about 3 mm.
Habitat Range
River bottoms; gravel stream chan- Alpine, Arctic: Eastern Brooks Range;
nels; hillsides; meadows; wet tundra Alaska Range; mountains of south-
in the mountains. ern Yukon and adjacent Northwest
Territories; southward to southwest-
ern Alberta, British Columbia, and
northwestern Montana (Map 48).
Discussion
Salix barrattiana is a distinctive alpine species characterized by forming low
thickets of grey, erect stems, leaves grey, sericeous-lanate which are crowded
on branchlets with short internodes and with conspicuously oily buds and
stipules. The latter characteristic, which was first described by Porsild (1951),
is very prominent although the nature of the oily substance has yet to be
determined.
This species is apparently related to S. Janata ssp. richardsonii from which
it is readily distinguished by its oily buds and stipules, densely grey white
sericeous pistils and more pubescent leaves.
Section 15. Villosae Rouy
38. SALIX CANDIDA Fliigge ex Willd.
S. candida Fliigge ex Willd. Sp. Pl. 4: 708. 1806.
S. candidula Nieuwl. Amer. Midl. Nat. 3: 225. 1914.
S. candida var. denudata Anderss. in DC. Prodr. 16 (2): 278. 1868. S. can-
dida f. denudata Rouleau, Natur. can. 71: 266. 1944.
Description of species
Shrubs 0.3-3 m tall; branches dark brown, glabrescent; branchlets densely
lanate to floccose, rarely sparsely pubescent. Leaves narrowly elliptic to nar-
rowly ovate, the largest mature leaves 5-8 cm long, 0.8-1.8 cm wide and
177
Villosae Rouy
155° 130°
‘Lily
65° f
Map 48 Salix barrattiana
3.3-6.8 times as long as broad; apex acute; base acute; margins subentire,
often distantly glandular, revolute; the upper side of mature leaves floccose
to glabrescent, drying dark green or brown, the veins impressed, the lower
side densely dull white lanate and floccose, the midrib prominent and yellow-
ish; petioles 3-9 mm long, light brown and pubescent; stipules narrowly
ovate, 2-3 mm long, lanate. Aments coetaneous, on short, leafy, floriferous
branchlets. Staminate aments 1-1.5 cm long, subsessile, floriferous branch-
lets 3-4 mm long; stamens 2, filaments 4-5 mm long, glabrous, distinct or
slightly connate at base; anthers 0.5-0.6 mm long. Pistillate aments 2.2-3
(5) cm long, floriferous branchlets 2-7 mm long; pistils 4-6 mm long, densely
dull white lanate, capsules glabrescent or sparsely lanate; styles 0.3-1 mm
long, red in life; stigmas about 0.2-0.3 mm long; stipes 0.1-1.2 mm long,
lanate; nectaries 1, adaxial, red in life, 0.4-1 mm long, 0.5-0.75-1 times as
long as stipe; bracts narrowly oblong, apex rounded, 1.2-1.5 mm long, pale
to dark brown, pubescent on both sides.
Habitat Range
Occasional in wet, usually alkaline, Boreal: East central Alaska; Fort
habitats including: Carex-Betula glan- Yukon; Tanana River southeast of
178
Villosae Rouy
160°
Map 49 Salix candida
dulosa fens; Larix laricina fens; Bet- Fairbanks, Alaska; Mayo, Whitehorse,
ula glandulosa-Salix thickets at edges Watson Lake, Yukon Territory; south-
of ponds and on river terraces. ward and eastward through the boreal
forest and northern prairies of Canada
and northern United States (Map 49).
Discussion
Salix candida is characterized by narrowly elliptic leaves, densely dull lanate
beneath and dark green or brownish above with dull white floccose pubes-
cence, and densely lanate branchlets and pistils.
This species reaches the northwestern end of its range in central Alaska
where its occurrence is sporadic.
It is related to S. alaxensis, which it sometimes resembles morphologically
(see that species), and to the eastern Siberian and central Asian species S.
krylovii Wolf with which it may be conspecific.
179
Plate XXXVIII
a B
Salix alaxensis (Anderss.) Cov. (A) Variety alaxensis, Argus 5580, (B) Variety /ongistylis
(Rydb.) Schneid., Argus 6495
180
Villosae Rouy
39a. SALIX ALAXENSIS (Anderss.) Cov. var. ALAXENSIS
S. alaxensis (Anderss.) Cov. Proc. Wash. Acad. Sci. 2: 280. 1900.
S. speciosa Hook. and Arn. in Hook. Fl. Bor.-Am. 2: 145. 1838, non Host. 1828.
(Type: Richardson 80 & 81, GH). S. speciosa 8 (var.) alaxensis Anderss. in
DC. Prodr. 16 (2): 275. 1868.
S. alaxensis var. obovalifolia Ball, J. Wash. Acad. Sci. 28: 443. 1938. (Type:
Dutilly 445, US).
Description of species
Shrubs 0.3-4 m tall; branches dark brown to chestnut brown with more or
less persistent grey villous-lanate indumentum; branchlets densely white or
yellowish villous-lanate with long straight and short curly trichomes; buds
villous. Leaves narrowly ovate to oblong or obovate to narrowly obovate, the
largest mature leaves 5-10.8 cm long, 1.5-3.5 cm wide and 2.1-3.8 times as
long as wide; apex acute; base narrowly cuneate; margins revolute, entire,
sometimes glandular and undulating; the upper side of mature leaves bright
green sometimes drying brownish, floccose to glabrescent, the lower side
densely white lanate; petioles (3) 5-15 (20) mm long, yellowish, tomentose
or glabrescent, in about 33 per cent the petiole base is inflated around the
bud; stipules 4-15 (20) mm long, linear, pubescent as the leaves, glandular.
Aments precocious or subprecocious, sessile on branches of previous year.
Staminate aments 3-3.5 cm long; stamens 2, filaments about 7 mm long,
glabrous, distinct or connate; anthers 0.6-0.9 mm long. Pistillate aments 6-15
cm long; pistils about 1.5 mm long, green, sparsely pubescent, capsules
about 4-5 mm long, tawny and glabrescent; styles 1.3-2.8 mm long; stigmas
0.5-1.8 mm long; stipes 0-0.3-0.4 mm long, lanate to sparsely pubescent;
nectaries 1, adaxial, 0.6-1 mm long, 1-1.33-2 times as long as stipe; bracts
ovate, apex acute or obtuse, 1.5-2.5 mm long, dark brown to black, villous
on both sides with long straight trichomes about 2 times as long as the bract.
Habitat Range
Gravel bars and terraces of rivers, Arctic, alpine and boreal: Throughout
streams, and lakes; alpine meadows. Alaska; absent in most of the Aleutian
Islands, some Bering Sea islands, and
in southeastern Alaska south of Gla-
cier Bay; throughout Yukon Territory;
British Columbia; eastward across the
Canadian Arctic; south in the Rocky
Mountains to Jasper National Park;
Asia: Yenisei River; eastward to the
Chukotsk Peninsula; southward to
Lake Baikal (Map 50).
181
Villosae Rouy
135° 130°
Map 50 Salix alaxensis var. alaxensis. Circles based on Skvortsov, 1966, as S. alaxensis
39b. SALIX ALAXENSIS var. LONGISTYLIS (Rydb.) Schneid.
S. alaxensis var. longistylis (Rydb.) Schneid. J. Arnold Arb. 1: 225. 1919.
S. longistylis Rydb. Bull. N.Y. Bot. Gard. 2: 163. 1901. S. alaxensis f. longi-
stylis Boivin, Natur. can. 93: 436. 1966. S. alaxensis ssp. longistylis Hult.
Ark. Bot. Al.-7: 37.4 4967.
Description of variety
Differs from var. alaxensis in tall shrubs or trees 2-10 m tall, basal diameter
about 3 dm; branchlets and branches glaucous, sometimes sparsely pubescent
but soon glabrate; buds often small and the petiole base non-inflated; bract
apex sometimes obtuse.
Habitat Range
Pioneer vegetation on river alluvi- Boreal, Pacific coastal, alpine and
um and glacial moraines; immature Arctic: Central Alaska; southern
forests; subalpine thickets; alpine Brooks Range; eastern Alaska Penin-
182
Villosae Rouy
160°
Map 51 Salix alaxensis var. longistylis
tundra; uncommon in arctic Salix sula; Kenai Peninsula; Kodiak Island;
thickets. mainland southeastern Alaska; Yu-
kon Territory; eastward to Hudson
Bay; south in Rocky Mountains to
northern British Columbia; Asian dis-
tribution uncertain (Map 51).
Discussion
Salix alaxensis is characterized by narrowly ovate to obovate leaves, bright
green above and densely white lanate beneath; the branchlets are densely
villous-lanate (var. alaxensis) or sparsely pubescent and glaucous (var.
longistylis). The flowering is precocious and the aments are large and bear
sparsely pubescent pistils with styles up to 2.8 mm long. This species is re-
lated to S. candida from which it is distinguished by its larger stipules (4-20
mm long), long styles (1.3-2.8 mm long) and very short stipes (0-0.4 mm
long).
The two varieties of S. alaxensis in Alaska and the Yukon, var. alaxensis
and var. /ongistylis, have been variously treated taxonomically. In 1951
Porsild argued for specific status for both taxa, stating that they were distinct
ecologically as well as morphologically; Hultén (1967) treated them as alti-
183
Plate XXXIX
Salix drummondiana Barr. Argus 6924
184
Villosae Rouy
tudinal subspecies; Raup (1959) and others preferred to maintain them as
varieties; Boivin (1966) reduced var. /ongistylis to a forma; and in 1966
Skvortsov treated them as one species and stated that the characteristics
which distinguish var. /Jongistylis occur at random in Asian populations.
Salix alaxensis var. longistylis may be distinguished from var. alaxensis by
a series of characteristics listed above which, although useful in distinguishing
the varieties, are highly variable in themselves and may be variously re-
combined with var. alaxensis characteristics. Branchlet pubescence and glau-
cescence are most important diagnostically, but at the same time are highly
variable. Within a population at College, Alaska, all degrees of pubescence
from densely villous to completely glabrous were observed. Similarly, varia-
tion in branchlet pubescence was noted in specimens from Old John Lake,
Sheenjek River and Mount McKinley National Park. However, it is not simply
branchlet pubescence that is characteristic of the taxa, but also the conspicu-
ous glaucescence on the branchlets of var. /ongistylis. It has been assumed
that this glaucescence does not occur in var. alaxensis, but this is difficult to
affirm because of the densely villous pubescence covering the branchlets.
Specimens of var. alaxensis from Ogotoruk Creek, Cape Beaufort, Seward
Peninsula, Old John Lake and the Little Delta River that have had the pubes-
cence partly worn off the stems reveal a glaucescence just as conspicuous as
in var. longistylis. If both taxa have glaucous branchlets, then the difference
is simply a matter of degree of pubescence, which has already been shown to
be highly variable.
Enlarged winter buds and inflated petiole bases are also characteristic of
var. alaxensis. These characteristics were the primary bases for the recogni-
tion of var. obovalifolia, which is treated here as a synonym of var. a/axensis.
Although inflated petiole bases and large buds are common in var. alaxensis,
and infrequently encountered in var. /ongistylis, they do occur in the latter
variety. | have collected specimens of var. /ongistylis with inflated petiole
bases at College, Alaska, and Peterson Creek, British Columbia, and | have
seen other specimens with this characteristic from the Matanuska Valley.
A series of specimens of S. alaxensis collected by C. Rouse at Egavik on
Norton Sound (ALA) show gradations from densely yellowish villous branch-
lets, large winter buds and inflated petioles (no. 76) to sparsely pubescent,
glaucous branchlets, large buds and inflated petioles (no. 78), to sparsely
pubescent, glaucous branchlets, small buds and uninflated petioles (no. 79).
Specimens such as nos. 76 and 78 have been determined as S. alaxensis var.
alaxensis > longistylis to indicate their intermediate character (Map 3).
One of the major arguments for recognizing these taxa is their partial
geographical and ecological separation. Var. alaxensis occurs in the Arctic
and at somewhat higher elevations in the mountains than does var. /ongi-
stylis; the latter taxon is more frequent throughout the boreal forest and does
not extend into the Arctic. While | am unable to accept Skvortsov’s view
(1966) that the taxa are completely confluent, | also cannot accept them as
distinct species. Since they are more or less confluent in some areas, and
since the characteristics that distinguish them are variable and often re-
combined in intermediate specimens, they are best treated as varieties.
185
Villosae Rouy
40. SALIX DRUMMONDIANA Barr.
S. drummondiana Barr. in Hook. Fl. Bor.-Am. 2: 144. 1838. (Type: Drummond
672, photo and frag. A).
S. bella Piper, Bull. Torr. Bot. Club 27: 399. 1900. S. drummondiana var. bella
Ball, Amer. Midl. Nat. 45: 743. 1951.
S. subcoerulea Piper, Bull. Torr. Bot. Club 27: 400. 1900. (Type: Cusick 1302,
US). S. drummondiana var. subcoerulea Ball, Amer. Midl. Nat. 45: 744. 1951.
S. pachnophora Rydb. Bull. Torr. Bot. Club 31: 403. 1904.
S. pellita auct. non Anderss.
Description of species
Shrubs 1-4 m tall; branches dark reddish brown, glabrous or glabrescent,
glaucous, glossy and brittle; branchlets brownish green to reddish brown,
glabrous or rarely sparsely pubescent, glaucous. Leaves elliptic, elliptic-obo-
vate to narrowly elliptic-obovate, the largest mature leaves 4.2-8.5 cm long,
1.1-2.6 cm wide and 3.2-6.2 times as long as wide; apex acute; base acute;
margins entire, distantly glandular dotted or glandular crenate, revolute;
immature leaves densely white sericeous; the upper side of mature leaves
dark green, sparsely pubescent with white or, rarely, ferruginous trichomes,
semi-glossy, the lower side densely to sparsely sericeous with white or white
and ferruginous trichomes, pale non-glaucous; midrib yellow and prominent;
petioles 2-12 mm long, yellow to reddish; stipules narrowly elliptic (0.2)
1.2-3.2 (7.2) mm long, pubescent abaxially or entirely glabrous, margins
glandular, deciduous. Aments precocious, sessile on branches and bearing
from one to several dark brown bracts at base. Staminate aments 2-2.7 cm
long; stamens 2, filaments about 6 mm long, glabrous, distinct or connate
at base; anthers about 0.4-0.6 mm long; nectaries 1, adaxial, 0.4-0.6 mm
long. Pistillate aments 2.5-8 cm long, rachis pubescent; pistils about 2.4 mm
long, densely sericeous, capsules 2-5.6 mm long, greenish tawny and seri-
ceous; styles (0.4) 0.7-1.3 mm long; stigmas 2, horseshoe-shaped or 4 linear
lobes, 0.3-0.4 mm long; stipes 0.6-1.4 mm long, pubescent; nectaries 2,
adaxial and abaxial, 0.6-0.8 mm long, 0.5-0.7 (1-2) times as long as stipe;
bracts ovate, apex acute, 1.2-2 mm long, brown to dark brown, pubescent
on both sides with long straight trichomes 2 times as long as bract.
Habitat Range
Salix thickets along streams; river Montane, boreal: Southeastern Yu-
margins; subalpine Salix thickets. kon; southward in the cordillera to
California and Nevada; eastward to
north central Alberta and Montana
(Map 52).
186
Villosae Rouy
120°
Mo Gilles
he
Map 52 Salix drummondiana
Discussion
Salix drummondiana is a minor component of the willow flora of Alaska and
the Yukon, entering that area only in southeastern Yukon. However, it is
important in that it may be related to S. alaxensis — it shows some signs of
intergradation with S. alaxensis var. longistylis in the Watson Lake area. The
two taxa may be distinguished as in Table 19.
Salix drummondiana is a member of a North American complex including
the eastern S. pellita Anderss. and the northwestern S. alaxensis. Salix pellita
is sometimes treated as synonymous with S. drummondiana, but because of
its distinctive leaf morphology (resembling S. viminalis L.), | am inclined to
recognize it as a distinct species although the two do appear to intergrade
in central Canada. Cronquist (1964) indicated that S. pe/lita was apomictic,
apparently because of the absence of staminate flowering specimens. | have
seen staminate flowering specimens from Saskatchewan and | am confident
that they occur elsewhere, but because of the precocious flowering of this
species, they are either not frequently collected or not recognized when they
are. This complex requires further study before any taxonomic revision can
be made.
187
Villosae Rouy
Table 19 Comparison table: Salix drummondiana and S. alaxensis var. longistylis
Characteristics drummondiana alaxensis var. longistylis
leaves sericeous beneath villous lanate
although sometimes beneath
+ lanate
style length (0.4) 0.7-1.3 mm 1.3-2.8 mm
stigma length 0.3-0.4 mm 0.5-1.8 mm
stipe length 0.6-1.4 mm 0-0.3-0.4 mm
pistillate nectaries 2 1
The identity of the type material of S. drummondiana has been a persistent
source of confusion (Raup 1934, 1959; Ball 1951) which can now be resolved.
Of the original specimens studied by Barratt, | have seen a photograph
and fragments from the sheet at Kew (Drummond 672, Rocky Mountains, A)
and two sheets at the New York Botanical Garden (one sheet Drummond,
H. B. and T. No. 2 and a second sheet simply labelled “Salix drummondiana,
Fl. Bor. Am."’). The material included on these sheets is heterogeneous, con-
taining flowering branches, an immature fruiting branch, and a mature
vegetative branch. Unfortunately the type at the Barratt Herbarium at Wes-
lyan University, Middletown, Conn., which was cited by Raup (1934) as an-
notated by Barratt himself, could not be relocated. But all the material does
have a consistency about it that suggests that all the specimens were ob-
tained from the same three plants.
The assumption that the three types of branches belong to the same
species was tacitly made by Barratt (Hooker 1838), and accepted by Schnei-
der (1919c) and Raup (1934, 1959). However, this assumption is false;
Barratt’s description is a composite one. The description of leaves is based
on the mature vegetative branch, and that of flowers and aments is based on
the flowering branches and the immature fruiting branch. Raup (1934, 1959)
was seriously misled by this assumption of taxonomic homogeneity of the
material when he concluded that the leaves belonged to S. alaxensis var.
longistylis and that the small aments and pistils were ‘in a state of arrested
or partial development’, and indicated hybridity. If we study the three ele-
ments of the original material without assuming that they are the same taxon,
it is evident that the flowering branches represent S. arbusculoides, that the
immature fruiting branch (on the Kew sheet only) represents S. drum-
mondiana, and that the mature branch represents either S. alaxensis var.
longistylis or S. drummondiana.
There are seven flowering branches on the three sheets that | have seen.
The branches are dark reddish brown, glossy and have a trace of glauces-
cence. The unfolding leaves are narrowly elliptic, glabrous above and seri-
ceous beneath, the margins are glandular serrulate with about 6 gland-tipped
serrulations per mm, and the stipules are narrowly elliptic and glandular. The
coetaneous aments are 1.2-1.7 cm long, and subsessile with 2-3 green bracts;
the pistils are 1.5 mm long, on stipes 0.4 mm long, and both are sericeous
with shiny, straight trichomes; the styles and stigmas are ca. 0.2 mm long.
This material can be positively identified as S. arbusculoides. The glandular
188
Villosae Rouy
serrulations on the leaf margins are very diagnostic, even in immature mate-
rial, and although leaves of S. alaxensis and S. drummondiana are sometimes
toothed, the serrulations are never as uniform or as numerous as in S. arbus-
culoides. The subsessile aments, the short styles and stigmas and the short,
but evident, stipes as well as the coetaneous flowering, are characteristic of
S. arbusculoides and serve to distinguish it from S. alaxensis and S. drum-
mondiana.
There is one mature vegetative branch on the Kew specimen and a frag-
ment of a branch bearing 2 leaves on one of the specimens at NY. The
branch is dark brown, glaucous, bearing narrowly obovate leaves 6.7 x 2.8
cm and 6.5 x 2.6 cm, with obtuse apices, acute bases and petioles 7-9 mm
long, glabrous above and densely lanate beneath, the midrib yellowish and
glabrous. The identity of this material is not unequivocal, and it could repre-
sent either S. alaxensis var. longistylis or S. drummondiana. As Raup (1934)
has noted, these taxa are difficult to distinguish vegetatively, and although S.
alaxensis var. longistylis has leaves densely lanate beneath (curly trichomes)
and S. drummondiana usually has leaves sericeous beneath (straight, silky
trichomes), there are many exceptions. For example, specimens with inter-
mediate pubescence are known from Watson Lake, Y.T. (Argus 5017), and
the taxon called S. bella Piper (= S. drummondiana) is characterized by
lanate leaves. It must be concluded that positive identification of this mate-
rial cannot be made with our present knowledge.
There is one immature fruiting branch, the centre branch on the Kew
specimen. | have been able to examine only a young shoot and an ament of
this specimen at A. The leaves are variously elliptic, densely white sericeous
beneath, sparsely pubescent above with margins entire and glandular dotted.
The aments are sessile, ca. 2.5 cm long; the pistils are densely sericeous,
3.2-4 mm long, with styles ca. 0.6 mm long; the stipes are ca. 1.2 mm long, and
the nectaries are about one third the length of the stipe. This branch compares
favourably with what is usually named S. drummondiana (S. subcoerulea) in
the Rocky Mountains.
Since the original material consists of three discordant elements, the name
must be rejected, ‘‘unless it is possible to select one of these elements as a
satisfactory type” (Article 70, Lanjouw, et al., 1966). The proto/ogue is based
on all of the elements and does not provide a clear basis for the selection of
a lectotype. However, in order to preserve current usage (Recommendation
7B, Lanjouw, et al., 1966), the centre branch on the Kew specimen is
selected as the lectotype. The flowering branches are redetermined as S.
arbusculoides and the mature vegetative branch as S. a/axensis var. longistylis
or S. drummondiana (if the latter, then a syntype).
189
Plate XL
Salix sitchensis Sanson. Vegetative specimen, Argus 6672 (left) and pistillate specimen,
Argus 6651 (right)
190
Sitchenses Bebb
Section 16. Sitchenses Bebb
41. SALIX SITCHENSIS Sanson
S. sitchensis Sanson, in Bong. Mem. Acad. St. Petersb. 6. 2: 162. 1832.
(Type: Lyall s.n. photo and frag. A).
S. coulteri Anderss. Ofvers. Vet. Akad. Forh. (Stockh.) 15: 119. 1858. S.
sitchensis f. coulteri Jeps. Fl. Calif. 1: 342. 1909. S. sitchensis var. coulteri
Jeps. Man. Fl. Pl. Calif. 265. 1923.
S. sitchensis var. congesta Anderss. in DC. Prodr. 16 (2): 233. 1868.
S. sitchensis var. denudata Anderss. in DC. Prodr. 16 (2): 233. 1868.
Description of species
Shrubs 0.5-3.8 m tall, sometimes prostrate or decumbent; branches dark
brown becoming greyish, indumentum usually persistent for 2 years; branch-
lets densely sericeous or velutinous becoming sparsely pubescent, brittle.
Leaves narrowly elliptic, narrowly obovate or obovate, the largest leaves
(321 45-35 .(12)..cmlongs.(1.3),» 1.9-3.2 (4.8) cm wide and (2.1) 24-3
(3.5) times as long as wide; apex round with an obtuse or sometimes acu-
minate tip, sometimes acute, often conduplicate when pressed; base cuneate;
margins distantly and inconspicuously glandular serrulate or irregularly
glandular crenate, crispate in some vigorous leaves, revolute; the upper side
of mature leaves sparsely sericeous or villous becoming glabrate, bright
green and dull, the lower side densely to sparsely sericeous with matted
straight trichomes appearing satiny, epidermis glossy, more vigorous leaves
may be sericeous-lanate; petioles (3) 4-10 (16) mm long, yellowish, veluti-
nous becoming sparsely so; stipules half ovate, minute to 0.4-1.5 mm long,
larger on vigorous shoots, pubescent as the leaves. Aments coetaneous,
rarely subprecocious, on leafy, floriferous branchlets. Staminate aments
2.2-3 cm long, floriferous branchlets 0.5-1 cm long, persistent after ament
drops; stamens 1, filaments 4.4-5.2 mm long, glabrous; anthers 0.6-0.7 mm
long. Pistillate aments (1.5) 3-6.5 (9.5) cm long, floriferous branchlets 1-2
cm long; pistils tear-shaped, 2.4-3.6 mm long, densely white sericeous, cap-
sules 3.6-5.6 mm long, tawny green to yellowish, sericeous or sparsely so;
styles 0.5-0.8 mm long; stigmas 0.1-0.2 mm long; stipes 0.8-1.4 mm long,
densely sericeous; nectaries 1, adaxial, 0.5-0.8 mm long, equal to or 0.5
times as long as stipe; bracts narrowly oblong to obovate, apex rounded,
1.5-2.4 mm long, brown, usually bicolour and darker toward apex, villous
on both sides.
Habitat Range
Gravel bars of rivers; glacial mo- Pacific coastal, montane: Coastal
raines; Salix and Alnus thickets; southern Alaska, from Kodiak Island
shrubby openings in forests. to southeastern Alaska and adjacent
191
Sitchenses Bebb
160° 135°
AK
Map 53 Salix sitchensis
British Columbia; southward to Wash-
ington and California (Map 53).
Discussion
Salix sitchensis is characterized by usually obovate leaves, satiny pubescent
beneath, with revolute vernation; slender pistillate aments with densely seri-
ceous pistils and staminate flowers with one stamen each. It may be confused
with vegetative specimens of the densely pubescent form of S. scouleriana
(see that species for discussion).
In southeastern Alaska Salix sitchensis is a pioneer species on glacial
moraine and gravel bars. In these habitats it may be a prostrate shrub
spreading as much as 7 m in diameter. It remains a component of later
successional stages, finally becoming the dominant species in mature Salix-
Alnus thickets. It then declines in importance as the forest becomes closed,
but may remain on gravelly stream margins and in disturbed openings within
the forest. It is one of the most conspicuous willows in coastal Alaska, but is
restricted to coastal habitats and is unable to cross divides such as the Three
Guardsman Pass, British Columbia (3,500 ft), north of Haines, Alaska. | know
of no case of its occurrence north or east of the coastal mountains.
192
Appendix A
Key to Codes
PROVINCE / STATE
1 ALBERTAs* CANADA
2 BRITISH COLUMBIAe CANADA
3 MANITOBA? CANANA
7 NORTHWEST TERR.* CANADA
11 SASKATCHEWAN? CANADA
12 YUKON TERRITORYe CANADA
14 ALASKAr UeSeAe
QUADRANGLES
ALBERTAse CANADA
1 2 BISTCHO LAKE QUAD
1 3 WHITESAND RIVER QUAD
1 & PEACE POINT QUAD
1 5S FITZGERALD QUAN
1 9 LAKE CLAIRE QUAD
1 10 FORT CHIPEwWYAN QUAD
1 20 WATERWAYS QUAD
1 22 WINAGAMI QUAD
1 24 PELICAN QUAD
1 29) TAWATINAW QUAD
1 32 EDSON QUAD
1 34 EDMONTON QUAD
1 36 CANOE RIVER QUAD
1 38 ROCKY wOUNTAIN HOUSE QUAD
1 41 GOLDEN QUAD
1 44 OYEN QUAD
1 4& FERNIE QUAD
BRITISH COLUMBIAr CaNADA
2 1 YAKUTAT QUAD
2 2 TATSHENSHINI RIVER QUAD
2 4% ATLIN QUAD
2 5S JENNINGS RIVER QUAD
2 0 MCDAME QUAD
2 7 RABBIT RIVER QIIAD
2? 12 TULSEQUAH QUAD
2? 13 DEASE LAKE QUAN
2 14 CRY LAKE QUAD
2 16 TUCHODT LAKES QUAD
2? 17 FORT NFLSOW QUAD
2 20 TELEGRAPH CREEK QUAD
2 21 SPATSIZI QUAD
2 22 TOODOGGONE RIVFR QUAD
2 24% TRUTCH QUAD
2 20 ISKUT RIVER QUAD
2 3U HALFWAY RIVER QUAD
2 31 CHARLIE LAKE QUAD
2 33 HAZELTON QUAD
2 35 PINE PASS QUAD
? 39 TERRACE QUAU
2 40 SMITHERS QUAD
2 41 FORT FRASER QUAD
? 44 GRAHAM ISLAND Quad
2 45 HECATE STRAIT Quad
2 46 DOUGLAS CHANNE! QUAD
2 49 PRINCE GEORGE QuAD
2 51 MOUNT ROBESON QuAD
2 52 MORESBY ISLAND QUAD
2 55 ANAHEIM LAKE QUAD
? 56 QUESNEL QUAD
2 62 MOUNT wADDINGTON QUAD
? 64 BONAPARTE LAKE QUAD
2 65 SEYMOUR ARM QUAD
2 72 ASHCROFT QUAD
?-74 LARDEAU QUAD
2 79 HOPE QUAD
2 8U PENTICTON QUAD
MANITOKA+e CANADA
3 52 BRANDON QUAD
SASKATCHEWAN? CANADA
TAZIN LAKE QUAN
PHELPS LAKE QUAD
WILLIAm RIVER QUAD
WATERHFN RIVER QUAD
SHELLBROOK QUAN
PRINCE ALBERT QUAD
MELFORT QUAD
HUDSON BAY QUAN
NORTHWEST TERRee CANADA
Te Wy
7133
7134
7135
7140
7152
7153
7170
7173
7161
7188
7208
7214
7218
7241
7243
7245
7254
7274
7275
7276
72835
7284
7285
7290
7297
7306
7307
7308
7309
7313
7315
7316
7329
7335
7338
7339
734uU
7341
7342
7345
7346
7347
7345
7349
7350
75607
7368
7369
7370
7371
7372
Tote
7374
7375
7376
7388
7391
7392
7394
7395
7396
7397
7402
7421
CAPE DALHOUSIE QUAD
MACKENZIE pELTa QUAD
STANTON QUAD
FRANKLIN BaY QUAD
CAMBRINGE KAY QUAD
AKLAVIk QUaD
CROSSLFY LAKES QUAD
BELL RTVER QUAN
TRAVATIILANT LAKE QUAD
COPPERMINE QUAN
OGDEN BAY yWUAD
FORT GOOD HOPE QUAD
SOLAN RIVER QUAD
MARA RIVER QUAN
NORMAN WELLS QUAD
FORT FRANKLIN QUAD
LEITH PENINSULA QUAD
PELLY LAKE QUAN
MOUNT EDUNI QUAD
CARCAJOU CANYON QUAD
FORT NORMAW QUAD
LAC DE GRAS QUAD
AYLMER LAKE QUAD
HEALEY LAKE QuaD
SCHULTZ LAKE QUAD
CORAL HARBUUR QUAD
NIDDERY LAKE QUAD
SEKwI MOUNTAIN QUAD
WRIGLEY LAKE QUAD
DAHADINNI RIVER QUAD
MARIAN RIVER QUAD
CARP LaKE wJAD
MACKAY LAKE QUAD
BEAR COVE QUAD
FROBISHER wUAD
NAHANNT QUAD
GLACIER LAKE QiAD
ROOT RIVER QUAN
CAMSELL BEWD QIJAD
BULMER LAKE QUAD
YELLOwKNIFE QUAD
HEARNE LAKE QUAD
SNOWDRIFT QUAD
FORT RELIANCE QUAD
LYNX LAKE QUAD
BEAVERHILL LAKF QUAD
FLAT RIVER QUAD
VIRGINIA FALLS QUAD
SIBBESTON LAKE QUAD
FORT SIMPSON QUAD
MILLS 1 AKE QUAN
FALAISF LAKE @iAD
SULPHUR BAY QUAD
FORT RESOLUTION QUAD
TALTSON LAKE wiiAD
NONACHO LAKE QUAD
FORT LIARD QUAN
TATHLINA LAKE QUAD
BUFFALO LAKE QUAD
FORT SMITH QUAN
HILL ISLAND LAKE QUAD
ABITAU LAKE QUAD
WHOLDATA LAKE RUAD
HYDE LaKE QUAD
FRANCES LAKE QIJAD-
YUKON TFRRITORY* CANADA
12
1?
12
12
12
12
12
12
12
12 10
12 11
if ie
12 13
12 14
12 15
12 16
1? 17
crnouvrunre
DEMARCATION POINT QUAD
HERSCHEL ISLAND QUAD
DAVIDSON MTSe QUAD
BLOw RIVER QUAN
OLD CROW QUAD
BELL RIVER QUAN
PORCUPINE RIVER QUAD
EAGLE RIVER QUAD
TRAIL RIVER QUAD
MARTIN HOUSE QUAD
OGILVIF RIvceR QUAD
HART RIVER QUAN
WIND RIVER QUAN
SNAKE RIVER QUAD
DAWSON QUAD
LARSEN CREEK QUAD
NASH CREEK QUAND
193
Appendix A
ALASKA?
14001
14002
14003
14004
14005
14006
14007
14008
14009
14010
14011
14012
14013
14014
14015
14016
14017
14018
14019
14020
14021
14022
14023
14024
14025
14026
14027
14028
14029
14030
14031
14032
14033
14034
14035
14036
14037
14038
14039
14040
14041
14042
14043
14044
14045
14046
14047
14048
14049
14050
14051
14052
14053
140546
14055
14056
14057
14058
14059
14060
194
NADALEEN RIVER QUAD
BONNET PLUME LAKE QUAN
STEWART RIVER QUAD
MCQUESTEN QUAD
MAYO QUAD
LANSING QUAD
NIDUERY LAKE QiIAD
SEKwI MOUNTAIN QUAD
SNAG QUAD
CARMACKS QUAD
GLENLYON QUAD
TAY RIVER wUAD
SHELDON LAKE QUAD
NAHANNI QUAD
KLUANE LAKe QUAD
AISHIHTIK LAKE QUAD
LABERGF QUAD
QUIET LAKE QUaAn
FINLAYSON LAKE QUAD
FRANCES LAKE QUAD
FLAT RIVER QUAN
MOUNT ST ELIAS QUAD
DEZADEASH QUAD
WHITEHORSE QUAN
TESLIN QUAL
WOLF LAKE wVJAD
WATSON LAKE QUAD
COAL RIVER QUAN
LA BICHE RIVER QUAD
FORT LIARD QUAN
UeSebe
DIXON FNTRANCE QUAD.
PRINCE RUPERT QUAD.
KETCHIKAN QUAD.
CRAIG QUAD.
PORT ALEXANDER QUAD.
PETERSRURG QUAN.
BRADFIFLD CANAL QUAD.
SUMDUM QUAD.
SITKA QUAD.
MTeFAIRWEATHER QUAD.
JUNEAU QUAD.
TAKU RIVER QUAN.
ATTU QUAD.
KISKA QUAD.
RAT ISt ANDS
GARELO!I ISLAND
ADAK Q.iAD.
ATKA QUAD.
SEGUAM QUAi)«
AMUKTA QUAI)
SAMALGA ISLAND QUAD.
UMNAK QUAD.
UMALASKA QUAD.
UNIMAK QUAD.
FALSE PASS QUAN.
SIMEONOF ISLAND QUAD.
STEPOVAK BAY QIIADe
PORT MOLLER QUAD.
COLD BAY QUAD.
CHIGNIK QUAD.
SUTWIK ISLAND QUAD.
TRINITY ISLANDS QUAD.
KAGUYAK QUAD.
KODIAK QUADe.e
KARLUK QUAI).«
UGASHIk QUAD.
BRISTOL BAY QUAD.
PRIBILOF ISLANNS QUAD.
HAGEMEISTER ISLAND QUAD.
NUSHAGAK BAY QUAD.
NAKNEK QUAie
MTe KATMAI QUAN.
AFOGNAK QUAD.
ATLIN «GUAD.
SKAGWAY QUAD.
YAKUTAT QUAD.
ICY BAY QUAD.
MIDOLETON ISLAND QUAD.
BLYING SOUND QUAD.
SELUOVTA QUAD.
ILIAMNA QUAL.
DILLINGHAM QUAN.
GOODNEwS QUAD.
KUSKOKWIM BAY QUAD.
CAPE MFNDENHALL WGUAD.
STe MATTHEW QUAD.
NUNIVAK ISLAND QUAD.
BAIRD TNLET QUAD.
BETHEL QUAD.
TAYLOR MTS. QUAD.
14061
14062
14063
14064
14065
14066
14067
14068
14069
14070
14071
14072
14073
14074
14075
14076
14077
14078
14079
14080
14081
14082
14083
14084
14085
14086
14087
14088
14089
14090
14091
14092
14093
14094
14095
14096
14097
14098
14099
14100
14101
14102
14103
14104
14105
14106
14107
14108
14109
14110
14111
14112
14113
14114
14115
14116
14117
14116
14119
1412U
14121
14122
14123
14124
14125
14126
14127
14128
14129
1413u
14131
14132
14133
14134
14135
14136
14137
14136
14139
14140
14141
14142
14143
14144
14145
14146
14147
14148
14149
14150
14151
14152
14153
LAKE Ci ARK QUAN.
KENAI QUAD.
SEWARD QUAU.
CORDOVA QUAD.
BFRING GLACIER QUAD.
MTe ST. ELIAS QUAD.
MCCARTHY QUAD.
VALDEZ QUAD.
ANCHORAGE QUAD.
TYONEK QUAi)e
LIME HILLS QUAN.
SLEETMUTE QUAD.
RUSSIAN MISSION QUAD.
MARSHALL QUAD.
HOOPER BAY QUane
BLACK QUAD.
KwWIGUK QUAI)e
HOLY CROSS QUAN.
IDITAROD QUAD.
MCGRATH QUAD.
TALKEETNA QUAD.
TALKEETNA mTSe QUAD.
GULKANA QUaDe
NABESNA QUAD.
TANACROSS QUAD.
MOUNT HAYES QUaDe
HEALY QUAD.
MOUNT MCKINLEY QUAD.
MEDFRA QUAD.
OPHIR QUAD.
UNALAKLEET QUAN.
ST» MICHAEL QUAD.
STe LAwRENCE QUAD.
NOME QUAD.
SOLOMON QUaDe
NORTON BAY QUAN.
NULATO QUA\I)e
RURY QUAD.
KANTISHNA RIVER QUAD.
FAIRBANKS WUAD.
BIG DELTA WUAD.
EAGLE QUAD.
CHARLEY RIVER QUAD.
CIRCLE QUAD.
LIVENGOOD wUAD.
TANANA QUAL.
MELOZITNA wQUAD.
KATEEL RIVER QUAD.
CANDLE QUAL.
RENDELEBEN QUAN.
TELLER QUAD.
SHISHMAREF QUAN.
KOTZEBUIE QUAD.
SELAWIK QUAD.
SHUNGNAK QUAD.
HUGHES QUAD.
BETTLES QUAD.
BEAVER QUAD.
FORT YUKON QUAN.
BLACK RIVER QUAD.
COLEEN QUAD.
CHRISTIAN wUAD.
CHANDAIAR QUAD.
WISEMAN QUAD.
SURVEY PASS QUAD.
AMBLER RIVER QUADe
BAIRD mTSe QUAN.
NOATAK QUAD.
POINT HOPE QUAN.
NE LONG MTSe QUAD.
MUSHEGIIR MTSe QUAD.
HOWARD PASS QUAD.
KILLIK RIVER QUADe
CHANDLFR LAKE QUAD.
PHILIP SMITH MTSe QUAN.
ARCTIC QUAD.
TABLE MOUNTAIN QUADe
DEMARCATION POTNT QUAN.
MT» MICHELSON QUAD.
SAGAVANIRKTOK QUAD.
UMIAT QUAD.
IKPIKPIIK RIVER QUADe
LOOKOUT RIDGE QUAD.
UTUKOK RIVER QUAD.
POINT t.AY WUAD.
WAINWRIGHT QUAN.
MEADE RIVER QUAD.
TESHEKPUK QUAD.
HARRISON BAY QUAD.
BEECHEY POINT QUAD.
FLAXMAN ISLAND QUAD.
BARTER ISLAND QUAD.
BARROW QUAD.
Key to Codes
SPECIES
122
122
39
45S
50
71
=
oOo
i)
wp =
105
105
109
113
116
36
118
LO ad
HYBRIDS
J
ane,
OFND-
ALAXENSIS VAR. ALAXENSIS
ALAXENSIS VAR. LONGISTYLIS
ARBUSCIJLOIDES
ARCTICA
ARCTOPHILA
ATHABASCENSIS
BABYLONICA
BARCLAYI
BARRATTIANA
BEBbDIANA
BRACHYCARPA SSPe BRACHYCARPA
BRACHYCARPA SSPe NIPHOCLADA
CANDIONA
COMMUTATA
CHAMISSONIS
ORUMMONDI ANA
FUSCESCENS
GLAUCA VAR. VILLOSA
GLAUCA VAR. ACIITIFOLIA
GLAUCA VAR. GLAUCA
HASTATA
HOOKERTANA
INTERIOR
LANATA SSP. RICHARDSONII
LASIANNDRA
MACCALLIANA
MONTICOLA
MYRTILLIFOLIA
NOVAE=ANGLIAE
NUMMULARIA
OVALIFOLIA VAR. OVALIFOLIA
OVALIFOLIA VAR. ARCTOLITORALIS
OVALIFOLIA VAR. CYCLOPHYLLA
OVALIFOLIA VAR. GLACIALIS
PENICELLARIS
PENTANDRA
PHLEBOPHYLLA .
PLANIFOLIA SSP. PULCHRA VARe PULCHRA
PLANIFOLIA SSP. PLANIFOLIA
PLANIFOLIA SSP. PULCHRA VARe YUKONENSIS
POLARIS
PYRIFOLIA
RETICULATA SSP. RETICULATA
RETICULATA SSP. GLABELLICARPA
RIG1DA
ROTUNDTFOLIA SSP. ROTUNDIFOLIA
ROTUNDTFOLIA SSP. DODGEANA
SCOULERIANA
SETCHELLIANA
SITCHENSIS
SPHENOPHYLLA
STOLONIFERA
ALAXENSIS VAR. ALAXENSIS X 103 LANATA
ALAXENSIS VAR. LONGISTYLIS X 122 ALAXENSIS
ARCTICA X 87 RARCLAYT
ARCTICA X 49 GLAUCA
ARCTICA X 83 OVALIFOLIA
ARCTICA X 118 STOLONIFERA
ATHABASCENSIS x 15 PEDICELLARIS
BARCLAYI X 103 LANATA
BARCLAYI X 32 COMMUTATA
BARCLAYI X 118 STOLONIFERA
BRACHYCARPA SSPe NIPHOCLADA X 49 GLAUCA
PEDICELLARIS X 71 ATHABASCENSIS
PHLEBOPHYLLA X 105 ROTUNDIFOLIA
PLANIFOLIA SSP. PULCHRA X 109 SCOULERIANA
ROTUNDTFOLIA SSPe ROTUNDIFOLIA X 92 PHLEBOPHYLLA
195
Appendix A
HERBARIA
A ARNOLD ARBORETUM, CAMBRIDGE, MASS.
ALA UNIVERSITY OF ALASKA, COLLEGE, ALASKA
BRY BRIGHAM YOUNG UNIVERSITY, PROVO, UTAH
CAN NATIONAL HERBARIUM OF CANADA, OTTAWA
CS COLORADO STATE UNIVERSITY, FORT COLLINS, COLO.
DAO CANADA DEPARTMENT OF AGRICULTURE, OTTAWA
FSLC U.S. FOREST SERVICE, COLLEGE, ALASKA
GH GRAY HERBARIUM, CAMBRIDGE, MASS.
GWA PERSONAL HERBARIUM, GEORGE W. ARGUS, AT OTF
ISC IOWA STATE UNIVERSITY, AMES, IOWA
LAV PERSONAL HERBARIUM, L. A. VIERECK, AT ALA
MICH UNIVERSITY OF MICHIGAN, ANN ARBOR, MICH.
MIN UNIVERSITY OF MINNESOTA, MINNEAPOLIS, MINN.
NA U.S. NATIONAL ARBORETUM, WASHINGTON, D.C.
NY NEW YORK BOTANICAL GARDEN, NEW YORK
OTF CANADIAN FORESTRY SERVICE, OTTAWA
RM ROCKY MOUNTAIN HERBARIUM, LARAMIE, WYO.
SASK UNIVERSITY OF SASKATCHEWAN, SASKATOON, SASK.
US U.S. NATIONAL MUSEUM, WASHINGTON, D.C.
UWO UNIVERSITY OF WESTERN ONTARIO, LONDON, ONT.
WIS UNIVERSITY OF WISCONSIN, MADISON, WIS.
WTU UNIVERSITY OF WASHINGTON, SEATTLE, WASH.
196
Appendix B
Specimens Cited
TAXON SPC S V HYB PROV QUAD LAT
3 BABYLONICA
1673 3 14 6 5648
4 BEBBIANA
1873 4 1 3 5854
1873 4 1 5 5908
1873 4 1 38 5258
1873 4 2 2 5935
1873 4 2 7 5959
1875 qy 2 14 5822
18735 4 2 16 5839
1873 4 2 16 5839
1873 4 2 16 5847
1673 4 2 24 5714
1873 4 2 24 5705
1873 4 2 24 5705
1873 4 2 24 §705
1873 q 2 24 5705
1873 4 2 24 5659
1873 4 2 040 5445
1873 4 2 049 5320
1673 4 2 049 5320
1873 4 7 #153 6833
1673 4 7 #170 6745
18673 4 7 #241) 6517
1873 4 7 339 6205
1873 4 7 339 6205
1873 4 7 339 6205
1673 4 7 339 6205
1673 4 7 368 6137
1673 4 7 370 6152
1673 4 7 370 6152
1873 4 7 370 «©6152
1673 4 7 394 6000
1673 4 11 331 _ 5207
1873 4 12 15 6404
1873 4 12 15 6412
1875 4 12 26 0214
1673 4 12 27 »6205
1873 4 12 32 6122
1673 4 12 35 6155
1373 4 12 40 6047
18673 y 12 40 6046
1873 4 12 40 6046
13873 Q 12 41 6043
1873 y 12 41 6043
1673 4 12 41 6043
14873 4 12 41 0043
1073 4 12 41 6043
1673 4 12 41 6043
1873 4 12 41 ©6039
1873 4 12 41 6033
1573 4 12 42 6022
1673 4 12 42 ©006
1873 4 12 43 6005
1873 4 12 43 6007
1673 4 12 44 6007
1873 4 12 44 6003
1873 4 12 44 6003
1673 4 12 44 6003
1673 4 12 44 6003
1873 4 12 44 6003
1873 4 12 44 6003
1873 & 12 44 6003
1673 4 12 44 6013
1873 4 14 35 5748
1873 4 14 41 5842
1873 4 14 42 5845
1873 4 14 42 5845
1873 4 14 42 58641
1873 et 14 50 5937
1873 4 14 50 5940
1873 a 14 62 6055
1873 “ 14 62 6033
1873 4 14 62 6009
1873 4 14 63 6929
1873 4 14 63 6029
1875 4 14 63 6029
1873 a 14 63 6029
1873 4 14 67 6126
1873 4 14 67 6130
LONG
13258
11135
11128
11322
13629
12750
12955
12419
12420
12458
12243
12235
12235
12235
12235
12211
12700
12245
12245
12829
13601
12651
12735
12735
12735
12735
12544
12123
12122
12123
11153
10638
13925
14021
14040
13618
13845
13238
13735
13730
13730
13503
13503
13503
13503
13502
13503
13502
13417
13351
13227
13040
12931
12843
12840
12840
12840
12840
12840
12840
12840
12840
15404
15640
15537
15537
15515
15127
15132
14939
15116
15130
14920
14950
14950
14950
14256
14300
LOCALITY
PETERSBURG
QUATRE FOUR R
SLAVE RIVER
WETASKIWIN
MT GLAVE AT T
ALA HWY MI 590
DEASE L VILLAG
SUMMIT PASS
TETSA RIVER
MACDONALD CR
SIKANNI R
BEATTON RIVER
BEATTON RIVER
BEATTON RIVER
BEATTON RIVER
BEATTON RIVER
MACLURE LAKEe
PRINCE GEORGEes
PRINCE GEORGE>
ANDERSON RIVER
HORNE Le
NORMAN WELLS
RED MOUNTAIN
RED MOUNTAIN
RED MOUNTAIN
RED MOUNTAIN
VIRGINIA FALLS
FORT SIMPSON
FT+ SIMPSON
FORT SIMPSON
FORT SMITH
SASKATOON
DAWSON
SIXTY MI ROAD
SNAG JUNCTION
CARMACKS
KLUANE LAKE
LOWER LAPIE R
PINE CREEK
ALSEK RIVER
ALSEK RIVER
WHITEHORSE
WHITEHORSE
WHITEHORSE
WHITEHORSE
WHITEHORSE
WHITEHORSE
MILES CANYON
ALA HWY MI 882
LITTLE ATLIN L
ALA HWY MI 793
RANCHERIA
RANCHERIA R
WATSON LAKE
WATSON LAKE
WATSON LAKE
WATSON LAKE
WATSON LAKE
WATSON LAKE
WATSON LAKE
WATSON LAKE
WATSON LAKE
UYAK BAY
KING SALMON
COVILLE LAKE
COVILLE LAKE
GROSVENOR LAKE
HOMER
HOMER
HOPE
KENAI
STERHWY MI 125
MOOSE PASS
COOPER*S LANDG
COOPER*S LANDG
COOPER*S LANDG
MCCARTHY
KENNICOTT GL
DATE
19
COLLECTOR NAME
967
927
927
939
967
968
969
943
968
968
943
943
943
943
943
943
967
967
967
965
962
953
939
939
939
939
970
955
955
939
927
962
932
966
956
944
944
944
944
944
944
944
932
932
932
944
958
958
966
943
966
948
968
946
956
956
956
956
966
966
956
966
938
965
954
954
954
961
967
941
951
967
941
943
941
939
961
957
CTITtrPIroaw GQITEZEOTIITECOOOOTIIIIIMMNTAYNOTILT
VSEeSsSenecer
TMPCCCFrFOCTMOAQDC eC KOODOADYVDGDVNODCUCUNITEYVESTPYS
eprztVv0eVv>
ras <=
Zeeeseezeesrre
=Zsesecreese FS &CcvDe
fget£tectztxtcuouronredecredesn
2 ma»
Ver
ARGUS
RAUP
RAUP
RAuUP
ARGUS
WELSH
RIGBY
RAUP
WELSH
WELSH
RAUP
RAUP
RAUP
RAUP
RAUP
RAUP
ARGUS
ARGUS
ARGUS
SCOTTER
CALDER
COpY
RAUP
RAUP
RAUP
RAUP
SCOTTER
CODY
CODY
RAUP
RAUP
ARGUS
BEAUCHAMP
WELSH
ARGUS
COLE
RAUP
PORSILD
RAUP
RAUP
RAUP
ANDERSON
BEAUCHAMP
BEAUCHAMP
BEAUCHAMP
PORSILD
STIELL
STIELL
WELSH
RAUP
WELSH
ANDERSON
WELSH
ANDERSON
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
SUDA
ANDERSON
HARMS
CAHALANE
SCHALLER
SCHALLER
LITTLE»
VIERECK
ANDERSON
KLEIN
VIERECK
ANDERSON
ANDERSON
ANDERSON
SHARPLES
LITTLE»
VIERECK
JRe
JR.
COL NO HF&8B + NO
6A39
GWA
ALA
ALA
ALA
GWA
GWA
OTF
ALA
GWA
GWA
ALA
ALA
ALA
ALA
ALA
ALA
GWA
GWA
5WA
GWA
NAO
DAO
ALA
ALA
ALA
GWA
GWA
ALA
GWA
ALA
ALA
GWA
ALA
GWA
GWA
TSC
ALA
TSC
OLA
ALA
GWA
ISC
ALA
ALA
ALA
1S
SWA
GWA
GWA
ALA
GWA
1SC
GWA
1Sc
ALA
ALA
GWA
GWA
5WA
BWA
GWA
GWA
1SC
ALA
1SC
ALA
ALA
GWA
GWA
1Sc
ISC
GWA
TSC
1Sc
TSC
ISC
GWA
ALA
149475
14476
14826
1151
19827
19830
19838
19833
19834
19829
19831
14861
14855
14862
24337
14825
14477
505
256211
19825
256172
19826
19832
256205
507
510
506
256171
19828
256184
256187
4823
4822
256194
32563
219362
2978
2981
256203
256168
256177
256176
256188
256191
6406
197
Appendix B
TAXON SPC S V HYB PROV QUAD LAT
1873
1873
1873
1873
1875
1873
1873
1673
1873
1875
1873
1675
1873
1873
1673
1873
1673
1873
1473
1873
1673
14673
1873
1873
1673
1873
1873
1473
1875
1673
1873
1873
1673
1873
1873
18673
1673
1873
1873
1873
1573
1873
1873
1873
1873
1875
1873
1873
1873
1473
1675
1873
1873
16735
1s73
1673
1673
1673
1873
1673
1873
1473
1873
ln75
15673
1573
1673
1673
lb75
1873
1673
1673
1673
1673
1673
1673
1673
1573
1873
1673
1673
1673
1673
1673
1873
1873
1673
1673
1673
1873
1873
198
FEF SSF SFE SSS SSS SFE FSF SC HSHEL SSE FFE FSCS SESE SF SEL SCSF FES LES FE FSCS SFE FF SEF SEF SES ELS SS SSFHE FS SCS SEF SESS HES SESS SESS SEL ELS
6130
6131
6153
6147
6149
6147
6132
6132
6132
6128
6128
6150
6148
o132
0140
6140
6140
o1135
6113
6145
6145
6145
6157
6130
6135
6137
6259
6259
6259
6259
6229
6232
6233
6233
6257
e257
6257
6258
6220
6226
6228
6218
6216
6216
6218
6258
0301
6323
6320
6321
6327
6328
6337
6347
6347
6347
6347
6347
6347
6355
0344
6343
6344
0344
6345
o 344
0344
6344
6344
6315
6307
6445
6445
6445
0445
6445
60445
6445
0450
6450
6452
6452
6454
6452
6453
6452
0454
6452
6450
6450
6453
LONG
14300
14426
14602
14511
14512
14511
14914
14914
14914
14922
14922
14731
14628
14914
14900
14900
14900
14954
14954
15005
15003
15003
15111
15934
15930
15930
15604
15604
15604
15604
15503
15337
15337
15337
15536
15536
155356
15535
15006
1511S
15115
14518
14523
14525
14518
14156
14148
14321
14236
14233
14228
14301
14355
14545
14545
14545.
14430
14430
14430
14452
14855
14856
14855
14855
14900
14855
14855
14855
14855
15442
15652
15530
15530
15530
15530
15530
15530
15530
14740
14740
14752
14752
14751
14750
14750
14752
14751
14750
14730
14730
14750
LOCALITY
KENNICOTT GL
CHITINA
ST ANNE LAKE
RICH HwY M87
RICH HwY M90
RICHHWY M 8768
MATANUSKA
MATANUSKA
MATANUSKA
EKLUTNA
EKLUTNA
SHEEP MT
CHICKALOON
MATANUSKA
MATANUSKA VY
MATANUSKA VY
MATANUSKA VY
ANCHORAGE
ANCHORAGE
WILLOW
WILLOW
WILLOW
SKWENTNA
ANIAK
ANIAK
ANIAK
TAKOTNA
TAKOTNA
TAKOTNA
TAKOTNA
BIG RIVER
FAREWELL Le
FAREWELL Le
FAREWELL Le
MCGRATH
MCGRATH
MCGRATH
MCGRATH
TALKEETNA
KAHILTNA GL
KAHILTNA GL
GAKONA
GULKANA
GULKANA
GAKONA
NORTHWAY
NORTHWAY JC
TANACROSS
FORTY MI HOUSE
FORTY MI HOUSE
FORTY WI HOUSE
TOK JUNCTION
JAN LAKE
DONNELY DOME
DONNELY DOME
DONNELY DOME
GEORGE LAKE
GEORGE LAKE
GEORGE LAKE
UPPER TANANA
MCKINLEY NAT
MCKINLEY NAT
MCKINLEY NAT
MCKINLEY NAT
MCKINLEY NAT
MCKINLEY NAT
MCKINLEY NAT
MCKINLEY NAT
MCKINLEY NAT
NIXON MINES
OPHIR
RUBY
RUBY
RUBY
RUBY
RUBY
RUBY
RUBY
FAIRBANKS
FAIRBANKS
SMITH LAKE
SMITH LAKE
GOLDSTREAM CR
COLLEGe
COLLEGE
FAIRBANKS
GOLDSTREAM CR
COLLEGE
FAIRBANKS
FAIRBANKS
COLLEGE
VVUVVVVUVUVUD
DATE
08
07
06
08
08
08
07
07
07
06
06
07
07
07
957
935
961
967
967
967
931
931
931
941
941
948
948
931
940
940
940
965
965
941
941
941
961
949
949
949
941
941
941
940
950
949
949
949
950
949
949
948
941
956
956
944
957
957
966
946
944
963
963
963
7
957
957
957
957
964
964
964
966
954
960
959
95n
950
939
939
959
959
949
949
931
931
931
931
Jey!
lel
931
940
940
956
956
957
957
957
956
957
95/7
956
956
957
COLLECTOR NAVE COL NO
L 4& VIERECK 228
J © ANDERSON 2013
Ek RTT wR. 1A363
L A VIERECK R460
L A VIERECK AYUSY
L A VIERECK R459
J P ANDERSON 796
J © ANDERSON 897
J © ANDERSON RO7
J © ANDERSON 6957
J P ANDERSON 6958
J P ANDERSON 10673
J P ANDERSON ins71
J P ANDERSON 796
L J PALMER 134
L J PALMER 101
L J PALMER 160
S i. WELSH 4141
S t WELSH 4143
J P ANDERSON 7008
J © ANDERSON 7010
J © ANDERSON 7099
E L LITTLEs JR. 18433
WH DRURY 1705
Ww H DRURY 1424
WH DRURY 1461 °
J © ANDERSON 7414
J P ANDERSON 7412
J © ANDERSON 74U14
Es SCAMMAN 1827
Ww H DRURY 4768
WH DRURY 2557
WH DRURY 2284
WH DRURY 2345
WH DRURY 3618
w H DRURY 1356
W H DRURY 1370
R Lt LAYDEN 14
J P ANDERSON 7720
L A VIERECK 1050
L A VIERECK 1050
J P ANDERSON R505
G w ARGUS 1042
G w ARGUS 1042
S t WELSH 5654
G W GASSER SeNe
J P ANDERSON 10195
J P ANDERSON aA7QA20
M BARTHOLOMEW 1863
M BARTHOLOMEW 3963
J NAVA 196
L Af SPETZMAN 879
L A SPETZMAN 2AT
G w ARGUS 1056
G w ARGUS 1056
G w ARGUS 1043
V L HARMS 3175
V L HARMS 3164
V t HARMS 3162
L A VIERECK 7738
G ¥ FROHNE 542n2
A MURIE 10
A MURIE 4
A MURTE 7
A MURIE 9
A NELSON 3527
A NELSON 3587
—E H RENZEL 1A
E 4H RENZEL 18
w 4 DRURY 3356
WU DRURY 3448
C 4 ROUSE 53
C 4 ROUSE 53
C 4 ROUSE &3
C 4 ROUSE 54
C H ROUSE 54
C 4 ROUSE 51
C 4 ROUSE S1
J P ANDERSON 6078
J P ANDERSON 64079
G w ARGUS 725
G wv ARGUS PAL
G ¥ ARGUS 10°94
G w ARGUS 1107
G ™ ARGUS 1NN4
G w ARGUS 725
G w ARGUS 10?°4
G Ww ARGUS 1107
G6 w ARGUS TAT
G “ ARGUS 9R4
G w ARGUS 1004
HERB + NO
GWA
TSC 256198
GWA
BWA
SWA
GWA
ALA 27349
ALA 27351
ISC 258201
TSC 256189
TSC 256174
TSC 256183
TSC 256170
TSC 256199
ALA 5169
ALA 5166
ALA 5160
TSC 246123
TSC 246122
SC 256173
ISC 256202
TSC 256204
SWA
CAN
CAN
CAN
ALA 509
TSC 256193
TSC 256192
GWA
CAN
CAN
CAN
CAN
CAN
CAN
CAN
TSC 256181
TSC 256178
ALA 11624
GWA
TSC 256207
ALA 8402
GWA
SWA
ALA 24997
TSC 256186
TSC 256206
ALA 25882
ALA 25897
ALA 23968
ALA 6872
ALA 6873
ALA 6720
SWA
GWA
ALA 32611
ALA 32605
ALA 32610
FSLC
ALA 21926
5WA
GWA
SWA
Sw
ALA 508
TSC 256190
TSC 228343
TSC 228344
CAN
CAN
ALA 25442
ALA 25461
ALA 2012
ALA 25469
ALA 2n13
ALA 25466
ALA 2014
TSC 256196
TSC 256195
ALA 4569
ALA 4568
ALA 6741
ALA 6721
ALA 6731
SwWA
Sw
GWA
SWA
OWA
SwWA
Specimens Cited
TaAxON SPC S V HYB PROV QUAD LAT LONG LOCALITY DATE
16735 4 14 1lvUg 0453 14751 COLLEGE 14 06
1673 4 14 100 6453 14751 COLLEG= 14 06
1673 4 14 100 6452 14750 COLLEGE 18 05
1873 4 14 10uU 6457 14740 FOX 09 06
1673 4 14 100 6455 14755 COLLEGE 05 06
1673 4 14 100 o452 14750 COLLEGE
1873 q 14 10u 6451 14751 ESTER 21 07
1473 4 14 10u 60452 14752 UNIV EXP FARM 26 05
1573 4 14 100 6452 14752 UNIV ExP FARM 10 09
1873 4 14 #10U 6452 14750 COLLEGE 14 07
1873 4 14 100 6450 14740 FAIRBANKS 17 06
18675 4 14 100 ©6450 14740 FAIRBANKS 17 06
1673 4 14 100 6450 14740 FAIRBANKS 06
1873 4 44 1200 6450 14740 FAIRBANKS 06
1875 4 14 10u 6%50 14740 FAIRBANKS 06
16735 4 14 100 60450 14740 FAIRBANKS 06
1873 4 14 100 6450 14740 FAIRBANKS 06
1873 4 14 100 6450 14740 FAIRBANKS 06
1873 4 14 100 6450 14740 FAIRBANKS 06
1873 4 14° #100 ©0450 14740 FAIRBANKS 06
1873 4 14 100 6450 14740 FAIRBANKS 17 06
1673 4 14 #10y 6450 14740 FAIRBANKS 17 06
1873 4 14 100 6450 14740 FAIRBANKS 20 06
1873 4 14 100 6450 14740 FAIRBANKS 20 06
1873 4 14 100 6450 14740 FAIRBANKS 20 06
1375 4 14 100 6450 14740 FAIRBANKS 06
1873 4 14 100 6450 14740 FAIRBANKS 20 07
1573 4 14 100 ©6450 14740 FAIRBANKS 06
1873 4 14 100 6450 14740 FAIRBANKS 20 07
1673 4 14 100 6450 14740 FAIRBANKS 20 07
1673 4 14 100 6450 14740 FAIRBANKS 06
1673 4 14% #$§4JO00 6450 14740 FAIRBANKS 06
1673 4 14 100 6450 14740 FAIRBANKS 06
1673 4 14 100 60450 14740 FAIRBANKS 06
1873 u 14 100 6450 14740 FAIRBANKS 06
1873 4 14 100 6450 14740 FAIRBANKS 06
1873 4 14 100 6450 14740 FAIRBANKS 20 06
1873 4 14 100 6450 14740 FAIRBANKS 06
1673 + 14 #100 6450 14740 FAIRBANKS 06
1873 4 14 100 0450 14740 FAIRBANKS 06
1873 4 14 #100 0450 14740 FAIRBANKS 06
1573 4 14° #100 6450 14740 FAIRBANKS 06
1873 4 14 100 6450 14740 FAIRBANKS 06
1673 4 44 100 6450 14740 FAIRBANKS 06
1873 4 14 100 6450 14740 FAIRBANKS 06
1673 4 14 104g 6451 14748 FAIRBANKS 2f- 07
1875 4 14 10u 6453 14751 COLLEGE 05 06
1673 4 14 10u 60447 14745 COLLEGE 06 06
18673 4 14 10y 6450 14740 FAIRBANKS 07 06
18673 uy 14 100 o447 14745 COLLFGE 23 07
1873 4 14 100 6452 14752 UNIV EX® FARM 29 07
1573 4 14 100 6452 14752 UNIV EXP FARM 29 07
1673 4 14 100 6452 14752 UNIV EXP FARM 29 07
1673 4 14 #4100 6452 14752 UNIV EXP FARM 17 06
1873 4 14 101 6456 14640 CHENA RIVER 06 07
14673 4 14 101 6456 14640 CHENA RIVER 06 07
1673 4 14 101 6410 14551 BIG DELTA 01 06
1573 4 14 101 56458 14657 UPPER CHENA R- 25 05
1873 4 14 $4101 6458 14657 UPPER CHENA R- 25 05
1673 4 14 104 6545 14430 CENTRAL 15 06
1873 4 14 104 ©6545 14430 CENTRAL T5706
1873 4 44 104 6545 14430 CENTRAL 15 06
1673 4 14 104 6532 14513 MILLER HOUSE \2 07
1873 4 14 104 6532 14513 MILLER HOUSE 112° 07
1873 4 14 104 6523 14554 STEEHWY MI 90+ 25 06
13873 4 14 105 6503 14726 CLEARY SUMMIT 16 07
15673 4 14 #105 6532 14833 LIVENGOOD 08 07
1873 4 14 #105 ©6532 14833 LIVENGOOD 08 07
1673 4 14 105 0532 14833 LIVENGOOD 17 06
1873 4 14 105 0532 14833 LIVENGOOD 17 06
1873 4 14 106 6500 15038 MANLEY HOT SPR 05 07
1573 4 14 106 6500 15038 MANLEY HOT SPR 25 06
15673 4 14 113 6634 14516 FORT YUKON 24 05
1873 4 14 120 6644 14334 BLACK RIVER O04 O7
1873 4 14 120 6644 14334 BLACK RIVER 06 07
1873 4 44 12u 6644 14334 BLACK RIVER 03 07
1573 4 14 120 6644 14334 BLACK RIVER Of O07
1673 4 14 #120 0644 14334 BLACK RIVER 06 07
1873 4 14 #120 0644 14334 BLACK RIVER 14 07
1873 4 14 121 ©0710 14155 HOWLING DOG RK 12 08
1673 4 14 121 ©6710 14155 HOWLING DOG RK 12 08
1873 4 14 121 6710 14155 HOWLINS DOG RK 12 08
1873 4 14 121 6710 14155 HOWLING DOG RK 12 O08
1673 4 14 121 6711 14155 HOWLING DOG RK 26 07
1673 4 14 121 6709 14140 PORCUPINE R ee OF
1873 4 14 121 ©6716 14138 PORCUPINE R 16 08
1673 4 14 121 6716 14138 PORCUPINE R 10 08
14673 4 14 121 6709 14140 PORCUPINE R 12 08
1873 4 14% $4121 6716 14138 PORCUPINE R 10 08
1873 4 14 121 6702 14255 PORCUPINE R 18 08
18673 7 14 121 6oF02 14235 PORCUPINE 8 18 08
COLLECTOR NAME
MAMRMAMRMRUNCCCECCECECHUKHHVNGDOMOHCHKHUNECCHMMMCCECFrFKrHM@eaFerOrCreEMUUASaCTreer Cee crercrerrrererreEeeeccerererererrr. Greece eee a ae
9 Mea Wo ee Mp Nee Ladies
ARGUS
ARGUS
GUILSHER
HARMS
HARMS
HARMS
HARMS
HARMS
JOHNSON
OKEDARA
PALVER
PALMER
PALMER
PALMER
PALMER
PALMER
PALMER
PALVER
PALMER
PALVER
PALMER
PALMER
PALYER
PALMER
PALME
PALMER
PALVER
PALYER
PALVER
PALMER
PALMER
PALME?
PALVER
PALMER
PALMER
PALVER
PALMER
PALMER
PALVER
PALMER
PALVER
PALMER
PALMER
PALMER
PALVER
CHUCK
SMITH
SMITH
SMITH
VIERECK
VIERECK
VIERECK
VIERECK
VIERECK
ARGUS
ARGUS
LITTLE»
VIERECK
VIERECK
BUCKLEY
BUCKLEY
BUCKLEY
SCAYMMAN
SCAMMAN
SHETLER
ANDERSON
ANDERSON
ANDERSON
SMITH
SMITH
ANDERSON
WELSH
EETTELE?
SHETLER
SHETLER
SHETLER
SHE TLER
SHETLER
SAETLER
BUCKLEY
BUCKLEY
BUCKLEY
BUCKLEY
BUCKLEY
RUCKLEY
BUCKLEY
DEAN
DEAN
DEAN
DEAN
DEAN
JR.
cOL NO
313
779
4
2658
24P6
3yu2
29945
3345
yang
2nn
20
1°09
141
163
175
172
190
162
143
2n9
ang
17A7
1790
HF-QR + NO
GWA
CWA
ALA 3102
ALA 32645
PLA 32629
ALA 32635
ALA 32623
"LA 32631
GWA
ALA 18738
BLA 3233
PLA 5246
ALA 5230
ALA 5232
ALA 5234
ALA 5230
ALA 5238
ALA 5243
ALS 5233
ALA 5240
ALA 5250
ALA 32132
ALA 5922
ALA 5792
ALA 594n
ALA 5245
ALA 5977
ALA 5237
ALA 5907
ALA 5973
ALA 5229
ALA 5235
ALA 5241
ALA 65242
ALA 5235
ALA 5°31
ALA 5184
ALA 5263
ALA 5249
PLA = 5248
ALA 5244
ALA 5247
ALA 5253
ALA 5251
ALA
ALA 26176
ALA 10736
ALA 10740
ALA 10742
FSLC 273
FSLC 47
aSkc
ESLE 43
FSLC 281
Sw
SWA
SWA
SWwA
SWA
ALA 5940
ALA 5041
ALA 26817
ALA 8399
Sw
ALA 3869
’SC 256197
TSC 256209
TSC 256208
ALA 10735
aWA
TSC 256175
TSC 246329
SWA
ALA 3AA1
ALA 3905
ALA 3270
ALA 3933
ALA 3908
ALA 4001
ALA 5035
ALA 5931
ALA 26759
ALA 5927
4LA 5995
ALA 5079
ALA 5027
ALA 24043
ALA 24098
ALA 2409%
ALA 21522
ALA 24105
199
Appendix B
TAxON SPC S V HYB PROV QUAD
1673
1873
1675
1873
200
e+e fe
CANDIDA
GUM annrnrnrnrrrrannronnrnnvcw
INTERIOR
14
14
14
14
NNNANN NNN NNN NP eee
lei
124
124
l2o
LAT
0723
6725
6725
0706
9925
5926
6125
5336
6815
6335
6105
6041
6007
0043
6043
6043
6043
6253
6433
6433
6400
6400
6400
6400
6400
0400
6400
6449
6625
6635
5908
5854
5642
5504
5936
5714
5714
6833
6728
6517
6251
6251
0132
6158
6152
6152
6005
6404
6404
6404
6404
6404
6404
6322
6010
6003
6003
6003
0002
6002
6324
6353
6403
6451
6451
6452
6451
6451
6451
6451
6451
6451
6452
6430
6430
6412
6443
6439
6439
6550
6550
6557
6530
LONG
14350
15007
15007
15815
12604
12605
12636
10554
13850
13545
13516
13736
12843
13503
135035
13503
13503
14133
14705
14703
14530
14530
14530
14530
14530
14530
14530
14405
14716
14505
11126
11135
11120
11717
12657
12243
12243
12828
13053
12651
12123
12123
12520
12145
12122
12122
12347
15925
13925
13925
13925
13925
13925
13642
13442
12855
12855
12855
12835
12835
14321
14451
15055
14743
14743
14750
14743
14743
14743
14743
14743
14743
14750
14905
14905
14910
14809
14908
14825
14404
14404
14408
15010
LOCALITY
SMALL LAKE
WISEMAN
WISEMAN
ONION PORTAGE
LIARD HOT SPR
LIARD HOT SPS
Se NAHANNI Re
EMMA LAKE
OLD CROW FLATS
MAYO
DAWSON RD
MACKINTOSH
WATSON LAKE
WHITEHORSE
WHITEHORSE
WHITEHORSE
WHITEHORSE
DEADMAN LAKE
SALCHA SLOUGH
SALCHA SLOUGH
CLEARWATER
CLEARWATER
CLEARWATER
CLEARWATER
CLEARWATER
CLEARWATER
CLEARWATER
CIRCLE
GEORGE LAKE
FORT YUKON
DADAAADADDA
SLAVE R
QTR FOURCHES R
FORT MCMURRAY
VALLE YVIEW
COAL R
SIKANNI R
SIKANNI R
ANDERSON RIVER
THUNDER RIVER
NORMAN WELLS
FORT SIMPSON
FORT SIMPSON
Se NAHANNI R
FTe SIMPSON
FTs SIMPSON
FORT SIMPSON
LIARD RIVER
DAWSON
DAWSON
DAWSON
DAWSON
DAWSON
DAWSON
STEWART R
CARCROSS
UPPER LIARD R
UPPER LIARD R
UPPER LIARD R
WATSON LAKE
WATSON LAKE
TANACROSS
TANANA R
BEARPAwW MT
FAIRBANKS
FAIRBANKS
COLLEGe
FAIRBANKS
FAIRBANKS
FAIRBANKS
FAIRBANKS
FAIRBANKS
FAIRBANKS
COLLEGE
NENANA
NENANA
NENANA
SAM CHARLEY IS
NENANA
WHISKEY IS
CIRCLE
CIRCLE
YUKON R
RAMPART
DATE
31
10
14
31
14
07
31
03
29
20
20
05
16
23
11
11
28
11
20
27
21
06
01
06
07
07
06
07
957
962
56?
967
956
968
970
965
970
967
960
957
943
944
944
944
944
96R8
922
922
956
956
956
956
956
956
956
963
961
963
927
927
939
956
94x
943
943
965
947
953
939
939
970
955
955
955
961
914
914
914
914
914
916
960
960
966
966
966
943
943
957
965
964
956
956
957
956
956
956
956
956
956
957
941
965
965
966
965
965
935
963
965
961
COLLECTOR NAMF
aoOnaDzrwW
CrraqmagngagaqmMoounrrrrizacwAayvnooung
Errerrriercer OaMOOOOOVOOOOFrFOTrFTFIAMVANGOFTrcCSTHYHYP YF FP rPzZTZTFEZOITICAZrPOTIITSeIrTIT
Siac. SS
eq nzee
er ia
Patz e£ectecztecetcCtetervrvrrece t€£tzrrsd
a> PUPYr.->>DpD
SHETLER
SROCKMAN
BROCKMAN
SCHWEGER
ARGUS
WELSH
SCOTTER
ARGUS
WELSH
PORSILD
CALDER
CHOFIELN
RAuP
PORSILD
PORSILD
PORSILD
PORSILD
WELSH
MURIE
MURIE
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
SPETZMAN
SPETZMAN
TRENT
RAyeP
RAUP
RAUP
ARGUS
RAuUP
RAUP
RAUP
SCOTTER
PORSILD
copy
RAU?
RAYS
SCOTTER
CODY
CODY
CODY
CODY
EASTWOOD
EASTWOOD
EASTWOOD
EASTWOOD
EASTWOOD
MALTE
CALDER
SPETZMAN
ARGUS
ARGUS
ARGUS
RAUP
RAuP
SPETZMAN
VIERECK
VIERECK
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
COLE
HARMS
SCHMUCK
VIERECK
VIERECK
VIERECK
ANDERSON
SPETZMAN
VIERECK
DEAN
cOL "O 4FRA + NO
ROP AF
141
247
74unh
12753
4R68
10478
6F5
25077
8310
11029
9147
10663
1663
9147
7ORL
794
797
B26
795
794
826
B?6
4970
4631
RB
563
564
9043
2AlL
190914
19160
10159
6908
16745
7492
9049
9070
12472
8157
B273
R391
11492
465
145
131
342
33
24909
4541
5032
5038
5027
10°70
10970
501
7781
7335
BRT
4x3
1090
431
BR7
BR7
BAT
423
423
1090
SeNe
3722
2u7
8037
7688
7642
25P1
4969
7707
78
ALA 4191
ALA 28480
ALA 28484
ALA 34721
ALA 4763
GWA
SWA
SWwA
BRY 91685
CAN 312744
ALA 17425
CAN 269643
CAN 280494
6H
6H
CAN
CAN
SwA
US 1119998
WS 11199748
BWA
SWA
SWA
ALA 4756
ALA 4593
ALA 4621
ALA 22457
CAN 299615
CAN 299616
ALA 25740
ALA 14488
ALA 14487
ALA 16405
ALA 4698
ALA 19513
ALA 19518
ALA 19617
SWA
GWA
SAO 21405
ALA 16404
ALA 16408
GWA
GWA
SWA
ALA 24395
GWA
CAN 45511
CAN 45508
CAN 45509
CAN 45506
CAN 45507
CAN 122155
ALA 17427
CAN
GWA
SWA
SWA
SWA
ALA 19519
CAN
FSLC
FSLC 304
‘EWA
SWA
SWA
SWA
ALA 4B24u
ALA 4758
ALA 22460
ALA 4517
4LA 22491
ALA 6716
Ysc 256691
ALA 32585
ALA 29849
GWA
GWA
FSLC
TSC 256690
CAN
ESUC
ALA 19084
Specimens Cited
TAXON SPC S V HYB PROV QUAD LAT
1873
1s73
1875
1673
1673
13873
1873
1873
1673
1873
1573
1673
15
1673
1673
1673
1575
1673
10673
lo73
1673
1673
1675
16
1875
19
1873
1873
1873
1873
13873
1873
1673
1373
1473
1873
1673
1873
1673
43875
1873
1673
1573
1873
1873
1873
20
1673
1673
1873
1673
14675
14673
1673
1673
1673
1673
1673
1873
1673
1873
1673
1673
1lo735
1673
1873
1873
1873
1673
1873
1873
1673
1873
1873
1673
12 14
12 14
12 44
12 14
12 14
12 14
l2 14
12 14
12 14
12 14
le 14
12 14
PEDICELLARIS
15 7
15 7
15 7
15 11
15 11
15 12
15 12
£5 12
15 12
15 12
PENTANDRA
16 14
PYRIFOLIA
19 1
19 2
19 2
19 5
19 7
19 id
19 7
19 i
19 7
19 7
19 i
19 7
19 7
19 7
19 7
19 7
19 7
19 7
19 12
19 12
RIGIDA
20 2
20 2
20 2
20 2
20 2
20 2
20 7
20 7
20 7
20 12
20 12
20 12
20 12
20 12
106
119
119
121
Lei
i21
121
121
121
121
121
121
345
369
588
0510
6634
0644
e711
6711
6704
6711
6711
6711
6704
6704
6715
v2e27
6112
6055
5237
5237
6003
6003
6003
6003
6003
5648
5430
5830
5320
4720
6310
o227
0227
0227
622U
6220
0220
6121
0110
6135
6131
6007
6055
6006
6612
6612
5936
5939
5714
5714
5530
5320
6517
0127
6005
6201
6043
6043
6003
0003
LONG
15204
14516
14431
14155
14155
14230
14155
14155
14155
14230
14230
14140
11421
12346
12330
10356
10356
12840
12840
12840
12840
12840
13258
11715
12245
12245
5256
11310
11421
11421
11421
11308
11308
11308
11739
11155
11045
10806
12348
12330
106235
13625
13625
12643
12656
12243
12243
12245
12245
12651
12542
12347
132350
13505
13503
12855
12855
PLANIFOLIA SSPe* PLANIFOLIA
™
©
ee ee ee ee ee
NVNNNNNNNNNNNNFE
i
FOOCOUF FFE NNNHARNCG
~
5854
5941
5941
5941
5936
5936
59558
5958
5958
5958
5953
5951
5951
5822
11154
13652
15632
13632
13627
13627
13206
13206
13206
13206
13127
12908
12908
12955
LOCALITY
TANANA
FORT YJKON
PORCUPINE
PORCUPINE
PORCUPINE
PORCUPINE
PORCUPINE
PORCUPINE
PORCUPINE
PORCUPINE
PORCUPINE
PORCUPINE
DADDADAADAADAD
YELLOWANIFE
YOHIN LAKE
BLUE BILL CR
MCKAGUE
MCKAGUE
WATSON
WATSON
WATSON
WATSON
WATSON
LAKE
LAKE
LAKE
LAKE
PETERS3URGe
VALLEYVIEW
PROPHET R
PRINCE GEORGE.>
BIG COUNTRY PD
GORDON LAKE
YELLOWANIFE
YELLOWKNIFE
YELLOWKNIFE
HEARNE LAKE
HEARNE LAKE
HEARNE LAKE
FT PROVIDENCE
RUTLEDGE Re
wW RUTLEDGE Le
PORTER LAKE
LIARD RIVER
LIARD RIVER
SCOTT LAKE
PALMER LAKE
PALMER LAKE
COAL RIVER
LIARD R
SIKANNI R
SIKANNI R
PINE RIVER+
PRINCE GEORGE.
NORMAN WELLS
FLAT RIVER
FORT LIARD
PELLY RIVER
WHITEHORSE
WHITEHORSE
UPPER LIARD R
UPPER LIARD R
QTR FOURCHES R
CHILKAT PASS-e
CHILKAT PASSe
CHILKAT PASSe
RD M68
RD M68
HAINES
HAINES
MORLEY
MORLEY
MORLEY LAKE
MORLEY LAKE
SWAN LAKE
LAKE
LAKE
CREEK
19
06
13
13
13
06
06
17
17
iy
17
31
CASSIAR RD M16 30
CASSIAR RD M16 30
DEASE L VILLAG 13
DODDNNNDWANAANANA DBNODWDW
925
961
926
951
957
957
251
S57
957
957
957
957
970
961
938
938
966
966
966
966
943
967
956
966
967
969
961
963
961
961
961
961
961
961
966
962
966
961
961
966
960
960
960
943
943
943
967
967
953
970
961
944
944
944
966
966
oT
967
967
967
956
956
960
960
960
960
960
960
960
969
COLLECTOR NAWF
GCCacececaeacec amr
alt ol call aul sel (oa al oe a | ona
LANDDOFP=zAOC
CCEFFEOECOHOOOCCCAADVNOAD
QOOPrPrrz Qn ztQoortTiIitc
AQCCCCGCcCataaoaaz
=>
i a og Oe
= €
PALMER
LITTLEs JR.
MURIE
BUCKLEY
BUCKLEY
BUCKLEY
BUCKLEY
BUCKLEY
BUCKLEY
BUCKLEY
BUCKLEY
BUCKLEY
LARSEN
SCOTTER
CODY
BREITUNG
BREITUNG
ARGUS
ARGUS
ARGUS
ARGUS
.RAUP
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
SCOTTER
LARSEN
THIERET
THIERET
COTTER
SCOTTER
SCOTTER
THIERET
CODY
SCOTTER
CODY
CODY
COpY
copy
CALDER
CALDER
CALDER
RAUP
RAuP
RAUP
ARGUS
ARGUS
CODY
SCOTTER
CODY
PORSILD
PORSILD
PORSILD
ARGUS
ARGUS
RAUP
ARGUS
ARGUS
ARGUS
TAYLOR
TAYLOR
CALDER
CALDER
CALDER
CALDER
CALDER
CALDER
CALDER
RIGBY
COL NO
1282
1Aqg*0
3
SeNe
nu
ats
SeNe
204
2n4
211
211
113A
SeNe
12651
11960
SeNe
SeNe
5054C
549
5046
5047
1146
6A37 *
260
4°67
6079
7481
1248
7290
6703
967
939
64
6616
16121
3043
15541
11626
11°83
16755
26099
26099
25282
11510
1n4n8
1158
669
6973
7625
12541
11493
ORG
1n662
9155
5076
5029
515
6°16
6A1R
6P17
ARB
BRA
24461
24463
24463
24u4ué6l
24uaul
24uRR2
24832
71
HERB + NO
ALA 2035
BWA
ALA 535
5WA
CWA
SWA
ALA 5029
ALA 5996
ALA 26779
ALA 5976
ALA 26763
ALA 5990
GWA
GWA
GWA
CAN 49130
CAN 49132
GWA
SWA
GWA
GWA
CAN 280860
SWA
5WA
GWA
SWA
GWA
DAO 36306
GWA
GWA
OWA
DAO 36304
LAO 36305
DAO 36303
GWA
DAO 36310
DAO 36308
DAU 36312
DAO 36309
DAO 36307
DAO 36311
DAO 36302
WTU206384
SWA
ALA 20057
ALA 20958
ALA 20056
SWA
SWA
‘AO
SWA
6WA
C4N 47607
CAN 47605
CAN 47606
SWA
GWA
ALA 14489
SWA
GWA
GWA
‘AO 32745
9490 32747
SwA
6WA
SA0 32775
"AO 32766
AO 19073
GWA
NAO 32770
NTF 1149
201
Appendix B
TAXON SPC S V HY¥B PROV QUAD LAT LONG LOCALITY DATE
1873 28 1 02 16 5831 12434 SUMMIT PASS 19 07
1573 28 1 v2 17 5848 12332 STEAMBOAT 08 05
2575 428. 1 02 17 5848 12332 STEAMBOAT 08 05
1873 28 1 v2 17 5848 12332 STEAMBOAT 08 05
1873 28 1 02 17 5848 12332 STEAMBOAT 08 05
1873 26 1 v2 17 5848 12332 STEAMBOAT 08 05
1873 28 1 02 17 58468 12332 STEAMBOAT 08 05
la7s)0 (2a 1 02 17 5848 12332 STEAMBOAT 08 05
La73S 128.1 v2 17 5848 12332 STEAMBOAT 08 05
Lu73 “128.1 02 24 5705 12235 BEATTON RIVER 13 06
1873 28 1 02 24 5705 12235 BEATTON RIVER 10 06
1673 28 1 02 24 5705 12255 BEATTON RIVER 10 06
1373 28 1 02 24 5705 12235 BEATTON RIVER 10 06
16735 “26.2 02 24 5705 12235 BEATTON RIVER 12 06
MA7S 28) 8 v2 24 5705 12235 BEATTON RIVER 10 06
1873 281 v2 31 5643 12133 BLUEBERRY 26 05
1575 2B 1 v2 79 4905 12011 CATHEDRAL L 04 08
1873 281 O07 241 6517 12651 NORMAN WELLS 04 06
is73 928 1 07 241 6517 12651 NORMAN WELLS 17 07
1A73 26 1 07 275 6432 12635 STERILE LAKE 23 07
ts73 ves 1 07 275 6431 12634 STERILE LAKE 01 07
1873-28 1 07 313 6335 11002 MUNN LAKE 03 08
1673, “ee. 1 07 316 6302 11057 INDAIN YT L 08 08
1873 28 1 07 338 6300 12902 0 GRADY LAKE 25 07
13873 28 1 07 338 6222 12842 SELWYN MTS 09 08
1873 281 07 339 6205 12735 BRINTNELL LAKE 20 06
is7s. _ 28 1 07 339 6205 12735 BRINTNELL LAKE 20 06
13873 28 1 O07 345 ©6227 11422 YELLOWKNIFE 04 07
1673 28 1 07 345 ©6227 11422 YELLOWKNIFE 02 07
in7s) 26,1 07 345 6227 11422 YELLOWKNIFE 23 06
1873 28 1 07 345 6227 11422 YELLOWKNIFE 13 06
fa75 268) 1 O07 345 6227 11422 YELLOWKNIFE 12 06
1673 28 1 U7 345 6227 11422 YELLOWKNIFE 28 06
1873 28 1 O07 345 6227 11422 YELLOWXNIFE 18 06
T7528) 2 O07 345 6227 11422 YELLOWKNIFE 23 06
1673 28 1 O07 345 6227 11422 YELLOWKNIFE 17 06
1673) 28 1 07 345 6227 11422 YELLOWKNIFE 13 06
1673 281 07 346 6©220 11308 HEARNE LAKE 17 06
1873. 228 2 07 349 6220 10625 LYNX LAKE 23 07
1873s 28 1 07 350 6218 10425 MANTIC LAKE 26 07
157328 1 7 368 6108 12427 Se NAHANNI Re 2 FT
1873028 1 07 369 6117 11947 TROUT RIVER 30 06
16735 p28 1 07 370 ©6147 12043 FORT SIMPSON 07 06
13873 28 1 07 370 652 12122 FORT SIMPSON 13 06
1873 281 07. 370 6152 12122 FORT SIMPSON 07 06
1873 28 1 07 370 6152 12122 FORT SIMPSON 27 06
1573 28 1 07 37u 60152 12122 FORT SIMPSON 13 06
1873 28 1 07 370 6152 12122 FORT SIMPSON 15 06
1875 261 07 S70 6152 12122 FORT SIMPSON 08 06
1673. 28 1 07 370 6152 12122 FORT SIMPSON 11 06
ls73 28 1 07 3570 6152 12122 FORT SIMPSON 07 06
1873 28 1 O07 371 6128 11801 HORN R DELTA 18 08
1873 28 1 7 $71 6117 11947 TROUT RIVER 30 6
$573 "528 07 573 6143 11453 LONG ISLAND 22 07
1673-28 1 07 374 6123 11245 ROCHER RIVER 01 08
1873-28 1 O07 374 6104 11307 MCCONNELL IS 03 08
1673 28 1 07 374 6118 11339 SLAVE RIVER 16 06
1673 28 1 07 375 6155 11010 N GAGNON LAKE 29 06
1673 26 1 07 375 6135 11045 E RUTLEDGE L 01 07
i573)" ees 1 07 576 6131 10806 PORTER LAKE 27 07
1473. 268 1 07 376 6131 10806 PORTER LAKE 27 07
1873 28 1 07 388 6014 12323 FORT LIARD 25 07
1473 28 1 07 391 6058 11720 KAKISA RIVER 16 06
1573 281 07 391 6058 11720 KAKISA RIVER 16 06
1875 28 1 07 391 6058 11720 KAKISA RIVER 16 06
1473 28 1 07 391 6056 11720 KAKISA RIVER 16 06
14673 28 1 U7 $391 ©0053 11721 KAKISA LAKE 21 06
1473 26) 1 07 392 6051 11542 HAY RIVER 28 05
iB73~ »28 1 07 392 6951 11542 HAY RIVER 03 06
1873 28 1 07 292 6051 11542 HAY RIVER 03 06
1673 28 1 07 592 ©6051 11542 HAY RIVER 11 06
1473 28 1 07 394 6003 11200 FT SMITH 20 07
1473 28 1 07 594 6003 11200 FT SMITH 20 07
1873 28 1 07 394 6002 11238 FT SMITH 02 06
1673 28 1 07 394 6000 11153 FT SMITH 31 05
1473 28 1 U7 394 6052 11034 LADY GREY L 31 07
1673 28 1 U7 395 6006 10623 SCOTT LAKE 14 08
1473 281 07 395 6006 10623 SCOTT LAKE 14 08
18735: «28 1 07 596 6047 10738 SPEARFISH LAKE 30 07
16735 26 1 07 397 ©6100 10500 SMALL TREE L 23 07
1873 28 1 07 402 6050 9425 MCCONNELL R 02 07
1873 281 07 402 6050 9425 MCCONNELL R 03 07
e795 +26 1 07 402 6050 9425 MCCONNELL R 06
1873 28 1 07 402 6050 9425 MCCONNELL R 26 06
1873 26 1 07 402 6050 9425 MCCONNELL R 10 07
fe73 26 1 U7 402 6050 9425 MCCONNELL R 10 07
1873 28 1 12 32 6115 13844 ALA HWY M1073 07 08
1873 28 1 12 32 6122 13843 KLUANE LAKE 08 07
1873 281 12 32 6122 13859 BURWASH 03 08
1673 281 12 34 6137 13553 CONGLOMERATE M 14 05
1673 28 1 12 34 6137 13553 CONGLOMERATE M 14 05
202
943
960
960
960
960
960
960
960
960
943
943
943
943
943
943
960
956
957
953
967
967
966
966
967
967
939
949
949
949
949
949
949
949
949
961
961
961
966
966
970
961
955
955
955
961
955
955
955
955
955
956
961
951
965
965
927
962
962
966
966
961
959
959
959
959
959
951
951
951
951
965
965
950
950
965
966
966
966
961
964
964
964
964
967
967
948
944
948
960
960
COLLECTOR NAWF
CCLTTLPTAARKRAAKAACZFSFAZTAZTTSFFTFSFCCCCCZTA*SOAOOMOTLFEAFBPOAFEEFZAATZLAFAOAZAADOQCCAWZAEAZAEARZEIITAZAAZAEZSZZFZAZACCIIIIAIIOCcCcCcccccirTt
<>>vpvr,vpr,> >
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<¢
(1S
PE Can er, es a eet
a
1S >
3 GEES PLES Ls
a ee
=<
r
a me Jae | Geel Gaels f'n Wey (ee Care oe ee
Pree
RAUP
CALOER
CALDER
CALDER
CALDER
CALDER
CALDER
CALDER
CALDER
RAUP
RAUP
RAuP
RAUP
RAyYP
RAUP
CALDER
CALDER
CODY
CODY
SIMMONS
SIMMONS
CODY
CODY
CODY
copy
RAU?
RAUP
COnY
Copy
CODY
CODY
CODY
CODY
CODY
HALL
THIERET
THIERET
SCOTTER
COpyY
CODY
SCOTTER
CODY
CODY
CODY
CODY
CODY
CODY
CODY
CODY
CODY
CODY
BIRD
CODY
LEwIS
CODY
Copy
RAUP
SCOTTER
SCOTTER
CODY
CODY
copy
THIERET
THIERET
THIERET
THIERET
THIERET
LEwIsS
LEwIS
LEwIsS
LEwIS
CODY
CODY
CODY
CODY
CODY
CODY
CODY
CODY
MAINI
MACINNES
MACINNES
MACINNES
MACINNES
MACINNES
MACINNES
RAUP
RAyeo
RAuUP
CALDER
CALDER
cOL NO
1966?
24U3A2
24359
24360
24361
2uURA?
24359
243460
24u3AlL
10075
1N0?7
innel
10013
10043
19022
H=QR + NO
ALA 20n4n
Gwa
SWA
GWA
GWA
“AQ 32778
DAO «32769
DAO 32768
NAO 32777
ALA 22N23
ALA 20036
ALA 20029
ALA 20935
ALA 20034
ALA 20030
DAN 32773
YAO 19320
NAD 32623
YAO 32583
YAO 32592
“40 32591
NAO 32580
S40 32578
NAO 32663
MAO 32574
ALA
ALA 14843
YAO 32627
"AQ 32626
NAO 32594
NAO 32625
NAO 32582
NAO 32630
NAO 32628
YAO 32629
NAO 32600
949 32601
YAO 32604
YAO 32575
YAO 26489
GWA
YAO 32620
AQ 32617
YAO 32613
AO 32612
NAO 32622
NAO 32619
NAO 32614
YAO 32616
JA0 32615
AO 32618
QAO 32593
LAO 3262
NAD 32590
YAO 32595
980 632599
ALA 14606
YAO 32602
‘ADO 32603
HAO. S257 7
NAO 32579
940 32606
NAO 32611
QAO 32607
940 32609
NAO 32610
YAO 32608
340 632585
NAO 32587
‘AO 32588
YAO 32586
NAD 32598
NAO 32597
YAO 32584
SAO 32575
YAO 32596
DAO 32581
YAO 26492
AO 26490
NAO 32605
Uwo
Uwo
UwO
UwO
Uwo
Uwo
ALA 20025
ALA 20031
ALA 20024
SWA
YAO 32757
Specimens Cited
TAXON SPC
1875 28
1875 28
1875 28
1873 28
1873 28
1873 28
1875 28
1873 28
1875 28
1875 28
1873 28
1875 28
1875 28
1873 28
1875 28
1875 28
1875 28
1873 28
1873 28
1873 28
1673 28
1875 28
1875 28
1875 28
1873 28
28 3 2 PLANIFOLIA SSPe
1873
1875
1875
1873
1873
1875
1873
18735
1873
1873
1873
1875
1873
"8735
1873
1873
1873
1875
18735
1875
1873
1875
1873
1675
1875
1875
1875
1875
18753
1875
1873
1873
1873
18753
1675
1873
1875
1875
1875
1875
18735
1875
1875
1875
1873
1875
18753
1873
1873
1875
1875
1873
1873
1873
1875
1673
18753
1673
1873
18753
1873
1873
28
28
28
28
28
S V HYB PROV QUAD LAT
ee
Ot OF Ot GI Gi Ol GI OF OF GI OI GI Ol GI OF GI Gi Gi GO OF Gl Gi G8 OF Gi GO! OF OF GI OF OI OF GO! OF Gi OF OF OF OF Oi OF GO! Gi OF OF OF Gf OF G8 G8 OF GI OF GI Gi OF GI Ol OF Ol OF
NNNNNNNNNNANN NDNA NNNNNNANNNNNNNNMNNNANNANNANNNNNNNANNNNANANMWNNM NNN NW NW NW
02
02
02
02
07
40
40
40
40
40
40
41
41
41
41
41
41
41
41
41
41
42
43
43
43
44
44
44
44
44
100
6045
6047
6047
6047
6047
6047
6010
6010
6010
6010
6043
60435
6038
6053
6049
6045
6003
6005
6005
6005
60035
60035
60035
6003
6007
PULCHRA
5941
5941
5941
5935
7015
6931
6938
6942
6956
6921
6921
6842
6842
6906
6819
6854
6852
6852
6906
6815
6815
6842
6842
6842
6842
6842
6813
6845
6820
6750
6300
62435
6812
6825
6725
6725
6725
6404
6404
6404
6404
6405
6300
6252
6111
6051
6022
6150
6142
6205
6225
6252
6335
6327
6323
6323
6347
6356
6356
6500
6431
6451
LONG
13615
13735
13735
13735
13735
13736
13442
13442
13442
13442
135035
135035
13500
13532
13519
13503
13218
13035
13035
13035
12840
12840
12852
12852
12931
VARe
13632
13632
13632
13629
12940
13348
13254
129
12858
12815
12813
13408
13408
13037
13340
13314
13205
13205
1352357
13500
13500
13408
13408
13408
13408
13408
13547
13412
13330
11510
12902
10909
13845
13800
14100
14100
14100
13925
13925
13925
13925
14019
13012
13015
13512
13747
13351
15617
15721
16343
14451
14128
14216
14228
14345
14330
14430
14655
14655
15038
15012
14743
LOCALITY
MENDENHALL R
PINE CREEK
PINE CREEK
PINE CREEK
PINE CREEK
MACKINTOSH
CARCROSS
CARCROSS
CARCROSS
CARCROSS
WHITEHORSE
WHITEHORSE
MACRAE
TAKHINI RIVER
ALA HWY M927
WHITEHORSE
ALA HWY M784
RANCHERIA
RANCHERIA
RANCHERIA
WATSON LAKE
WATSON LAKE
LIARD RIVER
LIARD RIVER
ALA HWY M667
YUKONENSIS
CHILKAT PASS»
CHILKAT PASS.
CHILKAT PASSe
MT GLAVE
CAPE DALHOUSIE
KIDLUIT BAY
TOKER PT
ANDERSON R
NICHOLSON PEN
ANDERSON R
ANDERSON R
REINDEER STAT
REINDEER STAT
ESKIMO LAKES
INUVIK
ESKIMO
ESKIMO
ESKIMO
ESKIMO
AKLAVIK
AKLAVIK
REINDEER
REINDEER
REINDEER
REINDEER STAT
REINDEER STAT
CANOE LAKE
MACK RE BR
INUVIK
COPPERMINE
O GRADY LAKE
FT RELIANCE
CROW FLATS
SAM LAKE
RAMPART HOUSE
RAMPART HOUSE
RAMPART HOUSE
DAWSON
DAWSON
DAWSON
DAWSON
SIXTYMI RD M21
CANOL RD M268
CANOL RD M259
LAKE LABERGE
ALA HWY M1028
LTL ATLIN L
KUSKOKWIM Re
SLEITMUT
MOUNTAIN VILGE
TOKSLA HWY M20
ALA HWY M1247
TAYLOR HWY M26
TAYLOR HWY M15
NUTZOTIN MTS
ALA HWY M1237
GEROGE LAKE
w FK LTL DELTA
w FK LTL DELTA
MANLEY HOT SPR
KANTISHNA R
FAIRBANKS
LAKES
LAKES
LAKES
LAKES
STAT
STAT
STAT
DATE
30
02
06
06
06
06
06
07
05
06
06
966
948
948
944
944
957
960
960
960
960
949
949
949
949
949
944
960
948
966
966
966
966
943
943
968
967
967
967
967
963
957
957
962
963
965
957
957
963
957
957
957
957
957
957
957
957
957
957
957
964
964
934
970
951
967
964
970
970
951
951
951
914
914
914
914
966
944
944
932
944
943
949
949
950
944
944
963
963
944
966
964
941
941
945
964
940
COLLECTOR NAMF
WMTITMNAOOAOQCCPFOUCCCCCOCCCEATIALZIGO
FPPrurrrroaonounCccezteaynPcrceczFeFEEZTAAZAAZEETEFTTEFTEAZAEDOOACAZEEDAGODAD
nM
GCrirRar naa eC crea 2s
TLS, SS. S.2 2 SSS SSS ee ee ee SS
RAAT 2 PhO NE soc ccc CC CC CC. 2s Pec Cee See
OU) SESS
ARGUS
RAUP
RAUP
RAUP
RAUP
SCHOFIELD
CALDER
CALDER
CALDER
CALDER
GILLETT
GILLETT
GILLETT
GILLETT
MITCHELL
PORSILD
CALDER
ANDERSON
ARGUS
ARGUS
ARGUS
ARGUS
RAUP
RAUP
WELSH
ARGUS
ARGUS
ARGUS
ARGUS
CODY
CODY
CODY
BARRY
PARMELEE
SCOTTER
SCOTTER
CODY
CODY
CODY
CODY
CODY
Cony
CODY
CODY
CODY
CODY
CopY
COopY
CODY
CODY
LARSEN
LARSEN
PORSILD
WELSH
FINDLAY
Copy
LARSEN
RIGBY
WELSH
LOAN
LOAN
LOAN
EASTWOOD
EASTWOOD
EASTWOOD
EASTWOOD
WELSH
PORSILD
PORSILD
BEAUCHAMP
RAUP
RAuUP
DRURY
DRURY
OAKENSON
ANDERSON
ANDERSON
NAVA
NAVA
RAUP
VIERECK
HARMS
PALYER
PALMER
SCAMMAN
VIERECK
ANDERSON
cOL NO
50R4
13026
13034
1182
11802
7587
24448
24930
24930
24448
4422
3197
3527
3227
13
9148
24465
10397
6017
6020
5042
5052
11040
11040
7483
HERR + NO
GWA
ALA 19913
ALA 19914
ALA 20039
GWA
CAN 269642
GWA
GWA
AO 32759
DAO 32758
DAO 32764
DAO 32761
DAO 32762
DAO 32763
DAO 32765
ISC 256265
NAO 32760
ISC 256266
GWA
GWA
GWA
GWA
ALA 20038
SASK 8652
GWA
GWA
GWA
5WA
SWA
.AO 32679
AO 32635
DAO 32636
DAO 32697
DAO 32676
GWA
DAO 32642
DAO = 55724
"AO = S711
NAO 32698
DAO 32693
DAO 32682
DAO 32691
DAO 32637
"AO 32692
NAO 32681
"AQ 32680
DAO 32685
DAO 32684
DAO 32678
D040 32686
GWA
GWA
GWA
ATF 1195
YAO 32689
DAO 32661
GWA
OTF 1194
OTF 1197
SWA
GWA
GWA
US 892177
US 2441253
A
US 892178
GWA
CAN 49581
TSC 256438
ALA 542
ALA 20059
ALA 20042
CAN
CAN
TSC 256416
1SC 256437
TSC 256473
ALA 23954
ALA 23952
ALA 20032
GWA
ALA 32607
ALA 5195
ISC 256450
GWA
FSLC 305
ISC 256444
203
Appendix B
TAxON SPC
i875 28
1a73 26
1875 28
1673 28
1673 28
1873 268
1875 28
1873 28
1673 28
1873 28
1873 28
1673 28
1673 28
1673 28
1873 28
1873 28
1673 28
1873 28
1673 28
1673 28
1873 28
1873 28
1873 28
1673 28
1873 28
1873 28
1873 28
1873 28
18673 28
1873 28
13873 28
1873 28
1673 28
1873 28
1673 28
1873 28
13873 28
1673 28
1673 28
1873 28
1673 28
1873 28
1673 28
1673 28
1673 28
1873 28
1673 28
1873 268
1873 238
1873 26
1673 28
1673 28
1673 28
1673 28
1673 28
1873 28
24 3 3 PLANIFOLIA SSPe
1673
1873
1673
1873
1873
1373
1873
1673
1873
1673
1673
1673
1673
1873
1673
1873
1675
1673
1673
1873
1673
1673
1673
1873
18673
1873
1673
1873
1673
1473
1873
204
28
28
28
28
28
28
28
28
28
28
268
28
28
28
28
268
28
28
28
28
28
28
28
28
28
268
28
26
28
28
28
Oo ol OG) OI OF OG OI OG) OI GI OI GI OI OI GI GF G8 OF Of OF GF OF OF OF GO OI OW OF OF OF Gi OF OW GI OI GI OF GI Gi OF Gy GF GW OF GI GI Gi OF OF OI GF OF GI
NVNAONNANNVNANNNANNNANNANNNNNANNANMNNANNNN NNN NNWNH NNN NH NW NHN /W PH W/W —/& f— PM LWA P— — PO f—O fo
wu OU Ul UFO OF OF OF OF OF OY GF OF OF OF OF OF GF Gi OF Gi OF OF GF OG! G8 GI OF Oi
<
OUUWoOWWoOoooUeoAaoawWwaoawoww Wo WW WwW & Oo
HYB PROV QUAD LAT LONG
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
02
v2
v2
v2
v2
v2
02
02
02
02
100
100
100
100
100
100
100
100
100
100
100
100
100
100
100
100
100
100
100
100
100
100
100
100
100
100
100
100
100
100
100
100
101
101
101
101
101
102
104
104
104
104
105
105
105
105
105
119
120
120
120
121
121
141
141
149
NOCOF ENN NNNNANNA WY
6452
6452
6452
6453
6453
6454
6452
6453
6452
6452‘
6452
6453
6452
6452
6452
6452
0452
6452
6451
6451
6451
6451
6451
6452
6448
6450
6451
6448
6448
6447
6447
6452
6454
6454
6454
6454
6454
6405
6531
6523
6527
6559
6512
6502
6502
6502
6502
6654
6644
6644
6644
0723
6723
6922
0922
7033
PULCHRA
5935
5935
5935
5935
5935
9935
5935
5936
5936
5935
5945
5938
5917
5917
5917
5956
5822
5822
5705
5705
9705
5659
5659
5659
70135
6935
6931
6931
6931
6827
6930
14750
14752
14752
14750
14750
14750
14755
14750
14750
14752
14752
14750
14750
14750
14750
14750
14752
14750
14743
14743
14743
147435
14743
14752
14756
14710
14658
14744
14744
14745
14745
14752
14625
14625
14625
14625
14625
14156
14513
14554
14526
14411
14805
14740
14740
14740
14740
14516
14334
14334
14354
14350
14350
15210
15210
15142
VARe
13629
13629
13629
13629
13629
13629
13629
13627
13627
13627
13230
13325
12951
12951
12951
12734
12248
12248
12235
12235
12235
12312
12312
12312
12940
13355
13348
13348
13348
13350
13347
LOCALITY
COLLEGE
SMITH L
SMITH L
COLLEGE
COLLEGE
GOLDSTREAM CR
ACE LAKE
COLLEGE
COLLEGE
SMITH L
SMITH L
COLLEGE
COLLEGE
COLLEGE
COLLEGE
COLLEGE
SMITH L
COLLEGE
FAIRBANKS
FAIRBANKS
FAIRBANKS
FAIRBANKS
FAIRBANKS
SMITH L
CHENA
FAIRBANKS
CHENA R
FAIRBANKS
FAIRBANKS
CHENA PUMP RD
CHENA PUMP RD
UNIV EXP FARM
CHENA RIVER
CHENA RIVER
CHENA RIVER
CHENA RIVER
CHENA RIVER
CHICKEN
STEES HWY M115
STEESE HWY M92
EAGLE CR CAMP
CIRCLE
WICKERSHAM DOM
FOX
FOX
FOX
FOX
FT YUKON
FISH HOOK CR
FISH HOOK CR
FISH HOOK CR
SMALL LAKE
SMALL LAKE
UMIAT
UMIAT
ATIGARU PT
PULCHRA
THREE GUARDSME
THREE GUARDSME
THREE GUARDSME
THREE GUARDSME
MT GLAVE
MT GLAVE
MT GLAVE
HAINES RD M160
HAINES RD MI60
THREEGUARDSMAN
ATLIN LAKE
SURPRISE LAKE
CASSIAR
CASSIAR
CASSIAR
ALA HWY M581
ALA HWY M258
ALA HWY M258
BEATTON RIVER
BEATTON RIVER
BEATTON RIVER
ALA HWY M128
ALA HWY M128
ALA HWY M128
CAPE DALHOUSIE
SUMMER ISLAND
RICHARDS IS
RICHARDS IS
RICHARDS IS
MACK R E BR
RICHARDS IS
DATE
957
956
956
957
957
966
956
957
957
956
956
957
966
965
965
965
965
965
933
933
933
933
931
956
955
966
966
965
965
964
964
964
966
966
966
966
966
951
945
957
966
965
965
965
964
964
964
949
957
957
957
957
957
964
964
947
967
967
967
967
967
967
967
956
956
967
954
960
956
956
956
960
960
960
943
943
43
960
960
960
963
963
957
957
957
957
966
COLLECTOR NAME
ATTKTNMNMNWHWECCK cK KKTCUMMMADOODOCFTOTTrFTrFTFKCMNNFFTFrFrre ce ce ceqaggagoaqgaeqgeqgaoqononenoneaea
MEEZEECEZEECCCIIICCCAAACCrAADDNDAGWADD
<€cc
PRCCOCOCCFrFVrKWVrVTFTrezgeTeELTeEZLEARELZEE
ZStEtttreprp>
NHNeveaDNANDWaADACrOCOCe’rT rely
SrPIreresSEtEztTeEZEtE
Caouncrr rae SVrree
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
HARMS
HARMS
HARMS
HARMS
“HARMS
PALMER
PALMER
PALMER
PALMER
PALMER
SMITH
SMITH
SUDA
VIERECK
VIERECK
VIERECK
VIERECK
VIERECK
VIERECK
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
SCAMMAN
SCAMMAN
SHETLER
TRENT
VIERECK
HARMS
HARMS
HARMS
HARMS
HARMS
COOK
SHETLER
SHETLER
SHETLER
SHETLER
SHETLER
VIERECK
VIERECK
BLACK
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
TAYLOR
TAYLOR
VIERECK
AITKEN
CALDER
TAYLOR
TAYLOR
TAYLOR
CALDER
CALDER
CALDER
RAUP
RAUP
Raye
CALDER
CALDER
CALDER
Copy
CopY
copy
CODY
copy
Copy
SCOTTER
COL NO
1198
301
3n1
1007
1006
5090
302
106
1108
301
320
1007
5106
3418
3377
3376
3346
BYuy
194
191
155
177
14
HE83 + NO
ALA 6715
ALA 4759
ALA 22459
ALA 6727
ALA 6725
GWA
GWA
BWA
GWA
GWA
BWA
GWA
GWA
ALA 32588
ALA 32659
ALA 32658
ALA 32652
ALA 32633
ALA 5265
ALA 5264
ALA 5266
ALA 5268
ALA 5972
ALA 10741
ALA 10744
GWA
GWA
FSLC
FSLC
FSLC
FSLC
FSLC 283
SWA
GWA
SWA
GWA
GWA
GWA
GWA
ALA 3789
ALA 32693
FSLC
ALA 32589
ALA 32587
ALA 32623
ALA 32622
ALA 32568
TSC 256418
ALA 3880
ALA 3906
ALA 3966
ALA 4169
ALA 4216
GWA
GWA
TSC 256428
GWA
GWA
GWA
GWA
GWA
GWA
5WA
NAO 32746
YAO 32749
GWA
SAO 32771
NAO 32743
NAO 32750
NAO 32744
DAO 32748
AO 32772
GWA
AO 32767
ALA 20027
ALA 20028
ALA 20037
GWA
AO 32774
NAO 32776
AO 32688
240 32683
AO 32669
940 32670
"AO 32667
"AD 32668
GWA
Specimens Cited
TaxON SPC S V HYB PROV QUAD LAT LONG LOCALITY DATE
1875 26 3 3 07 133 6826 13255 TUKTOYAKTUK 13 08
18753 28 3 3 07 133 6826 13255 TUKTOYAKTUK 13 08
1873 26 3 3 07 134 6947 13102 MCKINLEY BAY 15 08
1673 28 3 3 07 134 6925 12816 ANDERSON R 13 07
1873 28 3 3 07 134 6925 12816 ANDERSON R 23 07
14673 28 3 3 07 152 6857 13234 ESKIMO LAKE 06 08
1673 28 33 O07 152 6813 13554 CANOE LAKE 16 07
1875 28 3 3 O07 152 6813 13554 CANOE LAKE 16 07
1873 28 3 5 O7 152 6813 13554 CANOE LAKE 16 07
18735 28 3 3 O7 152 6813 13554 CANOE LAKE 18 07
18753 286 3 3 O07 152 6842 13408 REINDEER STN 27 06
1875 28 3 3 07 152 6842 13408 REINDEER STN 10 06
1873 28.3 3 07 152 6842 13408 REINDEER STN 12 06
1873 28 3 3 07 152 6842 13408 REINDEER STN 10 06
1873 28 3 3 07 152 6842 13408 REINDEER STN 13 07
16735 28 3 3 07 152 6842 13408 REINDEER STN 10 06
1875 28 33 07 152 6842 13406 REINDEER STN 16 06
167535 26 3 3 07 152 6820 13337 INUVIK 13 06
1875 2833 O07 152 6813 13547 CANOE LAKE 26 06
167535 28 3 3 07 152 6839 13406 REINDEER STN 03 08
1875 28 3 3 07 152 6848 13424 REINDEER STN 31 07
18735 26 3 3 07 153 6812 13115 HYNDMAN LAKE 07 07
1673 28 3 3 07 181 6750 11510 COPPERMINE 28 06
18735 26 3 3 07 181 6750 11510 COPPERMINE 24 06
187535 28 3 3 07 181 6750 11510 COPPERMINE 28 06
1675 28 3 3 07 181 6750 11510 COPPERMINE 28 06
1873 28 3 3 07 218 660% 10828 BATHURST INLET 24 07
1875 28 3 3 O07 283 6405 11115 MATTHEWS LAKE 30 06
1875 28 3 3 07 283 6405 11115 MATTHEWS LAKE 30 06
1873 28.3.5 U7 283 6405 11115 MATTHEWS LAKE 30 06
1673 28 3 3 07 283 6405 11115 MATTHEWS LAKE 21 06
1875 28 3 3 O07 285 6405 11115 MATTHEWS LAKE 30 06
1873 28 3 3 07 283 6405 11115 MATTHEwS LAKE 18 06
1875 26 3 3 07 283 6405 11115 MATTHEWS LAKE 30 06
1873 28 3 3 07 284 6445 10810 MUSKOX LAKE 07 08
1875 28 33 07 284 6405 10830 AYLMER LAKE 05 08
1873 28 3 3 07 285 6404 10759 THANAKOIE NAR 07 O08
1675 26 3 3 07 306 06310 13008 MACMILLAN PASS 31 08
1873 28 3 3 07 307 6331 12840 JUNE LAKE 02 08
1873 28 3 3 07 338 6202 12810 FLAT RIVER 08 08
1873 28 3 3 07 338 6202 12810 FLAT RIVER 08 08
1873 28 3 3 07 338 63035 12855 0 GRADY LAKE 28 07
18735 28 3 3 07 338 6300 12902 0 GRADY LAKE 25 07
1875 28 3 3 07 338 6202 12810 FLAT RIVER 08 08
1873 28 3 3 07 339 6242 12640 LTL DAL LAKE 06 08
1873 28 3 3 O07 239 6205 12735 BRINTNELL L 26 06
1873 28 3 3 07 339 6205 12735 BRINTNELL L 21 06
1873 26 3 3 07 367 6142 12710 HOLE IN WALL L 10 08
1873 26 3 3 07 367 6142 12710 HOLE IN WALL L 10 08
1873 28 3 3 12 4 6856 13712 SHINGLE POINT 15 07
1873 28 3 3 12 4 6850 13845 TROUT LAKE {7
1873 28 3 3 12 4 6825 13800 SAM LAKE 30” "6
1873 28 3 3 12 4 ©6840 13907 BABBAGE CREEK if
1875 26 3 3 12 4 6857 13705 BLOW RIVER 06 08
1673 28 3 3 12 5 ©6746 14015 OLD CROW MT 20 07
1873 28 3 3 12 15 6458 13825 OGILVIE MT 27 06
1675 28 3 3 12 15 6409 13953 SIXTYML RD M12 26 06
1673 28 3 3 12 21 ©6350 13731 BARLOW 05 06
1675 28 3 3 12 22 ©6357 13510 KENO HILL 06 08
1873 283 3 12 30 6249 13043 CANOL RD M245 21 08
1873 28 3 3 12 35 6140 13302 CANAL RD M105 11 06
1873 28 3 3 i2 49 ©6047 13735 BEAR CREEK 18 06
1873 28 3 3 12 4Q 6047 13735 BEAR CREEK 18 06
1875 28 3 3 12 43 ©6005 13035 RANCHERIA 22 08
1675 28 3 3 12 43 ©6005 13035 RANCHERIA 22 08
1875 28 3 3 14 29 5505 16202 BELKOFSKY
1873 28 3 3 14 32 5657 15410 AKHIOK 26 05
1873 28 3 3 14 34 5735 15400 UYAK BAY 26 05
1875 28 3 3 14 34 5713 15314 SHEEP ISLAND 06
1873 28 3 3 14 35 5732 15359 LARSON BAY
1673 28 3 3 14 35 5705 15425 OLGA BAY 18 05
1873 28 3 3 14 36 9732 15725 UGASHIK 25 07
1873 26 33 14 38 5635 16935 ST GEORGE IS 08 08
1873 28 3 3 14 39 5834 16146 CAPE PEIRCE ee ot
1675 28 33 14 39 5834 160146 CAPE PEIRCE 8 6
1873 263 3 14 39 5840 16102 HAGEMEISTER IS 22 06
18735 28 3 3 14 39 5840 16102 HAGEMEISTER IS 22 06
1873 28 3 3 14 41 5811 15600 CONTACT CREEK 06 09
1875 28 3 3 14 41 5844 15747 KVICHAK BAY 05 06
1873 28 3 3 14 41 5844 15747 KVICHAK BAY 05 06
1875 28 3 3 14 41 5844 15747 KVICHAK BAY 05 06
18753 28 3 3 14 41 5844 15747 KVICHAK BAY 08 06
18735 26.3 3 14 41 5842 15640 NAKNEK 25 06
1673 28 3 3 14 41 5845 15637 NAKNEK 03 07
1875 28 3 3 14 41 5841 15639 KING SALMON 06 07
1873 28 3 3 14 41 5843 15701 NAKNEK 03 07
1873 28 3 3 14 42 5835 15550 DUMPLING MT TR 29 05
1873 28 3 3 14 43 5804 15304 PORT VITA 31 05
1873 28 3 3 14 43 5805 15305 IRON CREEK 12 10
1873 28 3 3 14 50 5947 15150 ANCHOR POINT 13 06
1875 28 3 3 14 51 5920 15555 IGIUGIS 21 07
965
965
957
965
965
957
963
963
963
963
957
957
EST /
957
957
957
964
964
964
965
965
966
951
951
O51
951
949
953
553
953
953
953
953
953
953
965
965
944
967
967
967
967
967
967
967
939
939
967
967
963
970
970
970
964
964
966
966
960
949
944
944
944
944
966
966
933
940
93A
963
933
938
967
945
970
970
966
966
954
951
951
951
951
949
949
952
952
967
945
945
967
965
COLLECTOR NAME
<FZEZFOZZAIMMMMCADTTFcecoLTOACMoOOO TIP PSPcewMAVVIMMMHCAZLTLILZAZTZFASAZHPCCCCCccecececc#tsFsEROOOCCCZEAZEZFTFAEZEFEEZLENDODEOOD
AHHH EZTEVPrPSPCCcCCCCcCcKeccCecacucezeczE
an
)
riz VUTEECCSseEayA evra Trreekac te sO £oOGQocaG tka SOaDDI|Ao
PRBPELE IEEE
reac
SCOTTER
SCOTTER
copy
SCOTTER
SCOTTER
copy
CopY
CopY
CopY
Copy
CODY
copy
CODY
CODY
Copy
Copy
LARSEN
LARSEN
LARSEN
SCOTTER
SCOTTER
SCOTTER
FINDLAY
FINDLAY
FINOLAY
FINOLAY
WOODRUFF
CHILLCOTT
CHILLCOTT
CHILLCOTT
CHILLCOTT
CHILLCOTT
CHILLCOTT
CHILLCOTT
CHILLCoTT
LARSEN
LARSEN
PORSILD
CODY
‘CODY
CODY
CODY
CODY
SPICER
CODY
RAUP
RAUP
CODY
SPICER
PARMELEE
WELSH
WELSH
WELSH
YOUNGMAN
YOUNGMAN
PORSILD
WELSH
CALDER
GILLETT
PORSILD
PORSILD
RAUP
RAUP
ARGUS
ARGUS
GEIST
LOOFF
ANDERSON
GORDON
GEIST
LOOFF
STREUBEL
ANDERSON
DICK
DICK
PEGAU
PEGAU
CAHALANE
MULLER
MULLER
MULLER
MULLER
RAUP
RAUP
SCHOFIELD
SCHOFIELD
ESTABRO0K
EYERDAM
EYERDAM
VIERECK
HARMS
COL NO
6996
6996
10928
6931
6956
10518
12807
12813
12818
129074
9691
9570
9590
9577
10025
9567
7024
7026
7028
6973
6968
1n102
42
HERB
GWA
NAO
NAO
NAO
NAO
"AO
NAO
DAO
NAO
YAO
‘AO
NAO
AO
DAO
NAO
MAO
GWA
GWA
GWA
GWA
GWA
SWA
NAD
DAO
AO
DAO
NAO
40
AO
DAO
DAO
NAO
DAO
AO
NAO
GWA
GWA
rc
NAO
NAO
DAO
DAO
AO
NAO
NAO
ALA
ALA
DAD
DAO
NAO
RRY
OTF
OTF
CAN
CAN
CAN
GWA
GWA
NAO
CAN
CAN
GWA
ALA
GWA
GWA
ALA
+ NO
32632
32631
32633
32634
32645
32647
32649
32646
32651
32652
32653
32648
32589
256439
32657
32658
32659
32662
32664
32656
32655
14763
14764
32660
32666
32756
91146
1193
1196
32751
20041
29388
US 2441271
TSC
WIS
ALA
GWA
ALA
Tsc
GWA
GWA
SWA
ALA
Tsc
Tsc
tsc
TSsc
rsc
tsc
Tsc
BWA
ALA
ALA
tsc
Tsc
GWA
ALA
256468
29398
34727
256504
32702
219360
224500
224495
224497
224496
219353
219351
17432
34694
256432
256431
32560
205
Appendix B
TAXON SPC
1873 28
1873 28
1873 28
1873 28
1873 28
1873 28
1873 28
1873 28
13873 28
1873 28
1873 28
1873 28
13873 28
1873 28
1673 28
1873 28
1873 28
1873 28
1873 28
1873 28
1873 28
1873 28
1873 28
1873 28
1873 238
13873 28
1673 28
1873 28
1873 28
1673 28
1873 28
1873 28
18735 28
1873 268
18735 28
1873 28
1673 28
1873 28
1873 28
1873 28
1873 28
1873 28
1873 28
1873 28
1673 28
1673 28
1673 28
1873 28
1673 28
1473 28
1873 28
1873 28
1873 28
1673 28
1673 28
1873 28
1673 28
1673 28
1673 28
1873 28
18673 28
1873 26
1873 28
1673 28
1873 28
1673 28
1673 26
1673 28
14673 28
1873 28
1873 28
1873 28
1673 26
1673 28
1673 28
1473 28
18673 28
1873 28
15673 28
1873 28
1473 28
1673 28
1873 28
1873 28
1673 28
1873 28
1673 28
1673 28
1673 28
1673 28
1873 28
206
VuUWduUWwuuww Goo Uw ou UO WOW OI OO OO OF Ol OI OF OF OI OF OF GIO GI OI GF OF GI OF GF OF OF OF OF OF OF OF OF OF GO! OF GI OOF OF Gt OF OF GI OF OF OG! GI OF OF OF GO) OF GI OI GF OF Gt GI OF Gi GI GI GF GF GF Gl GI Oi OF GI OF Gt GF Oi
VUUUWUwwoUWwWoUWwwwwwuw wou wo W & & GO OF & OO Oi OF OO OI OI O OF OI OO) OF OO GG OG OF OF Ov OF Ot Gi GI GI Gi Gi OF Gi Gi GF Gi Gi Gi Gt Gi Gi Gi GF Gi Gt Gi Gi GO) Gi Gi Gt Gi Gi GI Oi Gi Oi Gi GH OF Gi
S
V HYB PROV QUAD
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
56
57
57
S7
62
62
63
63
68
68
69
69
69
69
69
69
69
69
69
69
69
69
69
71
71
71
72
Lie
73
73
73
73
Ue)
1
73
75
79
79
80
80
81
81
81
81
81
81
81
81
81
81
LAT
6024
6023
6015
6015
6033
6003
6029
6029
6108
6148
6135
6115
6115
6115
6115
6140
6140
6140
6140
6140
6116
6158
6116
6152
6152
6152
6139
6137
6131
6132
6131
6137
6128
6137
6131
6200
6259
6259
6247
6257
6219
6229
6240
6240
6225
6225
6225
6225
o225
6240
6240
6225
6206
6322
6315
6315
6330
6347
6345
6344
6344
6344
6355
0335
6332
6325
6325
6332
6335
6325
0325
6325
6325
6325
6325
6327
6327
6327
6301
6315
6310
6310
6352
6352
6329
6335
6335
6302
6302
6302
6302
LONG
17242
16611
16614
16614
15116
15140
14922
14922
14544
14512
14930
14920
14920
14920
14920
14900
14900
14900
14900
14900
14941
14708
14941
15433
15433
15433
15717
15930
15937
15927
15937
15930
15939
15930
16021
16600
15604
15604
15544
15536
15006
15128
15230
152350
15125
15125
15125
15125
15125
15230
15230
14451
14542
14232
14540
14540
14535
14430
14433
14855
14855
14855
14725
14935
15058
15030
15030
15058
14935
15030
15030
15050
15035
15035
15035
15045
15045
15045
15532
15517
15631
15631
16047
16047
16202
16230
16230
16333
16333
16333
16333
LOCALITY
ST MATTHEWS IS 16
MEKORYUK RIVER 17
MEKORY JK 22
MEKORY JK 17
KENAI 31
NINILCHIK 13
MOOSE PASS 01
MOOSE PASS 17
THOMPSON PASS 05
RICH HWY M90. OL
PALMER 06
CHUGACH MTS 11
CHUGACH MTS 11
CHUGACH MTS 11
CHUGACH MTS 11
MATANUSKA VY
MATANUSKA VY 24
MATANUSKA VY
MATANUSKA VY
MATANUSKA VY 24
FT RICHARDSON 12
GLENN HWY M134 28
FT RICHARDSON 12
HEAD OF BIG Re 4
HEAD OF BIG Re 2
HEAD OF BIG Re 3
SLEITMUT 11
ANIAK REGION 18
ANIAK REGION 28
ANIAK REGION 20
ANIAK REGION 28
ANIAK REGION 18
ANIAK REGION 28
ANIAK REGION 18
KALSKOS R 18
KOKECHIK BAY 03
TAKOTNA 27
TAKOTNA 27
MCGRATH 22
MCGRATH 13
TALKEETNA 11
CHELATNA LAKE 14
KUSKOKWIM R 01
KUSKOKWIM R 29
CHELATNA LAKE 21
CHELATNA LAKE 21
CHELATNA LAKE 21
CHELATNA LAKE 21
CHELATNA LAKE 21
KUSKOKWIM R 01
KUSKOKWIM R 29
TOKSLA HWY M20 24
GLENNALLEN 10
TAYLOR HWY MS 21
MCCALLUM CR 19
MCCALLUM CR 19
RAINBOw MT 09
GEORGE LAKE 10
ALA HWY M1380 10
MCKINLEY PARK 30
MCKINLEY PARK 30
MCKINLEY PARK
DRY CR 23
IGLOO CREEK 28
KANTISHNA 25
MULDROW GL 27
MULOROw GL 27
KANTISANA 25
IGLOO CREEK 28
MULDROW GL 27
MULDROw GL 27
WONDER LAKE 27
MCKINLEY R 21
MCKINLEY R 20
MCKINLEY R 22
MCKINLEY R 20
MCKINLEY R 05
MCKINLEY R 20
TAKONTA Re 23
NIXON FORK 10
OPHIR 26
OPHIR 26
UNALAKLEET 12
UNALAKLEET 12
ST MICHAEL 19
STUART ISLAND 29
STUART ISLAND 29
KOTLIK 07
KOTLIK 07
KOTLIK 07
KOTLIK 07
DATE
07
07
07
06
05
06
06
06
07
08
08
07
07
07
07
6
oooo
DHDADDDDNNDONDAND
938
965
965
949
951
967
944
941
935
967
947
948
948
948
948
940
940
940
940
940
965
965
965
950
950
950
949
949
949
949
949
949
949
949
930
960
941
941
949
949
944
961
961
961
956
956
956
956
956
961
961
944
967
963
957
957
965
964
944
939
939
940
962
956
956
956
956
956
956
956
956
958
958
958
958
958
958
958
950
950
940
940
938
938
938
931
931
931
931
931
931
COLLECTOR NAME
J
ic]
R
D
J
Le
J
J
J
L
3
E
E
E
L
L
L
E
iE
=)
S
iS
W
W
W
W
W
W
W
W
W
W
W
W
N
J
J
W
W
J
E
L
L
L
L
iC
L
L
L
L
J
L
J
G
G
V
V
H
A
A
L
L
G
G
G
G
G
G
G
G
L
L
L
L
L
L
L
W
W
E
E
J
J
J
Cc
c
Cc
Cc
Cc
C
P ANDERSON
BOS
PEGAU
SPENCER
CALDER
VIERECK
ANDERSON
ANDERSON
ANDERSON
VIERECK
DUTILLY
LEPAGE
LEPAGE
LEPAGE
LEPAGE
PALMER
PALMER
PALMER
PALMER
PALMER
WELSH
WELSH
WELSH
DRURY
DRURY
DRURY
DRURY
DRURY
DRURY
DRURY
DRURY
DRURY
DRURY
DRURY
MILLER
JONES
ANDERSON
ANDERSON
DRURY
DRURY
ANDERSON
LITTLE»
VIERECK
VIERECK
VIERECK
VIERECK
VIERECK
VIERECK
VIERECK
VIERECK
VIERECK
ANDERSON
VIERECK
NAVA
ARGUS
ARGUS
HARMS
HARMS
RAUP
NELSON
NELSON
PALMER
VIERECK
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
VIERECK
VIERECK
VIERECK
VIERECK
VIERECK
VIERECK
VIERECK
DRURY
DRURY
SCAMMAN
SCAMMAN
ANDERSON
ANDERSON
ANDERSON
ROUSE
ROUSE
ROUSE
ROUSE
ROUSE
ROUSE
>PUVUVVS Pr
2>UPr rE OPPYPrPPrFFUIEEVVMA VDITIOEOCETLEECLrCIrr rT ceccace
=
i ae
fr
TLoOPrPePrappe rvresteEe ttt e>
a Se See as ole on alo TB Ti |
COL NO
3947
SeNe
wi?
8
5004
8258
6471
6815
1891
8453
21963
23446
23446
23446
23462
395
121
395
394
121
4590
4462
4590
3977
3954
3911
1934
1463
1642
1577
1643
1440
1695
1505
260C
50
7445
THES
2172
1366
7711
19425
5234
5178
1021
1036
1024
1001
1036
5234
5178
8713
Be11
96
1034
1034
3570
3165
12758
35R8
358
422
5790
692
652
666
665
652
692
666
665
30°86
3231
3039
3063
3041
3156
3040
4662
3624
1830
1830
32678
3267
3447
22
22
35
34
35
34
HERB + NO
1SC 255907
ALA 32287
GWA
1SC 256417
GWA
6WA
ISC 256446
TSC 256447
ISC 256454
GWA
940 32754
DAO 32755
ISC
TSC 256470
TSC 256469
ALA 5961
ALA 5168
ALA 5962
ALA 5960
US 2441270
ALA 29907
TSC 246047
TSC 246931
mAN
CAN
CAN
CAN
CAN
CAN
CAN
CAN
CAN
CAN
CAN
ALA 2020
ALA 24422
ALA 545
ISC 256449
CAN
CAN
TSC 256448
GWA
GWA
GWA
GWA
GWA
ALA 3497
ALA 11612
ALA 11615
ESLC
FSLC
1SC 256007
GWA
ALA 23953
GWA
ALA 6732
ALA 32655
ALA 32612
ALA 20033
ALA 544
TSC 256451
ALA 5633
5WA
BWA
SWA
GWA
SWA
ALA 4559
ALA 4592
ALA 4905
ALA 4906
6WA
GWA
6WA
GWA
5WA
5WA
BWA
CAN
CAN
GWA
ALA 86393
TSC 256435
TSC 256466
1SC 256436
ALA 2055
ALA 25447
ALA 25429
ALA 25433
ALA 22040
ALA 2040
Specimens Cited
TaxON SPC S V HYB PROV QUAD LAT LONG LOCALITY
1673 26 3 3 14 92 6300 16250 PASTOLIK
16735 28 3 3 14 92 6300 16250 PASTOLIK
1875 28 3 3 14 92 6259 16318 PASTOLIK
1873 28 3 3 14 92 6259 16318 PASTOLIK
1873 28 3 3 14 93 ©0347 17145 GAMBELL
1675 28 3 3 14 93 6342 17029 SAVOONGA
1673 28 3 3 14 93 6330 17030 ST LAWRENCE IS
1673 28 3 3 14 93 6330 17030 ST LAWRENCE IS
1873 28 3 3 14 93 6330 17045 ST LAWRENCE IS
1873 28 3 3 14 93 6330 17030 ST LAWRENCE IS
18735 28 3 3 14 93 ©6330 17030 ST LAWRENCE IS
1873 28 3 3 14 93 6320 17136 BOXER BAY
1873 28 3 3 14 93 6320 17136 BOXER BAY
1873 28 3 3 14 93 6334 17053 KANGEE CAMP
1875 28 3 3 14 94 6430 16530 NOME
1673 28 3 3 14 94 6430 16530 NOME
1873 28 3 3 14 94 6435 16530 NOME
1673 28 3 3 14 94 6434 16533 NOME
1873 28 3 3 14 94 6435 16530 NOME
1673 28 3 3 14 95 6433 16302 GOLOVIN
1873 28 3 3 14 95 6433 16302 GOLOVIN
1873 26 3 3 14 96 6402 16055 EGAVIK
1873 28 3 3 14 96 ©6402 16055 EGAVIK
1873 28 3 3 14 96 6402 16055 EGAVIK
1873 28 3 3 14 96 6402 16055 EGAVIK
1873 2833 14 96 6402 16055 EGAVIK
1873 es 3 3 14 96 6402 16055 EGAVIK
18735 28633 14 100 6451 14743 FAIRBANKS
18753 26 3 3 14 101 6345 14554 DONNELLY DOME
1873 28 3 3 144 101 6345 14554 DONNELLY DOME
1875 28 3 3 14 101 6345 14554 DONNELLY DOME
1875 28 3 3 14 101 6345 14554 DONNELLY DOME
1875 2833 14 101 6345 14554 DONNELLY DOME
1875 28 3 3 14° #101 6433 14455 INDIAN CREEK
1873 28 3 3 14 104 6529 14526 EAGLE SUMMIT
1873 2833 14 104 6547 14656 VICTORIA MT
1873 26 3 3 14% #104 ©6531 14513 MILLER HOUSE
1873 2833 14° 104 6531 14513 MILLER HOUSE
1873 28 3 3 14 104 6522 14556 STEES HWY MB9
1873 28 3 3 14 104 6522 14556 STEES HwY M89
1873 28 3 3 14 104 6522 14556 TWELVE MILE CR
1873 28 3 3 14 #104 6523 14557 TWELVEMI SUMIT
1873 28 3 3 14 104 6522 14556 TWELVE MILE CR
1873 26 33 14° #104 6527 14526 EAGLE CREEK
1u735 26 3 3 14 104 6527 14526 EAGLE CREEK
1573 28 3 3 14 105 6512 14805 WICKERSHAM DOM
1873 28 3 5 14 105 6512 14805 wICKERSHAM DOM
1873 28 3 3 14 #105 6512 14805 WICKERSHAM DOM
1873 28 3 3 14 105 6512 14805 wICKERSHAM DOM
1873 28 3 3 14 #108 6448 14545 DONNELLY DOME
1673 28 3 3 14 110 6536 16313 IMURUK LAKE
1873 28 3 3 14 110 6536 16313 IMURUK LAKE
1873 28 3 3 14 111 6537 16805 WALES
18735 28 3 3 1% 111 6516 16622 TELLER
1875 28 3 3 14 111 6516 16622 TELLER
187535 28 3 3 14 111 6458 16804 KING ISLAND
1875 28 33 14% 111 6509 16553 IMURUK BASIN
13873 28 3 3 14 113 ©6654 16235 KOTZEBUE
15673 28 3 3 14 113 6655 16231 KOTZEBUE
1873 28 3 3 14 113 6655 16240 KOTZEBUE
1873 28 3 3 14 114 6658 16026 KIANA
1875 28 3 3 14 114 6616 16120 ELEPHANT POINT
1675 28 3 3 14 114 6616 16120 ELEPHANT POINT
1575 28 3 3 14 114 6616 16120 ELEPHANT POINT
18753 28 3 3 14 114 6616 16120 ELEPHANT POINT
1875 28 3 3 14 114 6616 16120 ELEPHANT POINT
1875 28 3 3 14 114 6616 16120 ELEPHANT POINT
1873 28 3 3 14 114 6©616 16120 ELEPHANT POINT
1875 28 3 3 14 114 6616 16120 ELEPHANT POINT
18673 2833 14° #114 6616 16120 ELEPHANT POINT
1873 28 3 3 14 114 6616 16120 ELEPHANT POINT
18735 26 3 3 14 114 6616 16120 ELEPHANT POINT
18735 28 3 3 14 #114 6616 16120 ELEPHANT POINT
1875 28 3 3 14 114 6616 16120 ELEPHANT POINT
1673 28 3 3 14 114 6617 16153 CHORIS PEN
18753 28 3 3 14 114 6617 16153 CHORIS PEN
1875 28 3 3 14 #114 6617 16153 CHORIS PEN
1873 28 33 14 120 ©6644 14334 BIG LAKE
18735 28 3 3 14 121 ©710 14210 PORCUPINE R
1873 28 3 3 14 121 ©6710 14210 PORCUPINE R
1675 28 3 3 14 125 6757 15312 EASTER CREEK
1873 28 3 3 14 #126 6743 15609 ISIAK LAKE
1873 28 3 3 14 128 6744 16432 KIVALINA
1673 28 3 3 14 128 6744 16432 KIVALINA
1875 28 3 3 14 128 6744 16432 KIVALINA
1873 28 33 14% 128 6744 16432 KIVALINA
1873 28 3 3 14 128 6744 16432 KIVALINA
1873 28 3 3 14 128 6744 16432 KIVALINA
1873 28 3 3 14 #128 6744 16432 KIVALINA
1673 28 3 3 14 128 6744 16432 KIVALINA
1673 28 3 3 14 128 6744 16432 KIVALINA
931
931
931
931
938
938A
933
933
933
933
933
933
933
933
938
964
966
968
966
938
924
931
931
931
931
931
931
933
965
964
965
964
964
956
965
953
940
945
957
957
957
957
957
966
965
964
965
965
965
967
947
947
938
938
959
968
938
966
966
931
931
931
931
931
931
931
931
931
931
931
931
931
931
931
931
957
957
957
965
960
938
960
960
960
960
960
960
960
960
COLLECTOR NAVE
rPrrrrrrreve CC YNNANANNAANQANANAAANNANNNNIONACAOSQcCCCCCTFTKeKeeeCCVHVNNNUMMOK Pee KK KFANNNNANFCAADABANCOVOCCOCOCOOVOCOOC CANN
VEEZLEEZ=ZZEVGIrICICC
Cece re ee ote be Le Wey
N
VVUvnDseCre rr 2
a face | JR So es a i a cage oe aga: aie’ ge os i rl la coe ee Ul aa Se | =
re]
ROUSE
ROUSE
ROUSE
ROUSE
ANDERSON
ANDERSON
GEIST
GEIST
GEIST
GEIST
GEIST
GEIST
GEIST
GEIST
ANDERSON
HELLER
PEGAU
PEGAU
PEGAU
ANDERSON
RYNNING
ROUSE
ROUSE
ROUSE
ROUSE
ROUSE
ROUSE
PALMER
HARMS
HARMS
HARMS
HARMS
HARMS
JOHNSON
HARMS
KLEIN
SCAMMAN
SCAMMAN
SHETLER
SHETLER
SHETLER
SHETLER
SHETLER
TRENT
TRENT
HARMS
HARMS
HARMS
HARMS
VIERECK
SIEH
SIEH
ANDERSON
ANDERSON
ANDERSON
HARBO
PEGAU
ANDERSON
ARGUS
WELSH
CLARK
ROUSE
ROUSE
ROUSE
ROUSE
ROUSE
ROUSE
ROUSE
ROUSE
ROUSE
ROUSE
ROUSE
ROUSE
ROUSE
ROUSE
ROUSE
ROUSE
SHETLER
BUCKLEY
BUCKLEY
STAENDER
LENT
ANDERSON
BUCKNELL
BUCKNELL
BUCKNELL
BUCKNELL
BUCKNELL
BUCKNELL
BUCKNELL
BUCKNELL
cOL NO
210
3546
2866
3538
2840
28f2
46
3858
SeNe
2046
3495
214AF
161AF
BT7AF
L20AF
59AF
466
5265
3878B
3473C
3663
34U73A
R329
Tis,
7
4936
3575
3523A
SeNe
27468
4708
5977
5736
SeNe
BOOANNF FFE INURK KNN UWI
337AF
nN
com )
ul
203
—oO
Fn
4626
> et Oe et ee
Frrenwoou
HERB
ALA
ALA
ALA
ALA
TSC
TSC
ALA
ALA
ALA
ALA
ALA
ALA
ALA
ALA
1SC
ALA
GWA
GWA
ALA
TSC
ALA
ALA
ALA
ALA
ALA
ALA
TSC
ALA
ALA
ALA
ALA
ALA
ALA
ALA
ALA
ALA
GWA
GWA
ALA
ALA
ALA
ALA
ALA
SLA
ALA
ALA
ALA
ALA
ALA
GWA
ISC
TSC
SC
TSC
TSC
ALA
GWA
1SC
SWA
GWA
ALA
ALA
ALA
ALA
ALA
ALA
ALA
ALA
ALA
ALA
ALA
ALA
ALA
ALA
ALA
ALA
ALA
ALA
GWA
ALA
ALA
ALA
SC
ALA
ALA
ALA
ALA
ALA
ALA
ALA
ALA
+ NO
2056
25446
2053
25448
256464
256463
29392
29393
29735
29391
29739
29425
29422
29389
256465
27072
32706
256467
2032
25431
2064
25427
2061
25444
256445
5982
32637
32602
32649
32644
32603
7064
32653
23092
3800
3774
3735
3758
3715
32695
30668
33341
32627
32582
32628
256433
256434
256459
256456
256462
24826
256457
10190
2069
25424
2072
25423
2071
25422
2070
2073
25436
25425
25428
2067
2068
2066
25432
2065
3882
5036
29337
256460
26559
26591
26557
26575
26593
26558
26592
26574
207
Appendix B
TAXON SPC
1873 28
13873 28
1873 28
1873 28
1873 28
1873 28
1873 28
1873 28
1873 28
1873 28
13873 28
1873 28
1873 28
1873 28
1873 28
1873 28
1873 28
1873 28
1873 28
1873 28
1873 28
1873 28
1473 28
1873 28
1873 28
1873 28
1473 28
1873 28
1873 28
1873 28
1873 28
1873 28
1873 28
1873 28
1673 28
1673 28
1873 28
1873 28
1873 268
1873 28
1873 28
1873 26
1873 28
14673 28
1873 28
1875 28
1875 28
1873 28
1873 28
1873 28
1a73 28
1873 28
1673 28
1873 28
1873 28
1673 28
1873 28
1873 28
1673 28
1873 28
1673 28
1673 28
1873 28
1573 28
1873 28
1673 28
1873 28
1673 28
1673 28
1873 268
1873 28
1873 28
1873 28
18673 28
1873 28
1873 28
1873 28
1873 28
1873 26
1673 28
1873 28
1873 28
1673 28
1673 28
1873 26
1673 26
1873 28
1873 268
1673 28
1873 28
1873 28
208
S
VoOO Ww OW OG OOO GI OO OI OF OF OF OF OF Gi OI OF OF OF GO) OF GO! OY OF OH OG! OF GI OI OF OF GI GI GO! GI Oi OI Ov OF Gi GF OF OF OF OF OF OF OF OF GI GI OF OF OF Gl OF GI OF OI OF Gl Of OF OF Gl GF Of OF Gi Of OF GF GH OF OF Gi OF GF Gi Gi OF OF OW
Gl Ot GG Gt Ge Gr GF GH OF! OOF OF GI GO) OF OF OF GI GI GH OG! GF OF GO OF OG) GIO) OF OO) OF OF GO Ge OF OW Gt GO! Gi Gi Ol GF Gi GI Gi OF OF Gr Gi OF OF OG) OF Gl GO Gi Gi GI Gi GF GI GI Or Gi GF OF Gi Gi GI OF Gl Gi GI Gi GF OF OF Of Ol Gi Or GI OF GH OF Gi GF GF Gi
V HYB PROV QUAD
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
128
128
128
129
129
129
129
129
129
129
129
129
129
129
129
129
129
129
129
129
129
129
134
134
134
138
141
141
141
141
141
141
141
142
142
142
142
142
142
144
144
145
145
145
145
145
145
146
146
146
147
147
147
147
147
147
148
151
151
4151
152
152
152
152
153
153
153
153
153
153
153
153
153
153
153
153
153
153
153
153
153
153
153
153
153
153
153
153
153
153
153
LAT
6744
6744
6744
6821
6853
6806
6806
6806
6806
6806
6806
6806
6806
6806
6806
6806
6806
6806
6806
6806
6806
6842
6805
6805
6805
6950
6923
6923
6923
6923
6923
6923
6923
6952
6945
6942
6939
6939
6942
6948
6956
6902
6902
6902
6902
6902
6902
7038
7015
7015
7030
7030
7030
7051
7028
7028
7059
7010
7010
7010
7005
7005
7005
7005
7120
7120
7120
7120
7120
7120
7120
7120
7120
7120
7120
7030
7114
7120
7120
7030
7030
7116
7117
7116
7117
7116
7117
7120
7114
7114
7114
LONG
16432
16432
16432
16647
16613
16545
16545
16545
16545
16545
16545
16545
16545
16545
16545
16545
16545
16545
16545
16545
16545
16615
15030
15030
15030
14220
15210
15210
15210
15210
15210
15210
15210
15350
15450
15450
15450
15450
15450
16050
15957
16350
16350
16350
16350
16350
16350
16002
16121
16121
15730
15730
15730
15848
15725
15700
15434
14650
14650
14650
14335
14335
14335
14335
15640
15640
15640
15640
15640
15640
15640
15640
15640
15640
15640
15600
15656
15640
15640
15600
15600
15643
15648
15640
15644
15640
15648
15640
15656
15656
15656
LOCALITY
KIVALINA
KIVALINA
KIVALINA
POINT HOPE
CAPE LISBURNE
OGOTORUK CREEK
OGOTORUK CREEK
OGOTORUK CREEK
OGOTORUK CREEK
OGOTORUK CREEK
OGOTORUK CREEK
OGOTORUK CREEK
OGOTORUK CREEK
OGOTORUK CREEK
OGOTORUK CREEK
OGOTORUK CREEK
OGOTORUK CREEK
OGOTORUK CREEK
OGOTORUK CREEK
OGOTORJK CREEK
OGOTORUK CREEK
CAPE DYER
ULO LAKE
ULO LAKE
ULO LAKE
NUVAGAPAK PT
UMIAT
UMIAT
UMIAT
UMIAT
UMIAT
UMIAT
UMIAT
IKPIKPJUK R
IKPIKPUK R
IKPIKPUJUK RIVER
IKPIKPUK RIVER
IKPIKPUK RIVER
IKPIKPUK RIVER
UTUKOK R
KAOLAK RIVER
CAPE BEAUFORT
CAPE BEAUFORT
CAPE BEAUFORT
CAPE BEAUFORT
CAPE BEAUFORT
CAPE BEAUFORT
WAINWRIGHT
NEVAT PT
NEVAT PT
MEADE RIVER PO
MEADE RIVER PO
MEADE RIVER PO
PEARD BAY
MEADE RIVER PO
MEADE RIVER PO
CAPE SIMPSON
BULLEN
BULLEN
BULLEN
BARTER ISLAND
BARTER ISLAND
BARTER ISLAND
BARTER ISLAND
BARROW
BARROW
BARROW
BARROW
BARROW
BARROW
BARROW
BARROW
BARROW
BARROW
BARROW
POINT BARROW
BARROW
BARROW
BARROW
POINT BARROW
POINT BARROW
POINT 3ARROW
BARROW
BARROW
BARROW
BARROW
BARROW
BARROW
BARROW
BARROW
BARROW
DATE
06
06
06
08
08
08
08
07
06
08
06
06
06
06
06
06
06
06
08
06
07
COLLECTOR NAME
960
960
960
938
938
966
966
959
959
959
959
959
959
959
959
959
959
859
960
963
963
963
966
949
948
966
966
966
966
966
947
959
959
959
959
959
947
958
966
966
966
966
966
966
938
948
948
966
966
966
947
960
960
946
966
966
966
966
966
966
951
938
966
966
966
966
966
966
966
966
966
966
946
946
947
950
960
947
959
950
950
950
950
950
949
952
952
952
MPMONMNCCCC CN VODDADADVDVNVNAAVNONANANANAVDCAHUADANAMNDNADADOOADVDANDVD ZECAANDADANDOCOVOVCVCODAKX<X<X<<MOOCCCFEPFYPYPPPPrrPrPPrrrrrgagaccrreTp
FCTLTETATITCTCITCTCORNRENNNVINSTEEACZLTETLEUVUATELSLETEENEZETNELEZSZZVE
StvevrrereTTTettsteTezTTeTseeeteEevs
NESE EEN
EZEEE
BUCKNELL
BUCKNELL
BUCKNELL
ANDERSON
ANDERSON
ARGUS
ARGUS
JOHNSON
JOHNSON
JOHNSON
JOHNSON
JOHNSON
JOHNSON
JOHNSON
JOHNSON
JOHNSON
JOHNSON
JOHNSON
JOHNSON
JOHNSON
JOHNSON
VIERECK
FLOCK
FLOCK
FLOCK
ARGUS
FROHNE
LEPAGE
SUDA
SUDA
SUDA
SUDA
SUDA
BLACK
GEIST
GEIST
GEIST
GEIST
GEIST
BLACK
SHUSHAN
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ANDERSON
IRVING
IRVING
ARGUS
ARGUS
ARGUS
BLACK
GEIST
GEIST
BLACK
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
BARKALOW
ANDERSON
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
BLACK
BLACK
BLACK
BLACK
GEIST
SCHOLANDER
SHETLER
THOMAS
THOMAS
THOMAS
THOMAS
THOMAS
TILLETT
WARD
WARD
WARD
COL NO HERB
ALA
ALA
ALA
7sc
TSC
SWA
GWA
GWA
GWA
GWA
AWA
GWA
GWA
GWA
BWA
GWA
GWA
ALA
ALA
ALA
ALA
ALA
ALA
ALA
ALA
GWA
ALA
NAO
GWA
GWA
BWA
GWA
BWA
msc
ALA
ALA
ALA
ALA
ALA
Tsc
WIS
GWA
GWA
GWA
SWA
GWA
GWA
SC
ALA
ALA
5WA
GWA
SWA
tsc
ALA
ALA
TSC
6WA
5WA
GWA
GWA
BWA
GWA
msc
mse
5WA
GWA
BWA
GWA
6WA
GWA
GWA
6WA
GWA
GWA
TSC
TSC
Isc
rsc
ALA
Isc
ALA
ALA
ALA
1sc
1SCc
TSC
TSC
ALA
Isc
Isc
+ NO
26574
26588
26590
256461
256441
9186
9171
9185
9168
13226
21742
21719
21721
21805
32752
256420
26024
28159
28952
27985
29310
256443
256442
3109
3108
256421
29035
26951
256423
256427
256440
256429
256424
256422
256425
28099
256426
17453
6548
6549
224853
224825
224852
256430
6565
224912
224876
Specimens Cited
TAXON SPC S V HYB PROV QUAD LAT LONG LOCALITY DATE
1873 2833 14 153 7116 15640 BARROW 04 07
1873 28 33 14 153 7119 15640 BARROW 26 06
1873 26833 14 153 7120 15640 POINT BARROW 07 07
18753 28 3 3 14 153 7117 15647 BARROW 12 07
32 COMMUTATA
1875 32 2 2 5935 13630 INSPECTOR CREE 12 7
1873 32 2 2 5935 13630 INSPECTOR CREE 12 7
1875 32 2 2 5935 13629 THREE GUARDSME 14 7
1873 32 = 2 5941 13632 NADAHINI RIVER 13 7
1875 32 2 2 5941 13632 NADAHINI RIVER 13 7
1873 32 2 2 5947 13636 HAINES RD M70 04% 08
1875 32 2 4 5910 13358 LLEWELLYN GL 15 07
1875 32 2 16 5831 12434 SUMMIT PASS 19 07
1875 32 2 24 5723 12248 BUCKINGHORSE R 03 09
1875 32 2 24 5723 12248 BUCKINGHORSE R 03 09
1875 32 2 65 5130 11830 REVELSTOKE 08 06
16735 32 2 65 5150 11815 KINBASKET R 23 05
1875 32 7 338 6257 12858 OGRADY Le Oe OT
1875 32 7 338 6202 12810 SEWLYN MTSe 8 8
1875 32 7 338 6202 12810 FLAT RIVER 8 8
1875 32 7 339 6241 12750 GRIZZLY BEAR L 10 8
1875 32 7 339 6205 12735 BRINTNELL LAKE 03 08
1873 32 7 339 6205 12735 BRINTNELL LAKE 18 07
2873 32 7 339 6205 12735 BRINTNELL LAKE 18 07
1873 32 7 339 6205 12735 BRINTNELL LAKE 18 07
1873 32 7 339 6205 12735 BRINTNELL LAKE 16 07
1875 32 7 339 6205 12735 BRINTNELL LAKE 18 07
1873 32 7 339 6205 12735 BRINTNELL LAKE 18 07
1873 32 7 339 6205 12735 BRINTNELL LAKE 16 07
1875 32 12 30 6257 13009 ITSI RANGE 3i 07
1873 32 12 30 ©6240 13112 CANOL RD M234 10 08
1875 32 12 30 6238 13108 CANOL RD M222 17 08
1875 32 12 30 6300 13025 CANOL RD M268 28 08
1673 32 12 35 6130 13302 CANOL RD M95 15 07
1875 32 12 35 6130 13302 CANOL RD M95 15 07
1875 32 14 10 9838 13625 GEIKE INLET 29 06
1873 32 14 10 5859 13606 MUIR INLETe GL O1 7
1873 32 14 10 5857 136035 MUIR INLETe GL 02 7
1875 32 14 10 5858 13606 MUIR INLETe GL 29 6
1875 32 14 10 5859 13606 MUIR INLETe GL O01 7
1873 32 14 10 5855 13603 MUIR INLETe GL 28 6
1875 32 14 10 5855 13603 MUIR INLETe GL 28 6
1673 32 14 10 5855 13603 MUIR INLETe GL 28 6
1873 32 14 10 5855 13603 MUIR INLETe GL 28 6
1875 32 14 lu 5855 13603 MUIR INLETe GL 28 6
1875 32 14 10 5855 13603 MUIR INLETe GL 28 6
1873 32 14 10 5855 13603 MUIR INLETe GL 28 6
1875 32 14 10 5855 13603 MUIR INLETe GL 28 6
1875 32 14 10 5855 13603 MUIR INLET? GL 28 6
1875 32 14 luo 5855 13603 MUIR INLETe GL 28 6
1875 32 14 11 5825 13540 GUSTAVUS. 04 07
1875 32 14 11 5825 13540 GUSTAVUS. 04 07
1875 32 14 11 5825 13540 GUSTAVUS. 04 07
1873 32 14 11 5825 13540 GUSTAVUS. 04 07
18753 32 14 11 5825 13540 GUSTAVUS. 04 07
1873 32 14 11 5825 13540 GUSTAVUS. 04 07
186735 32 14 11 5825 13540 GUSTAVUS. 04 07
1875 32 14 11 5825 13540 GUSTAVUS. 04 07
1875 32 14 11 5825 13540 GUSTAVUS. 04 07
1673 32 14 11 5839 134135 TAKU B NUNATAK 10 08
1873 32 14 23 5335 16650 UNALASKA IS 06 06
1673 32 14 34 5738 15220 ISTHMUS PT 18 06
1873 32 14 42 5819 15406 KUKAK BAY 01 07
1873 32 14 45 5928 13605 MOSQUITO LAKE 45 7
1875 s2 14 45 5928 13605 MOSQUITO LAKE 45 7
1875 32 14 45 5928 13602 MOSQUITO LAKEe 11 7
1875 32 14 45 5905 13524 DAVIDSON GL 05 07
1673 32 14 45 5937 13508 WHITE PASS 23 07
1873 32 14 45 5921 13625 FLOWER MT 11 07
18735 32 14 46 5932 13940 YAKUTAT. 23 6
1875 32 14 46 5932 13940 YAKUTAT. 23 6
1873 32 14 46 5933 13930 SITUK RIVER. 23 6
1873 32 14 46 5932 13940 YAKUTAT. 23 6
1873 32 14 46 5915 13830 TANIS LAKE 21 6
1873 32 14 46 5915 13830 TANIS LAKE 19 6
1873 32 14 46 5915 13830 TANIS LAKE 19 6
1873 32 14 46 5915 13830 TANIS LAKE i935 16
1875 32 14 46 5915 13830 TANIS LAKE ZS 6
1873 32 14 46 5915 13830 TANIS LAKE 19> 16
1873 32 14 46 5915 13830 TANIS LAKE 19) 6
1873 32 14 46 5915 13830 TANIS LAKE 19°56
18753 32 14 4@ 5915 13830 TANIS LAKE er '6
1873 32 14 46 5952 13945 YAKUTAT BAY. 14 6
18735 32 14 46 5952 13945 YAKUTAT BAY. 14 6
1873 32 14 46 5952 13945 YAKUTAT BAY. 14 6
18735 32 14 46 5952 13945 YAKUTAT BAY. 14 6
1675 32 14 46 5952 13945 YAKUTAT BAY 14 6
1873 32 14 46 5952 13945 YAKUTAT BAY 14 6
952
952
966
950
967
967
967
967
967
967
914
943
943
943
953
958
967
967
967
967
939
939
939
939
939
939
939
939
960
944
944
944
944
944
928
967
967
967
967
967
967
967
967
967
967
967
967
967
967
967
967
967
967
967
967
967
967
967
966
944
949
B99
967
967
967
929
914
909
967
967
967
967
967
967
967
967
967
967
967
967
967
967
967
967
967
967
967
COLLECTOR NAME
HU @
rrees
DODAPANAANAPN ANAND GOV ANANAODGVAMPTONAOD AMUAADVYAVVGVOAVAVNGVGVONOVOVAANOVDGOOOVOVGVGOOOCPHPPP FP PFPCIITIITITIr£e£eztenc rT Iirraqgnooo
= =
#85 853 €C CG
=
=
SESTEECTCFCTEETETTETETTEAZTETETEZEVAIMNAANr Ss E
=zt=z
EREEEZCZEEEECTC ETC LCZEELCEZEEE
WARD
WARD
WELSH
WIGGINS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
VIERECK
EASTWOOD
RAUP
RAUP
RAuUP
CALDER
TAYLOR
CODY
CopY
Copy
CODY
RAUP
RAUP
RAuUP
RAUP
RAUP
RAUP
RAUP
RAUP
CALDER
PORSILD
PORSILD
PORSILD
PORSILD
PORSILD
ANDERSON
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
BESCHEL
BEALS
LEPAGE
COVILLE
ARGUS
ARGUS
ARGUS
COOPER
EASTWOOD
SCHEUBER
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
COL NO
1094
10P1
5701
12428
HERB + NO
TSC 224877
ISC 224824
GWA
ISC 224823
GWA
GWA
GWA
GWA
GWA
GWA
A
ALA
GWA
ALA
SWA
SWA
DAO
NAO
NAO
NAO
ALA
ALA
GWA
GWA
SWA
GWA
ALA 14802
ALA 14807
US 2349263
TSC 256386
GH
GH
TSC 256387
GH
TSC 256388
GWA
6WA
GWA
SWA
GWA
GWA
GWA
GWA
GWA
GWA
GWA
GWA
GWA
GWA
GWA
GWA
GWA
GWA
GWA
GWA
GWA
GWA
GWA
SWA
US 2034651
CAN
US 373486
GWA
GWA
GWA
- 742232
19851
19852
58175
58178
14821
14809
209
Appendix B
TaxON SPC S V HYB PROV QUAD
1873
1873
1873
1673
1873
1873
1873
1873
1873
1873
1873
1873
1675
1473
1873
1673
1873
1873
1873
1873
1873
1873
1873
1873
1673
1873
1875
1873
1873
1873
18735
1673
36
1675
1673
1673
1873
1873
1875
18735
39
1873
1673
1873
1673
1873
1673
1873
1873
1873
1873
1875
1873
1873
1873
1673
1873
1473
1673
1673
1673
1673
1673
1873
1673
1673
1873
1873
1673
1875
18673
1673
1673
1873
1873
1873
1873
1673
14673
1675
1873
1675
1673
1873
1673
210
32 14
32 14
32 14
32 14
32 14
32 14
32 14
32 14
32 14
32 14
32 14
32 14
32 14
32 14
32 14
32 14
32 14
Se 14
32 14
32 14
32 14
32 14
32 14
32 14
52 14
32 14
32 14
32 14
32 14
32 14
32 14
32 14
SPHENOPHYLLA
36 7
36 14
36 14
36 14
36 14
36 14
36 14
ARBUSCULOIDES
39 1
39 1
39 1
39 2
39 2
39 fe
39 2
39 2
39 2
39 2
39 2
39 2
39 7
39 U
39 7
39 TA
39 i
39 if
39 i
39 1
39 7
39 7
39 A
39 i
39 i/
39 7
39 7
39 7
39 1/
39 11
39 11
39 12
39 12
39 12
39 12
39 12
39 12
39 le
39 12
39 12
39 12
39 12
39 12
39 12
LAT
595e
5952
9952
5952
5952
5952
5952
5910
5924
5924
9910
5924
5924
6040
6108
6108
6108
6110
6111
6112
6110
6102
6140
6140
6155
6152
6228
6226
6228
6303
6324
6324
70135
6445
6445
6430
6546
6950
7005
5907
5640
5310
5925
5845
5840
5849
5840
5705
5705
5658
5705
6916
6920
6923
6813
6819
6842
6820
6517
6307
6205
6205
6224
6158
6251
6251
6051
6050
5350
5341
6727
6404
6404
6335
6158
6110
6110
6122
6122
6103
6146
6103
6137
LONG
13945
13945
13945
13945
13945
13945
13945
13823
13900
13900
13823
13900
13900
14903
14544
14544
14547
14540
14547
14613
14540
14801
14900
14900
15425
15433
15115
15115
15115
14610
15025
15025
12940
16620
16620
16525
16855
14220
14335
Live7
11115
11338
12610
12540
12417
12507
12440
12235
12235
12238
12235
12815
12815
13310
13500
13340
13407
13330
12651
12630
12735
12735
12714
12145
12125
12123
11542
9425
10518
10515
13959
13925
13925
13554
14033
13845
13845
13905
13903
13831
138637
13831
13553
LOCALITY
BAY
BAY
BAY.
BAYe
BAY.
YAKUTAT
YAKUTAT
YAKUTAT
YAKUTAT
YAKUTAT
YAKUTAT BAY-
YAKUTAT BAY.
ALSEK RIVER
HARLEQUIN L
HARLEQUIN L
ALSEK RIVER
HARLEQUIN L
HARLEQUIN L
TUNNEL
THOMPSON -PASS
THOMPSON PASS
THOMPSON PASS
WORTHINGTON GL
WORTHINGTON GL
ROBE LAKE
WORTHINGTON GL
COLLEGE FIORD
MATANUSKA VY
MATANUSKA VY
HEAD OF BIG Re
HEAD OF BIG Re
KAHILTNA GL
KAHILTNA GL
KAHILTNA GL
TANGLE LAKES
THOROFARE R
THOROFARE R
CAPE DALHOUSIE
FEATHER RIVER
FEATHER RIVER
NOME
LTL DIOMEDE IS
NUVAGAPAK PT
BARTER ISLAND
SLAVE RIVER
WATERWAYS
WHITEMUD CREEK
LIARD HOT SPR
PETERSEN CREEK
TETSA RIVER
RACING RIVER
SUMMIT PASS
BEATTON RIVER
BEATTON RIVER
BEATTONe®RIVER
BEATTON RIVER
ANDERSON RIVER
ANDERSON R
TUKTOYAKTUK
AKLAVIK
INUVIK
MACKENZIE DELT
INUVIK
NORMAN WELLS
DAL Le
BRINTNELL L
BRINTNELL L
MACKENZIE MTSe
FORT SIMPSON
FORT SIMPSON
FORT SIMPSON
LOWER HAY R
MCCONNELL R
CANDLE LAKE
BIRCHBARK CR
RAMPART HOUSE
DAWSON
DAWSON
MAYO
WHITE R
KLUANE LAKE
KLUANE LAKE
BURWASA
BURWASA
KLUANE LAKE
PTARMIGANHEART
KLUANE LAKE
CONGLOMERATE M
DATE
14
14
14
14
16
16
16
10
02
14
967
967
967
967
967
967
967
965
965
965
965
965
965
932
935
958
967
957
967
967
O5i7
935
940
940
950
950
956
956
956
953
956
956
963
966
966
966
926
966
966
927
939
939
966
966
966
960
943
943
943
943
943
963
965
963
957
957
947
970
957
967
939
939
963
955
939
939
951
967
965
965
951
932
949
944
944
966
966
944
948
944
948
944
960
COLLECTOR NAMF
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VIERECK
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VIERECK
VIERECK
SETCHELL
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VIERECK
VIERECK
VIERECK
VIERECK
VIERECK
COOPER
PALMER
PALMER
DRURY
DRURY
VIERECK
VIERECK
VIERECK
GJAERVOLL
VIERECK
VIERECK
CODY
PEGAU
PEGAU
WELSH
PORSILD
ARGUS
ARGUS
RAUP
RAUP
RAUP
ARGUS
ARGUS
ARGUS
CALDER
RAUP
RAUP
RAUP
RAUP
RAUP
Copy
SCOTTER
CODY
CODY
CODY
COWAN
WELSH
CODY
CODY
RAUP
RAUP
ROWLANDS
CODY
RAUP
RAUP
LEwIS
MACINNES
ARGUS
ARGUS
LOAN
BEAUCHAMP
CALDER
ANDERSON
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ARGUS
ARGUS
RAUP
RAUP
RAUP
RAUP
RAUP
CALDER
cOL NO HFRB + NO
6150
6149
6161
6160
6188
6185
6184
7604
7610
7605
7604
7610
7605
613
1887
2165A
B4OB
2210A
8478
8752
2210A
224
3e3
385
3°66
3A85
1053
1051
1053
1911
1083
1083
131°9
wi
wi
5047
1676A
§912
5848
550
9040
9027
5014
6037
4971
24674
10511
10014
10074
10098
10073
12611
6933
13061
9673
2622
27
10015
9564
17566
9263
9317
528
A156
9104
9079
237
63
4753
4753
390
2AR9
9717
9314
5087
5NR6
12451
13318
12098
13651
12098
24407
GWA
GWA
GWA
SWA
GWA
GWA
GWA
SWA
GWA
GWA
FSLC
FSEC
FSEC
F
ISC
CAN
GWA
GWA
GWA
BWA
ALA
F.
ALA
ALA
CAN
PAN
BWA
ALA
ALA
CAN
BWA
ALA
AO
GWA
ALA
GWA
CAN
SWA
GWA
ALA
ALA
ALA
GWA
GWA
GWA
ALA
ALA
ALA
ALA
ALA
ALA
NAO
GWA
NAO
NAO
DAO
NAO
OTF
DAO
NAO
ALA
ALA
DAO
ALA
ALA
ALA
NY
uwo
GWA
GWA
GWA
ALA
ISC
rse
1SC
GWA
5WA
ALA
ALA
ALA
ALA
GWA
BWA
341
333
329
755140
256389
B4i14
823977
5966
5966
11629
11628
11611
32715
32708
46440
14478
14838
14842
17430
20068
20046
20051
20069
20050
32677
32675
32650
1215
14846
14845
24338
14840
14841
493
255938
255940
255864
20048
20044
20052
20049
Specimens Cited
TAXON SPC S V HY¥B PROV QUAD LAT LONG LOCALITY DATE COLLECTOR NAME COL NO HERB + NO
1875 39 12 35 6135 13302 CANOL RD M95 14% 06 944 A F PORSILD 10361 TSC 255937
1875 39 12 39 ©6057 13825 KLUANE L 15 07 944 H “ RAUP 12471 ALA 20047
1875 39 ae 39 ©057 13825 KLUANE L 15 07 944 H ™ RAUP 12471 GWA
1675 39 12 40 ©6045 13615 MENDENHALL R 30 06 966 G W ARGUS 5078 GWA
18735 39 12 4Q 6045 13615 MENDENHALL R 30 06 966 G W ARGUS 5076 GWA
1873 39 12 4u 6045 13615 MENDENHALL R 30 06 966 G W ARGUS 5079 GWA
1875 39 12 40 6047 13735 PINE CREEK 23 06 944 H “ RAUP 11859 ALA 20045
1875 39 12 40 6047 13735 PINE CREEK 23 06 944 H ¥ RAUP 11A59 GWA
1873 39 12 41 6010 13442 CARCROSS 13 07 949 J M GILLETT 3797 SE 255939
1875 39 12 42 6022 13351 LTL ATLIN L 19 08 943 H ™ RAUP 113P8 ALA 20053
1875 39 12 42 6022 13351 LTL ATLIN L 19 08 943 H ™ RAUP 113RB GWA
18735 39 12 42 ©6006 13231 ALA HWY M793 24 06 966 S L WELSH 5474 GWA
1875 39 12 44 6007 12848 WATSON LAKE 07 06 956 G W ARGUS 268 GWA
1875 39 12 44 ©6003 12855 UPPER LIARD R 25 06 966 G W ARGUS 5031 GWA
1875 39 14 41 5844 15701 SALMON CREEK 28 08 948 E LEPAGE 24118 YSC 255943
1875 39 14 63 6029 14922 MOOSE PASS 01 06 941 J P ANDERSON 6497 ISC 255792
1873 39 14 67 6126 14255 MCCARTHY 08 06 961 E L LITTLEe JR. 18375 GWA
1875 39 14 68 6157 14517 COPPER CENTER 08 06 961 E L LITTLEr JR. 1R3A4 GWA
1873 39 14 68 6140 14510 TONSINA 28 07 958 L A SPETZMAN 2151 CAN
1875 39 14 69 6150 14731 SHEEP MT 17 07 948 J P ANDERSON 19745 1SC 255860
18735 39 14 69 6150 14731 SHEEP MT 15 07 948 J P ANDERSON 19672 TSC 255941
i873 39 14 71 6158 15438 BIG RIVER 19 7 950 W H DRURY 4322 CAN
1873 39 14 73 6135 15915 RUSSIAN MISSe 2 7 949 W H DRURY 1738 CAN
1873 39 14 73 6137 15930 ANIAK 18 6 949 W H DRURY 1506 CAN
1675 39 14 73 6132 15927 ANIAK 20 6 949 W H DRURY 1568 CAN
1875 39 14 73 +6135 15932 ANIAK 17 6 949 WH DRURY 1429 CAN
1875 39 14 73 6135 15932 ANIAK 17 6 949 W H DRURY 1430 CAN
1873 39 14 73 6132 15927 ANIAK RIVER 20 6 949 W H DRURY 1581 CAN
1875 39 14 73 6132 16018 KALSKAG 25 05 933 W ® MILLER 261C SC 255944
1875 39 14 73 =6132 16018 KALSKAG 25 05 933 WR MILLER 2€1C ALA 2855
1875 39 14 78 6239 16012 ANVIK 13 06 924 J CHAPMAN 43 NY
1875 39 14 79 6259 15604 TAKOTNA 26 07 941 J P ANDERSON 7413 JSC 255867
1675 39 14 79 6259 15604 TAKOTNA 26 07 941 J © ANDERSON 7413 ALA 491
1875 39 14 80 6257 15536 MCGRATH 13. 6 949 WH DRURY 1355 CAN
1875 39 14 80 6257 15536 MCGRATH 7 6 950 W H DRURY 3614 CAN
1875 39 14 80 6257 15536 MCGRATH 7 6 950 WH DRURY 3619 CAN
18735 39 14 80 6257 15536 MCGRATH 7 6 950 WH DRURY 3620 CAN
1875 39 14 80 6250 15452 BIG RIVER 28 7 950 W H DRURY 4388 CAN
1875 39 14 80 6233 15337 FAREWELL Le 2 8 949 WH DRURY 2289 CAN
1873 39 14 80 6232 15543 KUSKOKWIM Re 20 7 949 W H DRURY 2139 CAN
1873 39 14 80 6257 15535 MCGRATH 15 06 961 E L LITTLEs JR. 1A446 SWA
1875 39 14 83 6218 14518 GAKONA 08 06 944 J © ANDERSON 8507 TSC 255861
1875 39 14 83 6218 14518 GAKONA 18 06 944 J P ANDERSON 8506 TSC 255862
1875 39 14 83 6218 14518 GAKONA 19 06 957 G W ARGUS 1043 SWA
1875 39 14 83 6218 14518 GAKONA 19 06 957 G WwW ARGUS 1043 ALA 8403
1875 39 14 83° 6216 14522 GULKANA R BR 10 06 967 L A VIERECK R208 GWA
1675 39 14 84 ©6238 14102 ALA HWY M1222 27 05 961 E L LITTLEse JR. 1R223 GWA
1873 39 14 85 6323 14321 TANACROSS 27 07 944 J P ANDERSON R791 ISC 255863
18753 39 14 85 6320 14236 TETLIN JCT 01 07 963 M BARTHOLOMEW 7063 ALA 25908
1875 39 14 85 6322 14232 TETLIN JCT 21 06 963 J NAVA 95 ALA 23945
1875 39 14 85 6330 14348 ROBERTSON R 26 07 957 L A SPETZMAN 905 ALA 6844
1875 39 14 85 6318 14338 TANANA R 09 07 957 L A SPETZMAN 470 ALA 6843
1875 39 14 86 6347 14545 DONNELY DOME 30 06 957 G W ARGUS 1062 GWA
1873 39 14 86 6356 14655 W FK LTL DELTA 03 06 941 L J PALMER 553 ISC 255869
1875 39 14 86 6356 14655 W FK LTL DELTA 06 941 L J PALMER 4O7 5629
1875 39 14 86 6356 14655 W FK LTL DELTA 03 06 941 L J PALMER 553 ALA 5158
1873 39 14 87 6351 14858 HEALY 22 07 939 J P ANDERSON 5708 I1SC 255870
1875 39 14 87 6351 14858 HEALY 22 07 939 J © ANDERSON 5708 ISC 255870
1875 39 14 87 63540 14930 TEKLANIKA R 21 07 956 G W ARGUS 614 SWA
1875 39 14 87 6355 14930 SAVAGE R 18 06 928 Y MEXIA 2015 ALA 10201
1875 39 14 87 6355 14930 SAVAGE R 18 06 928 Y MEXIA 2016 ALA 10202
1673 39 14 87 6338 14934 IGLOO CR CAMP 10 07 939 A NELSON 3766 ISC 255945
18735 39 14 87 6338 14934 IGLOO CR CAMP 10 07 939 A NELSON 3766 ALA 492
1873 39 14 87 6355 14725 DRY CREEK 09 07 962 L fF VIERECK 58R1 SWA
1875 39 14 87 6355 14725 DRY CREEK 25 06 962 L A VIERECK 5719 GWA
1875 39 14 88 6331 15003 TOKLAT R BR 22 07 965 S L WELSH 4855 ISC 247207
1875 39 14 89 6315 15520 NIXON FORK 12 6 950 WH DRURY 3726 CAN
1873 39 14 90 6310 15631 OPHIR 26 06 940 E SCAMMAN 1831 GWA
1875 39 14 94 6430 16530 NOME G GASSER ALA 25007
1873 39 14 94 6430 16530 NOME 27 06 954 C HELLER 1157 ALA 24322
1875 39 14 97 6445 15857 GALENA 18 06 961 E L LITTLEs JR. 18507 GwA
18735 39 14 97 6445 15657 GALENA 18 06 961 E L LITTLEe UR. 18598 GWA
1873 39 14 97 6420 158435 KALTAG 04 07 931 C H ROUSE 43 ALA 25460
1873 39 14 97 ©6420 15843 KALTAG 04 07 931 C H ROUSE 43 ALA 2076
1873 39 14 98 6445 15530 RUBY 03 07 931 C H ROUSE 50 ALA
18735 39 14 98 6445 15530 RUBY 03 07 931 C H ROUSE 47 ALA 2016
1875 39 14 98 6445 15530 RUBY 03 07 931 C H ROUSE 47 ALA 25426
1873 39 14 98 6445 15530 RUBY 03 07 931 C H ROUSE 50 ALA 25462
1873 39 14 98 6445 15530 RUBY 03 07 931 C H ROUSE 50 ALA 2011
1875 39 14 100 6450 14743 FAIRBANKS 10 05 940 J P ANDERSON 6080 ISC 255869
1873 39 14 100 6450 14743 FAIRBANKS 12 05 940 J P ANDERSON 6087 ISC 255868
1875 39 14 100 6448 14755 CHENA PUMP RD 22 06 956 G W ARGUS 411 GWA
1875 39 14 100 6453 14752 SMITH LAKE 13 06 956 G W ARGUS 2A68 GWA
1873 39 14 100 6451 14752 UNIV EXP FARM 14 06 956 G W ARGUS 332 GWA
1875 39 14 100 6453 14751 SMITH LAKE 22 06 956 G W ARGUS 430 GWA
1875 39 14 100 6453 14752 SMITH LAKE 13 06 956 G W ARGUS 286 ALA 4522
1875 39 14 100 6451 14752 UNIV EXP FARM 14% 06 956 G W ARGUS 332 ALA 4524
1875 39 14 100 6453 14751 SMITH LAKE 22 06 956 G W ARGUS 430 ALA 4512
1875 39 14 100 6453 14751 SMITH LAKE 22 06 956 G W ARGUS 430 ALA 224868
1873 39 14 100 6500 14740 FOX 02 06 964 V L HARMS 2631 ALA 32624
1873 39 14 100 6450 14745 FAIRBANKS 06 933 L J PALMER 179 ALA 5257
ot
Appendix B
TAXON SPC S V HYB PROV QUAD LAT LONG LOCALITY DATE
1a730—Ss (39 14% 100 6450 14743 FAIRBANKS 17 06
1873 46039 14 100 6450 14743 FAIRBANKS 06
1873 39 14 100 6450 14743 FAIRBANKS 20 07
18730 (339 14% 100 6450 14743 FAIRBANKS 20 06
13873 = 339 14 100 6450 14743 FAIRBANKS 20 06
1s73 0s (339 14 100 6450 14743 FAIRBANKS 06
1873s (339 14 100 6450 14743 FAIRBANKS 06
18730339 14 100 6450 14743 FAIRBANKS 06
i873 39 14 100 6450 14743 FAIRBANKS 06
18730339 14 100 6450 14743 FAIRBANKS 06
1a73s) 0399 144 100 6450 14743 FAIRBANKS 06
1873s (339 14% 100 6450 14743 FAIRBANKS 06
13873 39 14 100 6450 14743 FAIRBANKS 20 06
18730339 14% 100 6450 14743 FAIRBANKS 20 06
187339 14 100 6450 14743 FAIRBANKS 17 06
1873 = 339 144 100 6450 14743 FAIRBANKS 20 07
1673 0339 14% 100 6450 14743 FAIRBANKS 06
1873 39 14 100 6450 14743 FAIRBANKS 06
18730339 14% 100 6450 14743 FAIRBANKS 06
1873 39 14 100 6450 14743 FAIRBANKS 06
1873 39 144 100 6450 14743 FAIRBANKS
1373 339 14% 100 6451 14748 FAIRBANKS 21 07
18730339 14 100 6452 14752 SMITH LAKE 05 06
1873039 14 100 6451 14736 CHENA RIVER 07 06
1873 6039 14 100 6452 14752 UNIV EXP FARM 17 06
1873339 14 100 6452 14752 UNIV EXP JARM 29 07
1873 39 14 100 6448 14744 VAN HORN ROAD 26 05
1873 39 14 100 6448 14744 VAN HORN ROAD 26 05
1873 = 339 14 100 6448 14744 VAN HORN ROAD 24 07
1873 ©6039 14 100 6448 14744 VAN HORN ROAD 24 07
1873 339 144 100 6453 14749 COLLEGE 08 07
1873 39 14 101 6454 14625 CHENA RIVER 06 07
1873 = 339 144% 101 6412 14555 BIG DELTA 21 05
1873s (339 14 101 6453 14645 UPPER CHENA R 22 06
1873 39 14 101 6457 14657 UPPER CHENA R_ 25 05
1873 39 14 101 6457 14657 UPPER CHENA R- 25 05
1673 «039 14 102 6410 14146 FRANKLIN 14 07
1873 «39 144% 102 6410 14146 FRANKLIN 14 07
l1a73.0s 39 14 103 6515 14245 CHARLEY RIVER 08 08
1873 39 14 104 6532 14446 CENTRAL 20 06
1873 39 14 104 6534 14448 CENTRAL 29 05
1873 39 14% 104 6531 14513 MILLER HOUSE 12 07
1873 339 14 104 6531 14513 MILLER HOUSE 13 06
1873 39 14 104 6527 14526 EAGLE CR CAMP 12 06
1873 §=639 14 104 6558 14407 CIRCLE 11 07
1873 39 144 105 6531 14833 LIVENGOOD 08 07
1873 39 14% 105 6531 14833 LIVENGOOD 08 07
18730339 14 105 6507 14747 CHATANIKA R 03 08
1873 39 14 105 6507 14747 CHATANIKA R 03 08
1873 339 14 105 6506 14745 CHATANIKA R BR OW 06
1873 «39 14 105 6502 14740 ENGINEER CREEK 02 06
1873 339 14 105 6527 14815 TOLOVANA R 30 07
1873 339 14 105 6550 14750 LIVENGOOD 23 06
1873 39 14% 105 6531 14833 LIVENGOOD 19 06
1873 «6039 14 113 6655 16231 KOTZEBUE 11 08
187339 14 117 6654 15141 BETTLES
1373339 14 119 6634 14516 FORT YUKON 16 06
1873 «39 14 119 6634 14516 FORT YUKON 24 05
1873 339 14 120 6644 14334 BLACK RIVER 08 07
1873 339 14% 121 6715 14140 PORCUPINE R 20 07
1873 39 14 121 6715 14140 PORCUPINE R 19 07
1873 «39 144 121 6715 14140 RAMPART 21 07
1873 39 14 121 6708 14208 PORCUPINE R 16 08
1673 39 14 121 6716 14138 PORCUPINE R 10 08
1a73.— 339 14 121 6723 14350 SMALL LAKE 25 07
1873 39 14 121 6723 14350 SMALL LAKE 26 07
1873 39 14 121 6723 14350 SMALL LAKE 27 07
1873 39 14 121 6723 14350 SMALL LAKE 27 07
1873 39 14 124 6725 15006 WISEMAN 02 08
1873 «339 14% 134 6820 15100 KANAYUT CR 17 07
1873s 39 14 136 6804 14503 OLD JOHN LAKE 15 08
1873 39 14 137 6822 14355 SHEENJEK R 23 06
1873 39 14 137 6822 14355 SHEENJEK R 15 06
1873 39 14% 137 6©822 14355 SHEENJEK R 15 06
1873 86039 14 137 6822 14355 SHEENJEK R 15 06
1673 339 14 137 6822 14355 SHEENJEK R 15 06
18673 39 144% 141 6920 15210 UMIAT 22 06
1873 39 14 141 6920 15210 UMIAT 22 06
1673 39 14 141) 6920 15210 UMIAT 22 06
16730339 14 141 6922 15208 UMIAT 27 06
1873 39 14% 141 6923 15210 UMIAT 27 07
1873 39 14 141 6923 15210 UMIAT 27 07
1673 39 14 141 6923 15210 UMIAT 28 07
1873 «39 14 141 6923 15210 UMIAT 29 07
1873 39 14 141 6923 15210 UMIAT 28 07
1673 «39 14% 141 6922 15210 UMIAT 03 06
1873 «6339 1% 141 ©6922 15210 UMIAT 12 08
212
933
933
927
927
933
933
933
933
933
933
933
927
927
933
931
933
933
933
933
947
963
955
955
964
963
966
966
966
966
967
966
951
964
965
965
941
941
964
964
961
949
945
966
965
944
944
957
957
965
964
965
959
940
966
939
961
961
957
957
957
957
961
961
957
957
957
957
937
949
957
956
956
956
956
956 ~
951
951
951
953
966
966
966
966
966
964
964
COLLECTOR NAME
ON<<<<<YCCCOMDWDVDBDUFMNNNUHANCCCUMMOAaMA<KK KOOCOCFCMMMe<rCCFrrreearrrrrrrnunaarrrrrrrerrrrrrrereer eee
rmroeverrrerceterprrrrpry?mo FTeoecececeoeccececeoecocecoccc
urrreztectuvuvs=z
yNrrrarrase
Dp
oe ees pe
PALMER
PALMER
PALMER
PALMER
PALMER
PALMER
PALMER
PALMER
PALMER
PALMER
PALMER
PALMER
PALMER
PALMER
PALMER
PALMER
PALMER
PALMER
PALMER
PALMER
PEWE
SCHUCK
SMITH
SMITH
VIERECK
VIERECK
VIERECK
VIERECK
VIERECK
VIERECK
VIERECK
ARGUS
copy
VIERECK
VIERECK
VIERECK
ANDERSON
ANDERSON
VIERECK
HARMS
LITTLE
SCAMMAN
SCAMMAN
TRENT
VIERECK
ANDERSON
ANDERSON
ARGUS
ARGUS
HARMS
HARMS
HARMS
OS FARRELL
SCAMMAN
ARGUS
JOHNSON
LITTLE,
LITTLE,
SHETLER
BUCKLEY
BUCKLEY
BUCKLEY
DEAN
DEAN
SHETLER
SHETLER
SHETLER
SHETLER
SCAMMAN
SPETZMAN
SHETLER
KESSEL
KESSEL
KESSEL
KESSEL
KESSEL
BUCKLEY
BUCKLEY
BUCKLEY
SMITH
SUDA
SUDA
SUDA
SUDA
SUDA
WEST
WEST
UR.
JR.
JRe
cOL NO
151
1R7
1789
17A9
149
151
2766
18262
5161
3495
866
7723
8972
8972
B
18451
18215
386AF
118
113B
1A1
672AF
TiISAF
697AF
695AF
9N0
2720
AF161
S66A
$22
S22
$21
$21
1848
24166
24266
26366
27866
26266
7179
T5N7
HERB + NO
ALA 32134
ALA 5260
ALA 5170
ALA 5910
ALA 5185
ALA 5227
ALA 5228
ALA 5267
ALA 5261
ALA $258
ALA 5254
ALA $259
ALA $186
ALA $921
ALA 5961
ALA 5978
ALA 5256
ALA $255
ALA 5252
ALA 5262
1SC 255935
ALA 26179
ALA
ALA 10746
FSLC 271
FSLC 44
GWA
GWA
GWA
GWA
GWA
GWA
GWA
FSLC 314
FSLC
FSLC
ISC 255866
ALA 494
FSLC 285
ALA 32601
GWA
GWA
ALA 8395
ALA 32690
FSLC
Isc 255942
GWA
SWA
ALA 6793
ALA 32632
ALA 32648
ALA 32595
ALA 9632
GWA
GWA
YSC 255936
GWA
GWA
ALA 3909
ALA 5098
ALA 5092
ALA 5116
ALA 24095
ALA 24094
ALA 4095
ALA 4120
ALA 4105
ALA 4104
GWA
DAO = 32753
GWA
ALA 5043
ALA 3547
ALA
ALA 5044
ALA 22873
GWA
ALA 5033
ALA 26814
ALA 10653
GWA
GWA
SWA
GWA
GWA
GWA
SWA
Specimens Cited
TAXON SPC S V HYB PROV QUAD LAT
41
1873
18735
1873
1673
1875
1873
1673
1873
1873
1873
1873
1873
1873
1873
1873
1873
1873
18735
1873
1673
1873
1875
1873
1873
1873
1673
1873
1873
1873
1673
1873
18735
1873
1873
1873
1873
1873
18735
1873
1873
1873
1873
1873
1873
18735
1873
1873
1873
1873
1873
1873
1873
1873
1873
1873
1873
1673
1873
1673
1873
1873
1873
1873
1873
1873
1873
1873
1673
1873
1873
15673
1873
18673
1873
1873
1873
1873
1873
1873
1873
1873
1873
1873
1873
1673
1873
1873
Pe ee ee ee ee ee ee oe ee ee oe oo oe
5 EP a Oe 2 a i ee 0 0 2 ee a ee ee 0 0 2 a a a 0 2 2 ~~ 2 —2 2 2 2 i 2 ee ee
6920
6944
6835
6812
6833
6922
6847
6811
6857
6827
6838
6734
6404
6155
6324
6315
6325
6344
6344
6339
6339
6339
6339
6333
6323
6325
6344
6325
6344
6356
6335
6330
6330
6330
6330
6330
6325
6434
6428
6420
6420
6433
6410
6522
6527
6520
6528
6511
6545
6512
6512
0657
6600
6715
6715
6715
6725
6725
6725
6756
6806
6806
6806
6855
6855
6855
6828
6817
6822
6840
6822
6839
6836
6926
6930
6920
6920
0930
6930
6925
6902
6902
6902
6902
6902
7030
7030
LONG
13350
13120
13404
13115
12828
13932
14028
13947
13832
13845
13945
13832
13925
15425
14543
14540
14900
14855
14855
14932
14932
14932
14932
14934
14852
14900
14855
14900
14855
14728
14845
15004
15004
15004
15004
15004
15050
16533
16512
15843
15843
14455
14136
14556
14526
14550
14739
16535
16510
16645
16645
15326
14915
14140
14140
14140
15007
15007
15007
16222
16545
16545
16545
164350
16110
16110
15442
15125
14355
14345
14355
14100
14356
14347
14500
14500
14500
15130
15130
15210
15532
16350
16350
16350
16350
15730
15730
LOCALITY
MACKENZIE DELT
ATKINSON PT
MACKENZIE DELT
HYNDMAN L
ANDERSON R
FIRTH RIVER
FIRTH RIVER
TIMBER CRe
BABBAGE Re
DOG CREEK
BABBAGE RIVER
OLD CROW Re
DAWSON
HEAD OF BIG Re
CASTNER GL
RAINBOW MT
CANTWELL
MCKINLEY PARK
MCKINLEY PARK
TEKLANIKA R
TEKLANIKA R
TEKLANIKA R
TEKLANIKA R
SABLE PASS
WINDY CREEK
CANTWELL
MCKINLEY PARK
CANTWELL
MCKINLEY PARK
DRY CREEK
CANTWELL
TOKLAT R
TOKLAT R
TOKLAT R
TOKLAT R
TOKLAT R
WONDER L
NOME
NOME
KALTAG
KALTAG
GOODPASTER R
FRANKLIN
TWELVE MILE CR
EAGLE CR CAMP
STEES HWY M88
FOSSIL CREEK
DUCK CREEK
SALMON L ROAD
PORT CLARENCE
PORT CLARENCE
HELPMEJACK CR
DALL RIVER
RAPID R
RAPID R
RAPID R
WISEMAN
WISEMAN
WISEMAN
NOATAK R
OGOTORUK
OGOTORUK
OGOTORUK CREEK
PITMEGEA RIVER
DRIFTWOOD CR
DRIFTWOOD CR
KURUPA RIVER
ANAKTUVUK PASS
SHEENJEK R
SHEENJEK R
SHEENJEK R
FIRTH RIVER
SHEENJEK R
JAGO RIVER
SADLEROCHIT R
LAKE PETERS
LAKE PETERS
COLVILLE RIVER
COLVILLE RIVER
UMIAT
COLVILLE RIVER
CAPE BEAUFORT
CAPE BEAUFORT
CAPE BEAUFORT
CAPE BEAUFORT
MEADE RIVER
MEADE RIVER
AKE
CREEK
CREEK
DATE
14
21
17
07
05
1
12
1
13
951
966
951
966
965
970
970
970
970
970
970
970
949
950
967
965
956
956
956
959
960
959
960
953
960
939
939
939
939
962
964
956
956
956
956
956
958
968
949
931
931
956
941
957
966
964
953
968
969
901
901
901
901
957
957
957
939
939
949
961
966
966
966
959
969
969
952
949
“956
956
956
961
956
957
948
959
959
951
951
966
952
966
966
966
966
966
966
COLLECTOR NAME
DADNDONODP<XAAMNUYUTCOMBMBAAITr SPAANOOOArFCCCCCEZEENANRDAOVFCNVHCPONOMAFrADDAGAV GOFF SE EP ESP KEP PP rFPODOMVOArFrrFr=EccNNNNNYNAOA?PD yS
SZ. 2.25222 PAGS. 2a ze
PEELE EPP
n RP2anVETz
LeVvrerr
HVDVDEZE
ZaeEzezEtE
LINDSEY
SCOTTER
LINDSEY
SCOTTER
SCOTTER
WELSH
WELSH
WELSH
WELSH
WELSH
WELSH
RIGBY
CALDER
DRURY
VIERECK
VIERECK
ARGUS
ARGUS
ARGUS
MURIE
MURTE
MURIE
MURIE
MURIE
MURIE
NELSON
NELSON
NELSON
NELSON
VIERECK
VIERECK
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
VIERECK
PEGAU
SCAMMAN
ROUSE
ROUSE
JOHNSON
ANDERSON
SHETLER
TRENT
VIERECK
GJAEREVOLL
PEGAU
PEGAU
WALPOLE
WALPOLE
MENDENHALL
MENDEHALL
BUCKLEY
BUCKLEY
BUCKLEY
ANDERSON
ANDERSON
JORDAL
JOHNSON
ARGUS
ARGUS
ARGUS
SHETLER
PEGAU
PEGAU
HODGDON
SPETZMAN
KESSEL
KESSEL
KESSEL
LITTLE?
SCHALLER
CANTLON
SPETZMAN
SHETLER
SHETLER
CHAMBERS
CHAMBERS
SUDA
HODGDON
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
JRe
COL NO HERB + NO
CAN 215919
GWA
CAN 215920
GWA
GWA
OTF 1190
OTF 1212
BRY 91159
BRY 91476
OTF 1207
OTF 1189
BRY 92301
TSC 256394
CAN
GWA
GWA
GWA
GWA
ALA 4903
GWA
GWA
GWA
GWA
GWA
GWA
ALA 517
ALA 516
ISC 256398
ISC 256401
GWA
FSLC 295
GWA
GWA
ALA 4904
ALA 22497
ALA &750
GWA
GWA
GH
ALA 25465
ALA 2017
ALA 7064
ALA 536
ALA 3730
ALA 32692
FSLC 309
CAN
GWA
GWA
US 378703
US 378736
US 377286
US 377367
GWA
ALA
ALA 26776
ALA 27503
TSC 256400
TSC 256399
GWA
GWA
GWA
SWA
GH
GWA
GWA
GH
US 2032478
GWA
ALA 3679
ALA 5854
GWA
wis
6H
US 2032217
GH
CAN
GH
US 2312148
GWA
6H
GWA
GWA
GWA
GWA
GWA
SWA
213
Appendix B
TAXON SPC S V HYB PROV QUAD LAT
1873
1875
1875
1875
1875
45
1875
175
14873
1875
1873
1873
1673
1873
1873
1873
1673
18675
1873
1873
1873
1873
1873
1873
1873
1675
1873
1873
1873
1873
1873
1873
1873
1873
1873
1873
1873
1873
1873
1873
1873
1875
1873
1873
1873
1873
1875
1873
1873
1873
1873
1873
1873
1873
1873
1873
1873
1873
1873
1873
1873
1873
1873
1673
1873
1873
1673
1873
1873
1873
1873
1673
1873
1673
1873
1873
1873
1873
1673
1873
1873
1873
1873
1873
1673
1873
1673
18735
214
ARCTICA
14
14
14
14
14
147
147
147
147
147
001
002
002
002
002
134
134
140
297
297
339
339
339
339
7030
7030
7030
6953
7003
5935
5935
5935
5935
5935
5935
5935
5935
5935
5935
5935
5935
5935
5935
5935
5935
5935
5935
5935
5935
5935
5935
5800
5838
5831
5831
5831
5650
5500
5400
5400
6941
6933
6927
6940
6949
6905
6445
6447
6408
6408
6338
6346
6338
6345
6345
6305
6257
6257
6205
6205
6205
6205
6205
6205
6205
6205
6127
6050
6050
6050
6050
6050
6902
6826
6855
6413
6350
6357
6247
6203
6203
61035
61035
6103
6149
6149
6103
6132
6120
6155
6055
LONG
15730
15730
15730
15708
15715
13630
13629
13629
13629
13629
13629
13629
13629
13629
13630
13630
13630
13630
13630
13630
13630
13630
13630
13627
13627
13627
13628
13300
12442
12434
12434
12434
13150
12700
12800
12700
13455
13346
13302
12827
12859
10456
12928
12937
8317
8317
12719
12719
12707
6832
6832
12850
12858
12858
12735
12735
12735
12735
12735
12735
12735
12735
12405
9425
9425
9425
9425
9425
13943
13840
13710
14006
13731
13510
13104
12859
12859
13831
138631
13831
13835
13835
13831
13302
13300
13238
13843
LOCALITY
MEADE
MEADE
MEADE
MEADE
MEADE
RIVER
RIVER
RIVER
RIVER
RIVER
INSPECTOR CREE
THREE GUARDSME
THREE GUARDSME
THREE GUARDSME
MT GLAVE
MT GLAVE
MT GLAVE
MT GLAVE
MT GLAVE
INSPECTOR
INSPECTOR
INSPECTOR
INSPECTOR
INSPECTOR
INSPECTOR
INSPECTOR
INSPECTOR CREE
INSPECTOR CREE
THREEGUARDSMAN
THREEGUARDSMAN
THREEGUARDSMAN
HAINES RD M56
TULSEQUAH LAKE
MT ST GEORGE
SUMMIT PASS
SUMMIT PASS
SUMMIT PASS
STIKINE RIVER
SKEENA CROSS
MT THORNHILL
SMITHERS
HOOPER ISLAND
RICHARDS IS
TUKTOYAKTUK
ANDERSON RIVER
MICHOLSON IS
CAMBRIDGE BAY
CACHE CREEK
MOUNTAIN RIVER
CORAL HARBOR
CORAL HARBOR
RINGSTONE CR
RINGSTONE CR
N REDSTONE R
FROBISHER BAY
FROBISHER BAY
O GRADY LAKE
O GRADY LAKE
O GRADY LAKE
BRINTNELL
BRINTNELL
BRINTNELL
BRINTNELL
BRINTNELL
BRINTNELL
BRINTNELL
BRINTNELL
Se» NAHANNI Re
MCCONNELL
MCCONNELL
MCCONNELL
MCCONNELL
MCCONNELL
BRITISH MTS.»
SAM LAKE
BLOW RIVER
SIXTY MILE
BARLOW
KENO HILL
MT SHELDON
UPPER HYLAND L
UPPER HYLAND L
KLUANE LAKE
KLUANE LAKE
KLUANE LAKE
PTARMIGAN HRT
PTARMIGAN HRT
KLUANE LAKE
ROSE RIVER
ROSE RIVER
LOWER LAPIE R
SLIMS RIVER
CREE
CREE
CREE
CREE
CREE
CREE
CREE
nr ferret
DADRDADAD
RDe
966
966
966
960
956
967
967
967
967
967
967
967
967
967
967
967
967
967
967
967
967
967
967
967
967
967
967
958
960
943
943
943
954
954
954
966
966
965
965
965
959
963
963
948
948
963
963
963
948
948
967
967
967
939
939
939
939
939
939
939
970
965
964
964
964
-967
970
970
970
949
960
949
944
960
960
944
944
944
948
948
944
944
944
944
967
COLLECTOR NAME
ARGUS
ARGUS
ARGUS
GEIST
WIGGINS
KOODO@
=zzetez
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
VIERECK
VIERECK
VIERECK
VIERECK
HEUSSER
CALDER
RAUP
RAUP
RAUP
ANDERSON
CALDER
CALDER
CALDER
SCOTTER
SCOTTER
SCOTTER
SCOTTER
SCOTTER
THOMSON
JOHNSON
KVALE
CODY
CODY
JOHNSON
JOHNSON
JOHNSON
SENN
SENN
CODY
CODY
CODY
RAUP
RAUP
RAUP
RAUP
RAUP
RAUP
RAuUP
RAUP
SCOTTER
MACINNES
MACINNES
MACINNES
MACINNES
MACINNES
RIGBY
WELSH
WELSH
CALDER
CALDER
GILLETT
PORSILD
CALDER
CALDER
ANDERSON
RAUP
RAUP
RAUP
RAUP
RAUP
PORSILD
PORSILD
PORSILD
MURRAY
PrRPPrPtetteteTezeTEZEzATETETZEE
OPPPITIIITIICCCCrPCCCHWNCAARKRAAOGOTIITIATILCICEZLELIOMMMEEMMCAOMVDVODACCCCTIITICOFFTFTHA9DGVDDOAVHVDWHVHVVHVHVH}GOH}EO}GTES®*®
HAMM EeESssesVrPrmMerrerrrarwrrrree i eesese FF FSSECCCPrTEEECOC ZZTETCETETZYP IF vPVve ss EP
COL NO HERB + NO
5254
5262
5261
SeNe
13935
GWA
GWA
GWA
ALA 27726
US 2264189
GWA
GWA
GWA
GWA
GWA
GWA
GWA
GWA
GWA
GWA
GWA
GWA
GWA
BWA
GWA
GWA
GWA
GWA
GWA
GWA
GWA
GWA
GWA
GWA
ALA 19468
ALA 19516
ALA 19571
TSC 85492
GWA
GWA
GWA
GWA
GWA
GWA
GWA
BWA
wIS
DAO 32713
DAO 32712
1SC 255813
TSC 255814
AO 32709
DAO 32710
DAO 32711
ISC 255927
TSC 255815
DAD 32703
DAO 32700
A0 32704
ALA 16409
ALA 16415
ALA 16410
ALA 16399
ALA 16398
ALA 16411
ALA 16414
ALA 16416
GWA
UWO
UWO
UWO
UWO
UwWO
BRY 92366
OTF 1208
OTF 1209
DAO 32728
GWA
DAO 32729
TSC 255830
GWA
GWA
TSC 255822
ALA 19517
ALA 19472
ALA 19572
ALA 19479
BWA
TSC 255929
TSC 255820
1SC 255928
SWA
Specimens Cited
TAXON SPC S V HYB PROV QUAD LAT
1873
1873
1673
1873
1875
1875
18735
1875
1875
1873
1873
1875
1875
1873
1873
1675
1873
1873
1875
1873
1875
1873
1875
1675
1873
1875
1673
1873
1875
18735
1875
1873
1873
1673
18753
1875
1873
1875
1873
1873
1873
1873
1875
1875
1873
1875
1873
1873
1875
1873
1873
1873
1875
1873
1873
1873
1873
1873
1873
1875
1873
1875
18753
1875
1875
1873
1873
1875
1673
1873
1873
1875
1875
1875
1873
18753
1873
1875
1875
1875
1873
1873
1873
1873
1873
1875
18753
1675
1875
1873
1875
45
45
45
45
45
45
45
45
45
45
45
45
45
45
45
45
45
45
45
45
45
45
45
45
45
45
45
45
45
45
45
45
45
45S
45
45
45
45
45
45
45
45
45
45
45
45
45
45
45
45
45
45
45
4S
45
45
45
45
45
45
45
4S
45
45
45
45
45
45
45
45
45
45
45
45
45
45
39
39
39
39
39
41
43
010
010
010
010
010
010
010
010
010
010
010
010
010
010
010
010
010
010
010
010
010
010
010
6055
6055
6055
6057
6057
6037
6005
5854
5854
5854
5859
5859
5859
5859
5859
5858
5858
5859
5858
5858
5858
5858
5858
5858
5859
5859
5859
5859
5859
5857
5857
5857
5857
5857
5857
5857
$857
5857
5857
5857
5857
5855
5855
5855
5855
5855
5855
5855
5855
5832
5832
5817
5830
5830
5830
5818
5826
5826
5818
5818
5837
5839
5839
58535
5855
5853
5825
5817
5256
5255
5249
5255
5255
5255
5255
5255
5255
5157
5132
5132
5132
5132
5132
5132
$145
5156
5145
5155
$212
5212
5207
LONG LOCALITY
138435
138435
13843
13825
13825
13508
13040
136035
13603
13603
13606
13606
13606
13606
13606
13606
13606
13606
13606
13606
13606
13606
13606
13606
13606
13606
13606
13606
13606
136035
136035
136035
13603
13603
13602
13602
13602
13602
13602
13602
13602
136035
13605
13603
13603
13605
13603
136035
13603
13555
13555
13419
13415
13415
13415
13423
13435
13435
13423
134235
13418
13415
13413
13420
13420
13420
RIVER
RIVER
RIVER
SLIMS
SLIMS
SLIMS
SLIMS RIVER
SLIMS RIVER
MT MCINTYRE
RANCHERIA
MUIR INLET? GL
MUIR INLETe GL
MUIR INLET? GL
MUIR INLET? GL
MUIR INLET? GL
MUIR INLET? GL
MUIR INLET? GL
MUIR INLET? GL
MUIR INLET? GL
MUIR INLET? GL
MUIR INLET? GL
MUIR INLET? GL
MUIR INLET? GL
MUIR INLET? GL
MUIR INLETe GL
MUIR INLET? GL
MUIR INLET? GL
MUIR INLETe GL
MUIR INLET? GL
MUIR INLETe GL
MUIR INLET? GL
MUIR INLET? GL
MUIR INLET? GL
MUIR INLET? GL
MUIR INLETe GL
MUIR INLET? GL
MUIR INLET? GL
MUIR INLET? GL
MUIR INLET? GL
MUIR INLET? GL
MUIR INLET? GL
MUIR INLET? GL
MUIR INLET? GL
MUIR INLETe GL
MUIR INLET? GL
MUIR INLET? GL
MUIR INLET? GL
MUIR INLET? GL
MUIR INLET? GL
MUIR INLET? GL
MUIR INLETe GL
MUIR INLET? GL
BEARDSLEE IS
BEARDSLEE IS
MT ROBERTS
JUNEAU ICE FD
JUNEAU ICE FD
JUNEAU ICE FD
MT ROBERTS TR
MENDENHALL GLA
MENDENHALL GLA
MT ROBERTS TR
MT ROBERTS TR
SHOEHORN MT
TAKU B=NUNATAK
TAKU B=-NUNATAK
GILKEY GL
GILKEY GL
GILKEY GL
13445 INSPIRATION PT
13424 MT ROBERTS
17315€ATTU ISLAND
17255€ ATTU ISLAND
17318€ALEXAI POINT
17255€ ATTU
172558 ATTU
17255FATTU
17255€ATTU
172558 ATTU
17255EATTU
ISLAND
ISLAND
ISLAND
ISLAND
ISLAND
ISLAND
17739ELITTLE KISKA I
17900F AMCHITKA IS
179008 AMCHITKA IS
179008AMCHITKA IS
179002 AMCHITKA IS
17900€ AMCHITKA IS
17900E AMCHITKA IS
17645 ADAK ISLAND
17644 ADAK ISLAND
17645 ADAK ISLAND
17645 ADAK ISLAND
17408 ATKA ISLAND
17412 ATKA ISLAND
17430 ATKA ISLAND
967
966
967
944
944
949
948
967
967
967
967
967
967
967
967
967
967
967
967
967
967
967
967
967
967
967
967
967
967
967
967
967
967
967
967
967
967
967
967
967
967
967
967
967
967
967
967
967
967
928
928
925
952
952
952
967
967
967
967
967
966
966
966
958
958
958
928
967
945
941
944
945
945
945
945
945
945
937
934
967
967
967
967
967
950
950
944
966
949
947
945
COLLECTOR NAME COL NO HERS + NO
D F MURRAY 828 SWA
D F MURRAY 407 GWA
D F MURRAY 777 «GWA
H “ RAUP 12584 ALA 19570
H “ RAUP 12584 GWA
J ™ GILLETT 3426 AO 32731
J P ANDERSON inya2 TSC 256281
G w ARGUS 6394 GWA
G Ww ARGUS 6357 GWA
G WwW ARGUS 6394 GWA
G wW ARGUS 6523 GWA
G Ww ARGUS 6522 GWA
6 W ARGUS 6521 GWA
G W ARGUS 6487 GWA
G w ARGUS 6480 GWA
G W ARGUS 6449 GWA
G Ww ARGUS 6442 GWA
G w ARGUS 6484 GWA
G Ww ARGUS 6466 GWA
G W ARGUS 6465 GWA
6 W ARGUS 6446 GWA
6 w ARGUS 6451 SWA
G Ww ARGUS 6459 GWA
G Ww ARGUS 6458 GWA
G W ARGUS 6513 SWA
G Ww ARGUS 6505 GWA
G Ww ARGUS 6515 GWA
G W ARGUS 6514 GWA
G W ARGUS 6530 GWA
G Ww ARGUS 6556 GWA
G Ww ARGUS 6557 GWA
G wW ARGUS 6558 GWA
6 Ww ARGUS 6555 GWA
G wW ARGUS 6554 SWA
G W ARGUS 6432 GWA
G Ww ARGUS 6426 GWA
G w ARGUS 6425 GWA
6 W ARGUS 6423 GWA
G w ARGUS 6422 GWA
G Ww ARGUS 6420 GWA
G w ARGUS 6419 GWA
G Ww ARGUS 6379 GWA
G w ARGUS 6378 GWA
G wW ARGUS 6360 GWA
G w ARGUS 6381 GWA
G Ww ARGUS 6373 GWA
G6 W ARGUS 6372 GWA
6 w ARGUS 6380 GWA
G Ww ARGUS 63830 GWA
J P ANDERSON 728 SC 256483
J P ANDERSON wer TSC 255807
J P ANDERSON 2a242 ISC 255934
G Ww ARGUS 93 ALA 4737
6 W ARGUS 90 GWA
G W ARGUS °o1 GWA
6 Ww ARGUS 6635 GWA
6 W ARGUS 6661 GWA
G Ww ARGUS 6655 GWA
6 w ARGUS 6634 GWA
G wW ARGUS 6640 GWA
R F BESCHEL 15442 GWA
R & BESCHEL 15469 GWA
R — BESCHEL 15483B GWA
Cc HEUSSER 1 GWA
Cc HEUSSER 2 GWA
c HEUSSER 3 GWA
Y MEXIA 2014 ALA 10206
L A VIERECK 8622 GWA
J P ANDERSON 543 ISC 255826
F BEALE SeNe TSC 256494
P J SALAMUN AG7 WIS
G w SOULE 377 ISC 255796
G w SOULE 12 ISC 255801
G Ww SOULE 539 ISC 255824
6 Ww SOULE 375 YSC 255912
G w SOULE 2n2 ISC 255798
G Ww SOULE 61 1sc255803
JH STEENIS 4563 wis
HATFIELD 18C255916
wM KLEIN 2382 CS
w ™M KLEIN 2264 CS
w M KLEIN 2263 GWA
w M KLEIN 2264 GWA
WM KLEIN 2263 CS
T P BANK 4267 ALA 6700
T P BANK 4271 ALA 6715
P E€ GLASSBURG SeNe TSC 255911
V L HARMS 5252 ALA 32642
T P BANK 1228 ALA 6705
) OLIVER 31 ISC 255914
C tL YORK 45-436 ISC 255793
215
Appendix B
TaxON SPC S V HYB PROV QUAD LAT LONG LOCALITY DATE
1875 45 14 21 5259 16855 ANANIULIAK IS 25 06
1673 45 14 21 5255 16903 ANANIULIAK IS 15 07
18753 45 14 21 5250 16902 ANANIULIAK IS 06 07
1873 45 14 21 5255 16905 ANANIULIAK IS 05 07
1873 45 14 21 5245 16858 NIKOLSKI 12 07
1875 45 14 21 5240 16858 NIKOLSKI 10 08
1873 45 14 21 5245 16858 NIKOLSKI 13 08
1873 45 14 21 5245 16858 NIKOLSKI 04 08
1875 45 14 22 5310 16842 MT RECHESHNOI 18 07
1873 45 14 23 5335 16650 UNALASKA 30 05
1873 45 14 23 5335 16650 UNALASKA 23 07
1875 45 14 23 5335 16650 UNALASKA 23 07
1873 45 14 23 5335 16650 UNALASKA 21 08
1875 45 14 23 5352 16632 UNALASKA 04 07
1873 45 14 23 5335 16650 UNALASKA 04 O07
1673 45 14 23 5335 16650 UNALASKI 20 05
1873 45 14 23 5335 16650 UMNAK IS 20 08
1673 45 14 25 5459 16228 COLD BAY
1875 45 14 25 5459 16228 COLD BAY
1875 45 14 34 5746 15312 KODIAK IS 01 08
1873 45 14 34 5747 15300 PILLAR MT 14 06
1875 45 14 34 5712 15332 THREESAINTS BY 05 08
18735 45 14 35 5730 15413 KODIAK REFUGIU 18 06
1873 45 14 38 5710 17015 ST PAUL IS 18 07
1875 45 14 38 5710 17015 ST PAUL IS 08 07
1873 45 14 38 5710 17015 ST PAUL IS 08 07
1875 45 14 38 5710 17016 ST PAUL IS 08 07
1873 45 14 39 5834 16146 CAPE PEIRCE slp Weed
1875 45 14 39 5834 16146 CAPE PEIRCE » I eT 4
1873 45 14 39 5834 16146 CAPE PEIRCE cower
1873 45 14 42 5846 15556 IDAVAIN L 29 08
1875 45 14 42 $834 15548 DUMPLING MT 17 06
1873 45 14 42 5835 15551 DUMPLING MT 28 06
1875 45 14 42 5817 15509 BROKEN MT 06 08
1873 45 14 42 5835 15551 DUMPLING MT 01 08
1873 45 14 43 5805 15305 RASPBERRY IS 15 06
1873 45 14 43 58604 15304 RASPBERRY IS 25 08
1873 45 14 43 5804 15304 RASPBERRY IS 25 08
1875 45 14 43 5804 15304 RASPBERRY IS 25 08
18735 45 14 43 5804 15304 RASPBERRY IS 25 08
18753 45 14 43 5804 15304 RASPBERRY IS 25 08
1873 45 14 43 5804 15304 RASPBERRY IS 25 08
1873 4S 14 45 5927 13519 SKAGWAY 23 07
1875 45 14 45 5927 13519 SKAGWAY 20 08
1875 45 14 46 5915 13830 TANIS LAKEe 22 6
18673 4§ 14 46 5915 13830 TANIS LAKE- 22 6
1873 45 14 46 5915 13830 TANIS LAKEe 22 6
1873 45 14 46 5915 13830 TANIS LAKEe 22 6
1875 45 14 46 5915 13830 TANIS LAKE. 22 6
1875 45 14 46 5915 13830 AZIMUTH PEAKe 20 6
1875 4S 14 46 5915 13830 AZIMUTH PEAKe 20 6
1875 45 14 46 5915 13830 TANIS LAKEe 22 6
1873 45 14 46 5915 13830 TANIS LAKEe 22 6
1873 4S 14 Si 5940 15500 IGIUGIG 21 07
1873 45 14 51 5943 15510 ILIAMMA L 19 07
1873 45 14 52 5945 15841 LAKE KULIK 14 07
1873 45 14 56 6024 17235 ST MATTHEW IS 08 07
1873 45 14 56 6024 17238 ST MATTHEW IS 04 07
18735 4S 14 56 6024 17238 ST MATTHEW IS 12 07
1875 45 14 56 6024 17238 ST MATTHEW IS 02 07
18753 45 14 56 602% 17238 ST MATTHEW IS 02 07
1873 45 14 56 6024 17238 ST MATTHEW IS 02 07
1875 45 14 56 6024 17238 ST MATTHEW IS 14 07
1873 45 14 56 6024 17238 ST MATTHEW IS 18 07
1873 45 14 56 6024 17238 ST MATTHEW IS 14 07
1873 45 14 56 602% 17238 ST MATTHEW IS 01 07
1873 45 14 56 6024 17238 ST MATTHEW IS
1675 45 14 56 6024 17238 ST MATTHEW IS 18 07
1875 45 14 56 602% 17238 ST MATTHEW IS 18 07
1873 45 14 56 6024 17238 ST MATTHEW IS 18 07
1875 45 14 56 6025 17235 ST MATTHEW IS 18 07
1873 45 14 56 6024 17235 ST MATTHEW IS 08 07
1875 45 14 56 6024 17238 ST MATTHEW IS 04 07
1673 45 14 56 6024 17238 ST MATTHEW IS 14 07
1873 45 14 56 6024 17238 ST MATTHEW IS 18 07
1875 45 14 56 6024 17238 ST MATTHEW IS 18 07
1873 45 14 56 6024 17238 ST MATTHEW IS 18 07
1875 45 14 56 6030 17300 HALL IS 09 08
1873 45 14 56 6030 17300 ST MATTHEW IS 21 07
1873 45 14 56 6030 17300 ST MATTHEW IS 19 07
1875 45 14 57 6012 16656 NUNIVAK IS 15 07
1873 45 14 57 6002 16543 NUNIVAK IS 10 07
1673 45 14 57 6004 16716 NUNIVAK IS 11 08
1875 45 14 57 6004 16716 NUNIVAK IS 11 08
1873 45 14 62 6020 15018 SKILAK L 09 08
1875 45 14 62 6020 15018 SKILAK L 09 08
1873 45 14 63 6006 14926 SEWARD 13 06
1673 45 14 63 6055 14938 HOPE 07 06
1873 45 14 63 6055 14938 HOPE 07 06
18735 45 14 63 6032 14932 MOOSE PASS V 10 06
1875 45 14 63 6026 14952 STETSON CR 03 08
216
962
962
962
962
962
962
962
962
962
938
938
938
943
941
941
932
926
950
962
963
962
938
941
941
941
970
970
970
954
965
966
954
954
945
946
946
946
946
946
946
934
919
967
967
967
967
967
967
967
967
967
965
965
950
966
966
966
966
966
966
966
966
966
966
966
966
966
966
966
966
966
966
966
966
957
957
957
938
965
965
965
952
952
941
941
941
951
951
COLLECTOR NAME
CCCeccouougaoaaocougucoceececececc ace cececec cece eC CC CAC ce OAONOVNGMODONOVGAVOACCZEEZTFLAEZAOOONEOTCKrOTTrrTrcsWsETEESETEErFneccsseseseEeETrEEED
goccac vvd
==
VVeVBISIrceeerreeervree Sey Ceres ecle Ce Srl 2a CSS SS Vee Ce ea cw ys
rPPrpuvdV7ID
GORDON
JOHNSON
JOHNSON
JOHNSON
JOHNSON
JOHNSON
JOHNSON
JOHNSON
JOHNSON
ANDERSON
ANDERSON
ANDERSON
BEALS
COLE
COLE
EYERDAM
MILLER
WILLIAMS
WILLIAMS
CLARK
JOHNSON
NYBAKKEN
JOHNSON
ANDERSON
COLE
COLE
COLE
DICK
DICK
DICK
CAHALANE
CHAUVIN
ESTABROOK
SCHALLER
SCHALLER
EYERDAM
EYERDAM
EYERDAM
EYERDAM
EYERDAM
EYERDAM
EYERDAM
ANDERSON
ANDERSON
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
HARMS
HARMS
RAUP
HARMS
HARMS
HARMS
HARMS
HARMS
HARMS
HARMS
HARMS
HARMS
HARMS
HARMS
HARMS
HARMS
HARMS
HARMS
HARMS
HARMS
HARMS
HARMS
HARMS
HARMS
KLEIN
KLEIN
KLEIN
ANDERSON
BOS
BOS
BOS
KLEIN
KLEIN
ANDERSON
ANDERSON
ANDERSON
CALDER
CALDER
cOL NO
HERB
WIS
WIS
WIS
WIS
WIS
wIs
WIS
WIS
WIS
Isc
1s¢
msc
1sc
ioe
WIS
TSE
ALA
GWA
GWA
Tsc
WIS
GWA
WIS
tsc
WIS
WIS
WIS
GWA
GWA
GWA
msc
ALA
ALA
ALA
ALA
TSC
TSsc
Tsc
TSC
msc
mse
msc
Isc
rsc
GWA
6WA
GWA
GWA
GWA
GWA
GWA
GWA
GWA
ALA
ALA
Tsc
ALA
ALA
ALA
ALA
ALA
ALA
ALA
ALA
ALA
ALA
ALA
ALA
ALA
ALA
ALA
GWA
GWA
GWA
GWA
GWA
GWA
ALA
ALA
ALA
mse
ALA
ALA
ALA
ALA
ALA
msc
TSC
msc
GWA
GWA
+ NO
256453
255910
255915
255799
255922
255979
2036
256481
255825
219361
39951
32682
2983
2982
255917
255834
255835
255832
256482
255833
255827
255809
85493
32559
32593
219357
32671
32670
32672
32666
32667
32669
32676
32681
32677
32665
32675
32680
32679
32678
32678
21486
21487
6620
255795
32293
32291
32292
23023
23029
255909
255797
255925
Specimens Cited
TaxON SPC S V HYB PROV QUAD LAT
1873
1873
1875
16735
1675
1873
1873
1873
1875
18753
1673
1873
1875
1873
1875
1875
1873
1873
1673
1675
1873
1875
1873
1475
1873
1675
16735
1873
1873
1873
18673
18735
1675
1675
1875
1873
1875
1673
1873
1875
1873
1873
16735
1873
1875
1875
1875
1875
1873
1873
1873
1875
1875
18753
18673
1875
1875
1875
1875
1873
1873
1873
187535
1875
1873
1873
1673
1875
1873
1875
1673
1875
1873
18735
1873
1873
16753
1873
1875
1875
1873
1873
1873
1873
1873
1873
1873
1873
1875
1873
1873
63
63
63
63
63
64
64
67
67
67
67
67
68
68
68
68
68
68
68
68
68
69
69
69
69
69
69
69
69
69
6049
6006
6006
6024
6027
6100
6100
6137
6135
6135
6136
6136
6108
6110
6110
6145
6110
6108
6108
6145
6108
6245
6245
6145
6140
6140
6140
6104
6146
6146
6117
6146
6155
6155
6146
6146
6146
6146
6146
6146
6137
6137
6137
6200
6200
6259
6259
6256
6259
6233
6233
6228
6233
6203
6203
6340
6341
6323
6323
6310
6331
6333
6313
6335
6315
6315
6347
6347
6347
6347
6305
6305
6345
6330
6330
6310
6357
6357
6320
6325
6325
6302
6303
6351
6351
6351
6351
6342
6342
6345
6331
LONG
14933
14927
14927
14945
14820
14702
14702
14158
14207
14207
14156
14158
14544
14542
14540
14630
14540
14548
14610
14630
14548
14915
14915
14915
14900
14900
14900
14905
14918
14918
14858
14918
15425
15425
15808
15808
15808
15808
15808
15808
15930
15930
16600
16434
16434
15604
15604
15601
15606
15336
15336
15350
15336
14714
14714
14215
14215
14345
14345
14330
14258
14552
14539
14603
14530
14530
14545
14545
14545
14545
14545
14545
14554
14535
14535
14451
14658
14658
14535
14545
14545
14552
14557
14858
14858
14858
14858
14929
14929
14920
14956
LOCALITY
HOPE
MT MARATHON
MT MARATHON
COOPER L
NELLIE JUAN GL
HEATHER IS
HEATHER IS
WRANGELL MT
N WRANGELL MT
N WRANGELL MT
N WRANGELL MT
N WRANGELL MT
THOMPSON PASS
WORTHINGTON GL
WORTHINGTON GL
TAZALINA GL
WORTHINGTON GL
THOMPSON PASS
VALDEZ GL
TAZALINE GL
THOMPSON PASS
WILLOW CR
WILLOW CR
WILLOW RD SUMM
MATANUSKA VY
MATANUSKA VY
MATANUSKA VY
GIRDWOOD MINE
HATCHER PASS
HATCHER
EKLUTNA GL
HATCHER
HEAD OF
HEAD OF
CANOE MT-
CANOE MTe
CANOE MT.
CANOE MTeo
CANOE MT.
CANOE MTe
ANIAK
ANIAK
KOKECHIK BAY
KUSILVAK MT
KUSILIVAK MT
TAKOTNA
TAKOTNA
TAKOTNA MT
TAKOTNA
FAREWELL MTeo
FAREWELL MTe
FAREWELL
FAREWELL MTe
TALKEETNA MTS
TALKEETNA MTS
MT FAIRPLAY
TAYLOR HWY M34
NUTZOTIN MTS
NUTZOTIN MTS
SLIPPERY RK CR
SIXTYML RD M21
RAPIDS LODGE
GULKANA GL
PILLSBURY DOME
GULKANA GL
GULKANA GL
DONNELY DOME
DONNELY DOME
DONNELLY DOME
DONNELY DOME
PAXSON
PAXSON
DONNELLY DOME
RAINBOW MT
RAINBOW MT
SLATE CR
Ww FK LTL DELTA
Ww FK LTL DELTA
PAXSON
BLACK RAPIDS G
BLACK RAPIDS G
PAXON
TANGLE LAKE
HEALY
HEALY
HEALY
HEALY
SANCTUARY R
SANCTUARY R
SAVAGE R CAN
POLYCHROME PAS
COLLECTOR NAF COL NO HERB + NO
951 J A CALDER 6241 GWA
951 J 4 CALDER 49AR6 «GWA
951 J A CALDER 5854 GWA
952 D KLEIN 186 ALA 23N4A
957 L A VIERECK 2239 GWA
957 L A VIERECK 2313 ALA B405
957 L A VIERECK 2313 GWA
967 D © MURRAY 1013 GWA
967 R w SCOTT 1584 SWA
967 R w SCOTT 15A1 SWA
967 R w SCOTT 1742 GWA
967 R w SCOTT 1620 SWA
935 J P ANDERSON 1RA8B TSC 255806
947 DUTILLY 21339 SC 255913
957 L 4 VIERECK 2211 ALA 6383
957 L A VIERECK 21°3 ALA 6384
957 L A VIERECK 211 GWA
967 L A VIERECK RUST GWA
957 L 4 VIERECK 22°?B SWA
957 L 4 VIERECK 2193 GWA
967 L *A VIERECK R490) «GWA
931 J © ANDERSON 977 ALA 27309
931 J © ANDERSON 977 SC 255829
951 G “ FROHNE 5124 ALA 21793
940 L J PALMER 454 ALA 5220
940 L J PALMER 393 =6ALA 5987
940 L J PALMER 391 ALA 5964
957 L A VIERECK 2051 GWA
965 S Lt. WELSH 4710 TSC 247N95
965 S L WELSH 4705 TSC 247099
965 S L WELSH 41°96 ISC 246150
965 S L WELSH 4714 TSC 247141
950 Ww H DRURY 4112 CAN
950 WH DRURY 4131 CAN
949 Ww H DRURY 1R21 CAN
949 W 4 DRURY 1896 CAN
949 WH DRURY 1894 CAN
949 WH DRURY 1893 CAN
949 WH DRURY 1892 CAN
949 W H DRURY 1820 CAN
949 Ww H DRURY 1496 CAN
949 WH DRURY 1494 CAN
960 N © BLURTON JONES 68 ALA 24433
945 H C KYLLINGSTAD 22 SC 255A17
945 H C KYLLINGSTAD 21 Sc 255921
941 J P ANDERSON 7377 ALA 497
941 J P ANDERSON 7377 SC 255919
948 R L LAYDEN 177 Isc 255923
SHO E SCAMMAN 1823 ALA 8398
949 WH DRURY 2764 CAN
949 WH DRURY 2725 CAN
949 W H DRURY 2R26 CAN
949 W H DRURY 2721 CAN
941 J © ANDERSON 69°98 ALA 4OR
941 J P ANDERSON 6998 ISC 255816
948 JP ANDERSON 19805 TSC 255804
963 J NAVA 69 ALA 23944
944 H MW RAUP 12779 ALA 19467
944 H ™ RAUP 12779 =SWA
957 L A SPETZMAN 723 ALA 6837
966 S L WELSH 5593 GWA
935 J P ANDERSON 2220 ISC 25582h
935 J P ANDERSON SeNe TSC 255924
948 J © ANDERSON 1ne58 YISC 255821
957 G Ww ARGUS 1141 GWA
957 G w ARGUS 1138 GWA
957 G W ARGUS 1073 GWA
951 w J CODY 4871 SASK 33105
951 w J CODY 4873 GWA
951 W J CODY 4871 GWA
965 V L HARMS 4196 ALA 32661
965 V L HARMS 4199 ALA 32657
964 V L HARMS 2843 ALA 32604
965 V L HARMS 3561 ALA 32656
965 V L HARMS 3575 ALA 32651
941 L J PALYER ALA 5155
941 L J PALMER GAG ALA 5627
941 L J PALMER 519 ALA 5637
966 C PARKER RM36 ALA 32687
957 L A VIERECK 2134 GWA
957 L A VIERECK 2134 ALA 8413
968 S L WELSH R207 GWA
968 S L WELSH A312 GWA
939 J P ANDERSON 5763 TSC 255810
939 J P ANDERSON 5763 ISC 255810
939 J P ANDERSON 5761 SC 255933
939 J P ANDERSON 5761 ISC 255933
956 G Ww ARGUS 696 GWA
956 G6 W ARGUS 696 ALA 4731
965 B FABER SeNeo ALA 30967
939 A NELSON 3781 ISC 255A0A
217
Appendix B
TAXON SPC S V HYB PROV QUA; LAT LONG LOCALITY DATE
18735 45 14 87 6331 14956 POLYCHROME PAS 14 07
15873 45 14 87 6323 14856 CANTWELL 10 07
1875 45 14 87 6335 14843 CARLO MT 30 07
1873 45 14 87 ©6336 14845 CARLO wT 30 07
1875 45 14 87 ©6356 14728 DRY CREEK 25 06
18753 45 14 88 6325 15020 CAMP EIELSON 22 07
1873 45 14 88 6325 15020 CAMP EIELSON 22 07
1873 4S 14 88 6325 15020 CAMP EIELSON 22 07
1875 45 14 88 6325 15020 CAMP EIELSON ee O07
1873 45 14 88 6332 15058 CAMP DENALI 23 07
1875 45 14 88 6325 15010 HWY PASS 21 07
1873 45 14 8&8 ©6325 15020 CAMP EIELSON 2? 07
1875 4S 14 88 6328 15010 HWY PASS 21 07
1873 45 14 88 6325 15020 CAMP EIELSON 27 07
1875 45 14 88 6328 15010 HWY PASS e107
1873 4S 14 88 6325 15020 CAMP EIELSON Hae LOTS
1875 45 14 88 6325 15020 CAMP EIELSON 22 07
1875 45 14 88 6325 15020 CAMP EIELSON MP ATA
1873 45S 14 86 6325 15020 CAMP EIELSON 2, 07
1873 45 14 88 6330 15100 KANTISHNA 03 07
1873 4S 14 88 6326 15033 MCK PK RD M68 19 06
1873 45 14 88 6326 15033 MCK PK RD “68 19 06
1873 45 14 88 ©6326 15033 MCK PK RD M68 19 06
1873 45 14 8&8 6328 15052 WONDER LAKE 19 07
1673 45 14 88 6328 15052 WONDER LAKE T9507
18673 45 14 88 6326 15012 MCK PK RD M63 28 06
1873 45 14 8B 6325 15015 MCK PK 8D M64 30 06
1673 45 14 88 6326 15012 MCK PK RD M63 27 06
1873 45 14 88 6325 15020 CAMP EIELSON 14 08
1873 45 14 88 6327 15050 WONDER L 21 07
1873 45 14 88 6324 15025 GLACIER CR 10 07
1675 45 14 88 6324 15025 GLACIER CR 10 07
1873 45 14 88 6324 15025 GLACIER CR 10 07
1873 45 14 88 6324 15025 GLACIER CR 10 07
1873 45 14 88 6324 15032 MULDROW GL 12 08
1873 45 14 88 6325 15020 MT EIELSON LUA
1873 45 14 88 6327 15050 WONDER L 02 08
1873 45 14 88 6326 15035 MCK PK RD M70 12 08
1873 45 14 92 6329 16202 ST MICHAEL 19 06
1873 45 14 93 6342 17029 SAVOONGA 27 06
1873 45 14 93 6321 17136 BOXER BAY 12 08
1875 45 14 93 6321 17136 BOXER BAY 12 08
1873 45 14 93 6342 17029 SAVOONGA 03 07
1875 45 14 93 ©6342 17029 SAVOONGA POF
18675 45 14 93 6342 17029 SAVOONGA 03 07
1873 45 14 93 ©6342 17029 SAVOONGA LENO?
1873 4S 14 93 6334 17053 KANGEE CAMP
1873 45 14 93 ©6322 17117 POWOOILIAK 17 08
1873 45 14 93 6322 17117 POWOOILIAK 17 08
1873 45 14 93 0322 17117 POWOOILIAK 17 08
16753 45 14 93 6342 17029 SAVOONGA 11 07
1873 4S 14 93 6333 17002 ST LAWRENCE IS
1875 45 14 93 ©6335 17010 BOXER R VALLEY 22 06
1673 45 14 94 ©6446 16523 MT DISTIN 04 07
1873 45 14 94 6430 16530 ANVIL MT 08 06
1873 45 14 94 6430 16530 NOME 17 06
1873 45 14 94 6430 16530 NOME 16 06
1873 4S 14 94 6446 16630 CAPE WOOLLEY 16).
1875 45 14 94 6430 16525 NOME 14 07
1873 45 14 94 6430 16530 NOME 14 07
1875 45 14 95 6458 16310 WHITE MYT 05 09
1673 45 14 95 6458 16310 WHITE MT 05 09
1873 45 14 99 ©6500 15038 MANLEY HOT SPR 26 06
1873 45 14 99 ©6500 15038 MANLEY HOT SPR 26 06
14673 45 14 100 6453 14803 ESTER DOME 20 07
1873 45 14 101 6425 14435 GOODPASTOR R 08 07
1873 45 14 101 6409 14550 BIG DELTA 24 06
1873 45 14 104 6529 14525 EAGLE SUMMIT 18 07
1875 45 14 104 6523 14555 TWELVE MI SUMM 15 07
1873 45 14 104 6525 14520 EAGLE SUMMIT 30 06
1873 45 14 104 6530 14525 EAGLE SUMMIT 12 06
1873 45 14 104 6530 14525 EAGLE SUMMIT 12 06
1875 45 14 104 6530 14525 EAGLE SUMMIT 23 07
1873 45 14 104 6©530 14525 EAGLE SUMMIT 12 06
1873 45 14 104 6529 14525 EAGLE SUMMIT 07 07
1873 45 14 #104 6529 14525 EAGLE SUMMIT 23 06
1873 45 14 104 6531 14513 MILLER HOUSE 04 06
16735 45 14 104 6517 14629 SOURDOUGH CR 09 08
1873 45 14 104% 6517 14629 SOURDOUGH CR 09 08
1875 45 14 104 6531 14513 MILLER HOUSE 04 06
1873 45 14 104 6531 14513 MILLER HOUSE 04 06
1673 45 14 104 6530 14525 EAGLE SUMMIT 25 07
1673 45 14 104 6530 14525 EAGLE SUMMIT 2007
1673 45 14 105 ©6505 14730 PEDRO JOME 14 07
1873 45 14 105 6505 14730 PEDRO DOME 14 07
1873 45 14 105 6505 14730 PEDRO DOME 14 07
1873 45 14 105 6519 14818 TATALINA 28 05
1873 45 14% 105 6519 14818 TATALINA 28 05
1873 45 14 105 6503 14726 CLEARY SUMMIT 15 06
1873 45 14 105 6519 14818 TATALINA 28 0S
1873 45 14 105 6519 14818 TATALINA 28 05
218
939
927
967
967
96?
956
956
956
956
956
956
956
956
956
956
956
956
956
956
954
960
960
960
939
939
962
962
962
947
959
958
958
958
958
956
956
956
956
938
938
933
S35
933
B35
933
933
933
933
933
933
933
933
960
938
954
954
954
969
966
966
966
966
965
965
967
956
965
935
967
959
940
940
951
940
945
953
953
953
953
953
964
964
956
956
956
958
953
954
958
953
COLLECTOR NAMF
NNNNNADATTHNNNMNMNMMMMMMAKCCHUPFPTNUNDDNNDANNCMODOOVOOVCDOOOOOCCFFFFFrFrrFrrmMaAAADPrrPrPPADADOVDODOAOONOOOOOOrIrIVrwr >yS
UMD EZEZLEFTFLELTSEALZZTEEZVVIYvSevrvFerrvrvd
ah=
SU aes le
YP9PEZEErPPDMDINAD
DDa”?D
NELSON
PALMER
VIERECK
VIERECK
VIERECK
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
FROHME
MURTE
MURIE
MURIE
NELSON
NELSON
RICHEY
RICHEY
RICHEY
SCAMMAN
SCHENE
VIERECK
VIERECK
VIERECK
VIERECK
VIERECK
VIERECK
VIERECK
VIERECK
ANDERSON
ANDERSON
GEIST
GEIST
GEIST
GEIST
GEIST
GEIST
GEIST
GEIST
GEIST
GEIST
GEIST
GEIST
SAUER
ANDERSON
HELLER
HELLER
HELLER
PEGAU
WELSH
WELSH
PEGAU
PEGAU
WELSH
WELSH
VIERECK
JOHNSON
WELSH
ANDERSON
HARMS
OS FARRELL
SCAMMAN
SCAMMAN
SCAMMAN
SCAMMAN
SCAMMAN
SCAMMAN
SMITH
SMITH
SMITH
SMITH
SMITH
VIERECK
VIERECK
ARGUS
ARGUS
ARGUS
SMITH
SMITH
SMITH
SMITH
SMITH
COL NO HFRa + NO
S7AL
1940
AUIBA
R415
5718
647
645
635
627
648
627
579
631
674
632
6%6
637
679
674
54318
ALA
ALA
5WA
BWA
SWA
GWA
GWA
SWA
6WA
SWA
GWA
GWA
BWA
SWA
ALA
ALA
ALA
ALA
ALA
ALA
SWA
SWA
SWA
TSC
ALA
ALA
ALA
ALA
BWA
ALA
FSLC
BWA
GWA
GWA
GWA
ALA
ALA
ALA
TSC
TSC
ALA
ALA
ALA
ALA
ALA
ALA
ALA
ALA
ALA
ALA
ALA
ALA
wt
TSC
ALA
ALA
ALA
GWA
GWA
5WA
SWA
ALA
1SC
ALA
SWA
ALA
Isc
SC
ALA
ALA
ALA
SASK
GWA
GWA
5WA
5WA
ALA
ALA
SWA
GWA
GWA
FSLC
FSLC
ALA
ALA
SWA
ALA
ALA
ALA
GWA
GWA
495
5196
4551
4733
4729
4740
4727
21788
255951
496
26101
26108
26099
25317
11608
11626
11606
255831
255791
29381
29512
29455
29406
29407
29405
29478
29457
29456
29712
29770
29717
255926
26943
24329
24330
32700
246963
29999
27147
246878
255805
34633
9627
8397
34211
10614
10728
300
298
22444
4901
10615
10644
10729
Specimens Cited
TaxON SPC S V HYB PROV QUAD LAT
1673 45
1673 45
1873 45
1873 45
1873 45
1873 45
1873 45
1873 45
1873 45
1673 45
1873 45
1873 45
1873 45
1873 45
1673 45
1673 45
1873 45
1873 45
1873 45
1673 45
1873 45
1673 45
1873 45
1573 45
1873 45
1873 45
1873 45S
1473 45
1373 45
1873 45
1873 45
1873 45
1873 45
1873 45
1873 45
1873 45
1873 45
1873 45
1873 45
1873 45
1873 45
1473 45
1673 45
1673 45
1673 45
1873 45
1873 45
1873 45
1873 45
1873 45
1873 45
1873 45
1873 45
1673 45
1873 45
1873 45
1873 45
1873 45
1873 45
1673 45
1873 45
1873 45
1875 45
1873 45
1873 45
1873 45
1873 45
1873 45
1873 45
1873 45
1873 45
1873 45
1873 45
1873 45
1873 45
1873 45
1873 45
1873 45
1673 45
1873 45
1873 45
1873 45
1673 45
1873 45
1873 45
1873 45
1873 45
1873 4&5
1873 45
1873 45
1873 45
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
105
111
111
rae
111
111
1l1l
73
its
113
124
124
126
129
129
129
129
129
129
129
129
129
129
129
129
129
129
129
129
129
129
129
129
129
129
129
129
129
129
129
129
136
136
136
137
137
137
138
138
138
138
138
138
138
138
138
138
138
139
140
145
145
145
145
145
145
145
145
145
145
145
145
145
145
145
145
145
145
147
147
147
147
147
148
148
150
yay
151
151
151
151
6503
6533
6458
6458
60458
6459
6459
6604
6654
6655
6725
6725
6743
6821
6819
68e1
6806
6806
6806
6806
6806
6806
6815
6815
6815
6810
6810
6810
6810
6810
6810
6810
6810
6810
6810
6840
6840
6840
e842
6819
6819
6804
6804
6804
6822
6822
6822
6950
6950
6950
6950
6950
6950
6950
6950
6950
6950
6950
6924
6950
6902
6902
6902
6902
6902
6902
6902
6902
6902
0902
6902
6902
6902
6902
6902
6902
6902
6902
7030
7030
7030
7030
7031
7049
7049
7012
7010
7010
7010
7010
7010
LONG
14726
16751
16804
16804
16804
16801
16801
16242
16235
16240
15006
15007
15609
16647
16640
16647
16545
16545
16545
16545
16545
16545
16600
16600
16600
16540
16540
16540
16540
16540
16540
16540
16540
16540
16540
16537
16537
16637
16615
16640
16640
145035
14503
14503
14355
14355
14355
14220
14220
14220
14220
14220
14220
14220
14220
14220
14220
14220
14608
14815
16350
16350
16350
16350
16350
16350
16350
16350
16350
16350
16350
16350
16350
16350
16350
16350
16350
16350
15730
15730
15730
15730
15724
15442
15442
14716
14650
14650
14650
14650
14650
LOCALITY
CLEARY SUMMIT
TIN CITY
KING ISLAND
KING ISLAND
KING ISLAND
KING ISLAND
KING ISLAND
DEERING
KOTZEBUE
KOTZEBUE
WISEMAN
WISEMAN
ISIAK LAKE
POINT HOPE
POINT HOPE
POINT HOPE
OGOTORUK CREEK
OGOTORUK CREEK
OGOTORUK CREEK
OGOTORUK CREEK
OGOTORUK CREEK
OGOTORUK CREEK
CAPE THOMPSON
EBRULIKORUK CR
CAPE THOMPSON
OGOTORUK CR
OGOTORUK CR
OGOTORUK CR
OGOTORUK CR
OGOTORUK CR
OGOTORUK CR
OGOTORUK CR
OGOTORUK CR
OGOTORUK CR
OGOTORUK CREEK
KUKPUK RIVER
KUKPUK RIVER
KUKPUK RIVER
KIPALOG CREEK
POINT HOPE
POINT HOPE
OLD JOHN LAKE
OLD JOHN LAKE
OLD JOHN LAKE
SHEENJEK R
SHEENJEK R
SHEENJEK R
NUVAGAPAK
NUVAGAPAK
NUVAGAPAK
NUVAGAPAK
NUVAGAPAK
NUVAGAPAK
NUVAGAPAK
NUVAGAPAK
NUVAGAPAK
NUVAGAPAK PT
NUVAGAPAK PT
CANNING RIVER
SAGAVANIRKTOK
CAPE BEAUFORT
CAPE BEAUFORT
CAPE BEAUFORT
CAPE BEAUFORT
CAPE BEAUFORT
CAPE BEAUFORT
CAPE BEAUFORT
CAPE BEAUFORT
CAPE BEAUFORT
CAPE BEAUFORT
CAPE BEAUFORT
CAPE BEAUFORT
CAPE BEAUFORT
CAPE BEAUFORT
CAPE SEAUFORT
CAPE BEAUFORT
CAPE BEAUFORT
CAPE BEAUFORT
MEADE RIVER PO
MEADE RIVER PO
MEADE RIVER PO
MEADE RIVER PO
MEADE RIVER PO
TESHEKPUK
TESHEKPUK
SHAVIOVIK R
BULLEN
BULLEN
BULLEN
BULLEN
BULLEN
PT
PT
PT
PT
PT
PT
PT
PT
PT
954
938
938
938
959
968
969
938
951
966
939
962
960
938
938
938
966
966
966
966
966
966
960
960
960
959
959
959
959
959
959
959
959
959
961
964
964
963
960
960
966
957
954
954
956
956
956
966
966
966
966
966
966
966
966
966
966
966
947
958
966
966
966
966
966
966
966
966
966
966
966
966
966
966
966
966
966
966
966
966
966
960
947
947
947
966
966
966
966
COLLECTOR NAME
DOMODANODCTDDOVN YVNYNAIGYV AND VODVNAPNGYV VDAVDAYVAGVVNGVDYVVDGVACLE AOAANDAVGAVD GOANANGVNAVGOVOAAIWAONWNUMNYHNTFTPTFTITLIADY YS YS KF FPF KrF KPrTrrAaA*MNMOOQDCECCVACYUMCADWOACCCH
EESSESEPOHVNSF LCT LCSEL ESSE SCLC SES SELL LESESECSSE PSH CSE ETE EE
GuvvVu a
0m
ur
ge Lhe 2S LS Ses Se Se So oy
AAD rerwvDVIVA
SMITH 2363
ANDERSON 4BA1
ANDERSON 36078
ANDERSON 3697B
HARBO 8
PEGAU 3948
PEGAU 12769
ANDERSON 4808
SCAMMAN 6460
WELSH 5735
ANDERSON 5877
BROCKMAN 2620
LENT 15
ANDERSON 4602
ANDERSON 4601
ANDERSON 4577
ARGUS 5952
ARGUS 5049
ARGUS 5966
ARGUS 5969
ARGUS 5948
ARGUS 5965
BELSON SeNe
BELSON
BELSON
JOHNSON 5
JOHNSON 213
JOHNSON 615
JOHNSON 3R2
JOHNSON 213
JOHNSON 5
JOHNSON 213
JOHNSON 615
JOHNSON 3R2
JOHNSON 35
MELCHIOR 655
MELCHIOR 654
MELCHIOR 446
VIERECK 4170
VIERECK 4031
WELSH 5797
SHETLER AF161
SMITH 2563
SMITH 2563
KESSEL Su4
KESSEL Siy
KESSEL S17
ARGUS 5915
ARGUS 5887
ARGUS 5876
ARGUS 5920
ARGUS 5914
ARGUS 58R2
ARGUS 5917
ARGUS 5BA6
ARGUS 5918
ARGUS 5905
ARGUS 5829
SPETZMAN 328
THOMSON
ARGUS 5530
ARGUS 5589
ARGUS 5607
ARGUS 5647
ARGUS 5660
ARGUS 5524
ARGUS 5522
ARGUS 551
ARGUS 5571
ARGUS 5578
ARGUS 5528
ARGUS 5529
ARGUS 5561
ARGUS 5475
ARGUS 5666
ARGUS 5526
ARGUS 5527
ARGUS 5525
ARGUS 5256
ARGUS 5227
ARGUS 5257
ARGUS 5263
GEIST
BLACK SeNe
BLACK SeNe
SPETZMAN 2168
ARGUS 5732
ARGUS 5742
ARGUS 5736
ARGUS 5752
ARGUS 5758
COL NO HERB
SWA
ISC
Isc
GWA
ALA
SWA
SWA
Tsc
SWA
GWA
1sc
ALA
ALA
Tsc
Tsc
rsc
SWA
GWA
GWA
GWA
GWA
GWA
ALA
ALA
ALA
ALA
ALA
ALA
ALA
SASK
GWA
GWA
GWA
GWA
ALA
SWA
GWA
GWA
ALA
ALA
GWA
SWA
ALA
GWA
ALA
ALA
SWA
GWA
GWA
GWA
GWA
GWA
GWA
GWA
GWA
GWA
GWA
SWA
Tsc
wIs
GWA
GWA
GWA
GWA
GWA
GWA
GWA
GWA
GWA
GWA
GWA
GWA
GWA
SWA
GWA
SWA
GWA
GWA
GWA
GWA
GWA
GWA
ALA
Isc
TSsc
Tsc
GWA
SWA
GWA
GWA
GWA
+ NO
255794
24810
256587
255819
28481
29326
256405
255811
255932
28806
28797
28829
9174
9182
9175
9204
33104
9175
9204
17671
13231
13232
10738
10738
5051
5045
255802
27630
256552
255920
250234
219
Appendix B
TAXON SPC S V HYB PROV QUAD LAT
1873
1873
1473
1873
1873
1873
1873
1873
1873
1873
1873
1873
1873
1873
1873
46
1873
1673
1873
1873
1873
1873
1873
1873
1873
1873
18673
1873
1873
1873
1873
1873
1873
1873
1873
1873
1873
1873
1873
1873
1873
1873
1873
1673
1673
1873
1873
1873
1873
1873
1873
1873
1673
1873
1873
1673
1873
1873
1873
1873
1873
1673
1873
1673
1873
1873
1873
1873
1873
1673
1873
1673
1873
1873
1473
1473
1873
1673
1873
1673
1673
18673
1873
1873
1873
1873
1873
1673
220
45 14
45 14
45 14
45 14
45 14
45 14
45 14
45 14
45 14
45 14
45 14
45 14
4S 14
45 14
45 14
FUSCESCENS
4o iA
46 u
46 u
46 ti
46 i/
46 7
46 7
46 iz
46 v
46 if
46 7
46 i
46 12
46 12
46 12
46 14
46 i4
46 14
46 14
46 14
46 14
46 14
46 14
46 14
46 14
46 14
46 14
46 14
46 i4
46 14
46 14
46 14
46 14
46 14
46 14
46 14
46 14
46 14
46 14
46 14
46 14
46 14
46 14
46 14
46 14
46 14
46 14
46 14
46 14
46 14
46 14
46 14
46 14
46 14
46 14
46 14
46 14
46 14
46 14
46 14
46 14
46 14
46 14
46 14
46 14
46 14
46 14
46 14
46 14
46 14
46 14
46 14
151
151
151
151
151
151
151
151
151
151
152
152
152
152
152
7010
7010
7010
7010
7010
7010
7010
7010
7010
7010
7005
7005
7005
6839
6839
6932
6930
6942
6924
6813
6845
6839
6745
6745
6050
6050
6050
6825
6718
6451
5520
5705
5705
5705
5705
5732
5834
5858
5858
5852
5844
5842
5819
5819
5833
5947
5937
5940
5923
5904
5903
5903
5958
0025
6025
6024
6025
6025
6015
6009
6015
6015
6015
6015
6015
6024
6024
6015
6000
6023
6031
6033
6033
0033
6033
6010
6128
6156
6156
6113
6124
6124
6140
6140
6147
6147
6152
LONG
14650
14650
14650
14650
14650
14650
14650
14650
14650
14650
14335
14335
14335
14100
14100
13349
13347
12856
13257
13554
13412
13405
13601
10050
9425
9425
9425
13800
13748
13625
16248
15425
15425
15425
15425
15725
16146
15711
15711
15700
15701
15639
15406
15406
15547
15150
15333
15606
15610
15826
14826
15832
16658
17235
17235
17242
172355
17235
16648
16643
16614
16614
16614
16614
16614
16610
16610
16614
16600
15118
15046
15116
15116
15116
15116
14930
14922
14710
14710
14954
14929
14929
14900
14900
15010
15010
15433
LOCALITY
BULLEN
BULLEN
BULLEN
BULLEN
BULLEN
BULLEN
BULLEN
BULLEN
BULLEN
BULLEN
BARTER ISLAND
BARTER ISLAND
BARTER ISLAND
FIRTH RIVER
FIRTH RIVER
RICHARDS IS
RICHARDS IS
ANDERSON RIVER
TUKTOYAKTUK
CANOE Le
MACK R E BR
REINDEER ST
HORNE Le
AN
N
KANGOWAN LAKE
MCCONNELL R
MCCONNELL R
MCCONNELL R
SAM LAKE
BELL RIVER
DEMPSTR RD
COLD BAY
OLGA BAY
OLGA BAY
OLGA BAY
OLGA BAY
UGASHIK
CAPE PEIRCE
COPENHAGEN
COPENHAGEN
KOGGIUNG
NAKNEK
KING SALMON
KUKAK BAY
KUKAK BAY
BROOKS FALL
STERL HwY M
ILIAMNA BAY
IGIUGIG
ILIAMNA
SNAG POINT
KANAKANAK
KANAKANAK
DUCHIKMIUT
MATTHEW
MATTHEW
MATTHEW
MATTHEW
MATTHEW
NASH HdR
TWIN MT
MEKERYUK
MEKERY JK
MEKERYUK
MEKERYUK
MEKERYUK
CAPE ETOLIN
CAPE ETOLIN
MEKERYUK
NUNIVAK IS
KASILOF
MS7
CR
CR
S
159
STERL HWY M71
KENAI
KENAI
KENAI
KENAI
SEWARD
EKLUTNA
GLENN HWY M
GLENN HWY M
ANCHORAGE
BIRCHWOOD
BIRCHWOOD
MATANUSKA
MATANUSKA
128
128
FLAT
FLAT
VALL
VALL
WILLOW CREEK
WILLOW CREE
HEAD OF BIG
K
Re
DATE
08
08
08
08
08
08
08
08
08
08
08
08
08
06
06
966
966
966
966
966
966
966
966
966
966
966
966
966
961
961
947
966
965
965
963
934
965
962
966
967
967
967
970
964
966
971
938
940
938
938
967
970
959
959
953
952
952
B99
899
919
967
902
965
965
951
950
946
965
B99
B99
899
966
957
938
965
929
929
965
966
965
932
932
949
954
898
968
951
951
951
951
962
941
944
944
948
948
948
940
940
940
940
950
COLLECTOR NAME
MMADADADDADDANODNAND
a ee
2rrr er rm CCCrFUCCCYHEVONOADADZAZOCVCKAMAANBMWCAMCc STFHPYNUNFTEMPPCTFTOMMMMsEDVUNAAAUNCOPZOODD PS
n
uvrNAA 4 ral. ae P=EnNncEezEeeE
>
met
aT
Dv
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
LITTLEs
LITTLE?
JRe
JR.
PORSILD
SCOTTER
SCOTTER
SCOTTER
CODY
PORSILD
SCOTTER
CALDER
SVERRE
MACINNES
MACINNES
MACINNES
WELSH
YOUNGMAN
PORSILD
WILLIAMS
LOOFF
LOOFF
LOOFF
LOOFF
STREUSEL
DICK
MOSSMAN
MOSSMAN
MULLER
SCHOFIELD
SCHOFIELD
COVILLE
COVILLE
MILLER
VIERECK
GORMAN
HARMS
HARMS
MULLER
RAUP
STEWART
BOS
COVILLE
COVILLE
COVILLE
HARMS
KLEIN
ANDERSON
BOS
MILLER
MILLER
PEGAU
PEGAU
PEGAU
ROUSE
ROUSE
SPENCER
STETTENHEIM
EVANS
WELSH
CALDER
CALDER
CALDER
KLEIN
ROWINSKI
ANDERSON
ANDERSON
ANDERSON
DUTILLY
LEPAGE
LEPAGE
PALMER
PALMER
J PALMER
PALMER
DRURY
cOL NO
5737
5766
5733
5748
585
5809
5807
5806
5757
5753
5859
5A4U8
5846
18485
18484
HERB + NO
GWA
GWA
BWA
GWA
GWA
GWA
GWA
GWA
GWA
GWA
GWA
GWA
6WA
GWA
GWA
CAN 46460
GWA
GWA
GWA
NAO
5WA
GWA
DAO
CAN 307168
Uwo
Wo
Te)
OTF 1205
CAN 283830
CAN 303396
GWA
NA 319642
A
DAO 32786
NAO 32787
ALA 34728
SWA
ALA 23579
ALA 23585
TSC 224499
US 2331516
NY
US 373487
A
NS 1072534
6WA
US 420054
ALA 32608
ALA 32592
US 2176386
CAN
US 2232021
ALA 32290
NS 2440623
US 373490
US 373490
BWA
ALA 6600
TSC 255A84
ALA 32285
ALA 2770
ALA = 2023
5WA
GWA
ALA 32698
TSC 255879
TSC 255880
ISC 255894
MICH
'S 342369
GWA
US 2331432
NY
6H
TSC 255883
ALA 25631
TSC 255888
ISC 256013
SWA
ISC 255893
TSC 255892
ISC 255891
ALA 5986
ALA 5631
NA
NIA
CAN
Specimens Cited
TAxON SPC S V HYB PROV QUAD LAT LONG LOCALITY DATE
1873 46 14 71 6155 15425 HEAD OF BIG Re 10 7
1873 46 14 73 6137 15930 ANIAK 18 6
1873 46 14 73 6134 15934 ANIAK REGION 28 6
1873 46 14 73, 6134 15934 ANIAK REGION 28 6
1873 46 14 73 6135 15915 ANIAK = aT
1875 46 14 75 6155 16550 SCAMMON BAY 16 06
1875 46 14 75 6143 16608 KOKECHIK BAY 19 07
1873 46 14 75 6159 16558 IGIAK BAY 03 07
1875 46 14 75 6159 16558 IGIAK BAY 03 07
1875 46 14 77 6205 16343 MOUNTAIN VILGE
1673 46 14 79 6227 15800 FLAT IG 46
18673 46 14 80 6247 15544 KUSKOKWIM Re rat a
1875 46 14 81 6225 15125 LAKE CHELATNA 21 06
1873 46 14 81 6225 15125 LAKE CHELATNA 21 06
1875 46 14 83 6233 14526 RICH HwY M150 25 06
18753 46 14 83 6233 14526 RICH HWY M150 25 06
1673 46 14 86 6311 14533 ISABEL PASS 10 07
1873 46 14 86 6338 14443 HORN HOUNTAIN 06
1875 46 14 86 6342 14430 UPPER DRY CR 03 08
1873 46 14 86 6342 14430 UPPER DRY CR 03 08
18735 46 14 87 ©6350 14925 MCK PK RD M78 31 07
1875 46 14 87 6315 14915 SUMMIT 27 07
1873 46 14 87 6309 14926 SUMMIT 21 07
1875 46 14 88 6326 15040 MCKINLEY PARK 05 07
1673 46 14 88 6327 15050 WONDER L 29 07
1873 46 14 88 6327 15050 WONDER L 29 07
18735 46 14 89 6315 15520 NIXON FORK 12-6
1873 46 14 89 6315 15517 NIXON FORK 10" 6
18735 46 14 89 6315 15517 NIXON FORK 10 6
1875 46 14 89 6315 15517 NIXON FORK 10 6
16753 46 14 90 6310 15631 OPHIR 26 06
1873 46 14 91 6352 16047 UNALAKLEET 12 06
1873 46 14 91 6352 16047 UNALAKLEET 27 08
1873 46 14 91 6352 16047 UNALAKLEET 16 08
1875 46 14 91 6352 16047 UNALAKLEET 16 08
1875 46 14% 92 6329 16202 ST MICHAEL 19 06
1873 46 14 92 ©6329 16202 ST MICHAEL 08 06
1873 46 14 92 6329 16202 ST MICHAEL 08 06
18735 46 14 92 ©0329 16202 ST MICHAEL 08 06
1875 4o 14 92 6329 16202 ST MICHAEL 08 06
lo73 46 14 92 0302 16318 PASTOLIAK 05 07
1873 46 14 92 6302 16333 KOTLIK 10 07
1875 46 14 92 ©6302 16333 KOTLIK 10 07
1875 46 14 92 6302 16333 KOTLIK 07 07
18753 46 14 92 - 6302 16333 KOTLIK 07 07
1875 46 14 92 6302 16333 KOTLIK 07 07
1873 46 14 92 6302 16333 KOTLIK 07 07
1873 46 14 92 6302 16333 KOTLIK 07 07
1673 46 14 92 6302 163335 KOTLIK 07 07
18735 46 14 92 6335 162350 STUART IS 29 07
1873 46 14 92 6335 16230 STUART IS 20) O17
1875 46 14 93 6330 17135 ST LAWRENCE IS
146735 46 14% 93 6320 17136 BOXER BAY
18735 46 14 93 ©320 17136 BOXER BAY
1875 46 14 93 6320 17136 BOXER BAY
1873 46 14 93 6330 17030 ST LAWRENCE IS
1873 46 14 93 ©6302 16850 PUNUK ISLANDS
1875 46 14 94 6430 16530 NOME 18 06
14673 46 14 94 6430 16525 NOME
1875 46 14 94 6433 16535 NOME 14 07
13875 46 14 94 6440 16521 NOME 07 09
18735 46 14 94 6433 16535 NOME 14 07
1873 46 14 94 6433 16535 NOME 06 08
1875 46 14 94 ©6430 16525 NOME 01 07
1875 46 14 94 6430 16525 NOME 28 07
1873 46 14 94 6430 16525 NOME 21 06
1873 46 14 95 6433 16302 GOLOVIN 17 06
18673 46 14 96 6402 16055 EGAVIK 11 08
1673 46 14 96 6402 16055 EGAVIK 11 08
1873 46 14 96 6402 16055 EGAVIK 11 08
1873 46 14 96 6402 16055 EGAVIK 11 08
1673 46 14 96 ©6402 16055 EGAVIK 11 08
1873 46 14 96 6402 16055 EGAVIK 11 08
1873 46 14 100 6454 14751 GOLDSTREAM CR 24 06
1675 46 14° 100 6454 14751 GOLDSTREAM CR 24 06
1873 46 14 100 6452 14747 COLLEG= 05 07
1873 46 14° #100 6454 14751 GOLODSTREAM CR 24 06
1873 46 14° 100 6452 14747 COLLEGE 05 07
13875 46 14 100 6452 14747 COLLEGE 07 07
1873 46 14 #100 6452 14747 COLLEGE 07 07
1673 46 14 100 6448 14755 CHENA RIVER 21 06
1873 46 14 100 6452 14747 COLLEGE 06 06
18735 46 14 #101 6425 14520 BIG DELTA 03 07
1673 46 14 104 6527 14526 EAGLE CR CAMP 12 06
1875 46 14 110 6536 16313 IMURUK L 16 07
1873 46 14 111 ©6516 16622 TELLER 06 08
1873 46 14 111 6515 16630 PORT CLARENCE 02 08
1473 46 14 #113 6655 16231 KOTZEBUE 11 08
1873 46 14 113 6655 16231 KOTZEBJE 17 Of
1873 46 14 113 6655 16240 KOTZEBUJE 09 07
18673 46 14 128 6708 16345 CA KRUSENSTERN 12 07
950
949
949
949
949
961
952
960
960
949
949
956
956
947
947
935
957
957
957
953
967
967
956
956
956
950
950
950
950
940
938
894
920
920
938
931
931
931
931
923
926
926
931
931
931
931
931
eel!
931
931
933
933
933
Leis,
933
934
954
914
966
968
966
966
929
929
929
938
931
931
931
931
931
931
956
956
957
956
957
966
966
955
967
961
966
947
949
901
966
951
966
960
COLLECTOR NAME
TFreeZzezrzzumezezztzee
>rorroan
ZGUMUMMDAMECCADTFNAAYVGVAIGAVGAVGVIANANANNANCHNANADABDVNDONOOVCSOOSONNANNNNANANNYF PETIIICrrVFrncmezeetezerrrrrerprrree
LES
Prpdy
IrIriLIe-rresnr
iv]
VG G
ELZAZAZET LE Pte fe 12
DRURY 4222
DRURY 1516
DRURY 1666
DRURY 1656
DRURY 1735
HULTEN
HUMPHREY 13
JONES 70
JONES am
KYLLINGSTAN
DRURY 1780
DRURY 2160
VIERECK 1017
VIERECK 1017
DUTILLY 20435
DUTILLY 20435
ANDERSON i171
SPETZMAN
SPETZMAN 1019
SPETZMAN 1019
MURIE 1
VIERECK 8393
VIERECK R4NS
VIERECK 1195
VIERECK 1572
VIERECK 1572
DRURY 3728
DRURY 3A74
DRURY 3625
DRURY 3638
SCAMMAN 1832
ANDERSON 3265
FUNSTON 232
PALMER 74
PALMER 79
ANDERSON 3448
MASON 6043
MASON 6044
MASON 6044
MASON 6043
MILLER 4C
PORSILD BA7
PORSILD 867
ROUSE 33
ROUSE 33
ROUSE x2
ROUSE 32
ROUSE 36
ROUSE 3
ROUSE 23
ROUSE 23
GEIST SeNe
GEIST 1
GEIST 12
GEIST 14
GEIST SeNe
GEIST SeNe
HELLER 1107
HILL RY
PEGAU w9
PEGAU 24868
PEGAU w9
PEGAU WB
THORNTON 69
THORNTON 610
THORNTON 695
ANDERSON 34228
ROUSE 12
ROUSE 12
ROUSE Al
ROUSE al
ROUSE 20
ROUSE 20
ARGUS 4ug
ARGUS 440
ARGUS 1n°4
ARGUS 440
ARGUS 1094
ARGUS 5195
ARGUS 5194
SMITH 8
VIERECK R2n1
HERRICK
TRENT 966
SIEH 50
SCAMMAN 54460
WALPOLE 1674
ARGUS 5979
SCAMMAN 642
WELSH 5733
SHACKLETTE 6499
COL NO HF8B + NO
CAN
CAN
CAN
CAN
CAN
US 2384884
MICH
ALA 24424
ALA 24425
TSC 256222
CAN
CAN
ALA 11630
SWA
NA 329375
NA 329375
TSC 255896
US 2349423
ALA 06870
US 2384222
GWA
SWA
5WA
ALA 11627
ALA 3496
GWA
CAN
CAN
CAN
CAN
BWA
TSC 25586
NY
ALA 2038
ALA 2037
TSC 255885
US 1789423
US 1789424
5H
GH
ALA 2022
6H
CAN
ALA 2043
ALA 2047
ALA 2048
ALA 25453
ALA 2046
ALA 25443
ALA 2054
ALA 25457
ALA 29717
ALA 29454
ALA 29397
ALA 29399
ALA 29366
ALA 29365
ALA 24325
US 539161
ALA 32705
5WA
GWA
GWA
US 1438551
US 1438552
US 1438551
TSC 255881
ALA 2n49
ALA 25454
ALA 2039
ALA 25456
ALA 2079
ALA 25440
ALA 22445
ALA 4738
ALA 06800
SwWA
CWA
SWA
GWA
ALA 10747
GWA
ALA 18535
ALA 32691
TSC 255887
aH
JS 378786
GWA
5H
SWA
JS 2388069
221
Appendix B
TAXON SPC S V HYB PROV QUAD LAT LONG LOCALITY DATE
1873 46 14 128 6708 16345 CA KRUSENSTERN 12 07
1873 46 14 129 6810 16540 OGOTORUK CR 31 07
1873 46 14 129 6810 16540 OGOTORUK CR 2on0i7
1873 46 14 129 6810 16540 OGOTORJK CR 12 08
1873 46 14 129 6810 16540 OGOTORUK CR 22 06
1873 46 14 129 6810 16540 OGOTORUK CR 30 06
1873 46 14 129 6810 16540 OGOTORUK CR 22 07
1873 46 14 129 6810 16540 OGOTORUK CR 31 07
1873 46 14 129 6810 16540 OGOTORUK CR 24 06
1873 46 14% 129 6810 16540 OGOTORUK CR me Be ONY
1873 46 14 129 6810 16540 OGOTORUK CR 25 407
1873 46 14 129 6810 16540 OGOTORUK CR 12 08
1873 46 14 129 6810 16540 OGOTORUK CR 22 06
1873 46 14% 129 6810 16540 OGOTORUK CR 30 06
18673 46 14 129 6810 16540 OGOTORUK CR 24 06
1873 46 14 129 6810 16540 OGOTORUK CR 24 06
1873 46 14 129 6810 16540 OGOTORUK CR 22 07
1873 4o 14 129 6810 16540 OGOTORUK CR Si. 07
1873 46 14 129 6810 16540 OGOTORUK CR 16 08
1873 46 14 130 6852 16430 PITMEGEA R 27 06
1873 46 14 131 6847 16000 LAKE NOLUK 26 07
1873 46 14 131 6847 16000 NOLUCK LAKE 26 07
1873 46 14 133 6822 15439 KURUPA LAKE 14 07
1873 46 14 136 6804 14503 OLD JOHN LAKE 08 08
1873 46 14 138 6950 14220 NUVAGAPAK PT 09 08
1873 46 14 138 6950 14220 NUVAGAPAK PT 09 08
1873 46 14 138 6950 14220 NUVAGAPAK PT 09 OB
1873 46 14 138 6950 14220 NUVAGAPAK PT 09 08
1873 46 14 138 6950 14220 NUVAGAPAK PT 09 08
1873 46 14 138 6950 14220 NUVAGAPAK PT 09 08
1873 46 14 $4138 6950 14220 NUVAGAPAK PT 09 08
1873 46 14% 139 6935 14445 SADLEROCHIT R 17 08
1873 46 14 141 6930 15200 UMIAT 10 08
1873 46 14 141 6923 15210 UMIAT 28 07
1873 46 14 141 6923 15210 UMIAT 28 07
1873 46 14 145 6902 16350 CAPE BEAUFORT 23 07
1873 46 14 145 6902 16350 CAPE BEAUFORT 23 07
1873 46 14 145 6902 16350 CAPE BEAUFORT 23 07
1873 46 14 #4145 6902 16350 CAPE BEAUFORT 23 07
1873 46 14° #145 6902 16350 CAPE BEAUFORT 23 07
1873 46 14 145 6902 16350 CAPE BEAUFORT 23 07
1873 46 14 145 6902 16350 CAPE BEAUFORT 23 07
1873 46 14 145 6902 16350 CAPE BEAUFORT 23 07
1873 46 14 146 7038 16002 WAINWRIGHT 04 08
1873 46 14 147 7030 15730 MEADE RIVER PO 16 07
1873 46 14 147 7030 15730 MEADE RIVER PO 16 07
1873 46 14 147 47030 15730 MEADE RIVER PO 16 07
1873 46 14 147 #=+:.(7030 15730 MEADE RIVER PO 16 07
1873 46 14 147 7030 15730 MEADE RIVER PO 16 07
1873 46 14 147 7030 15730 MEADE RIVER PO 16 07
1873 46 14 147 =%7030 15730 MEADE RIVER PO 16 07
1673 46 14 147 7030 15730 MEADE RIVER PO 16 07
1873 46 14 147 +7030 15730 MEADE RIVER PO 16 07
1873 46 14 147 7030 15730 MEADE RIVER PO 16 07
1573 46 14 147 #=+.‘(7030 15730 MEADE RIVER PO 16 07
1673 46 14% 147 #7030 15730 MEADE RIVER PO 16 07
1873 46 14 147 +7048 15815 PEARL BAY 05 08
1873 46 144 152 7005 14335 BARTER ISLAND 07 08
1873 46 14 152 7005 14335 BARTER ISLAND 07 08
1873 46 14 152 7005 14335 BARTER ISLAND 07 OB
1873 46 14 $4153 7123 15628 POINT BARROW 22 06
1873 46 14 153 7119 15643 BARROW 31 2oi7
1873 46 14 153 7119 15643 BARROW 31 407,
49 1 GLAUCA VARe GLAUCA
1673 BS) 0 7 133 6931 13348 RICHARDS IS 10 07
1873 49 O11 7 375 6103 11004 THEKULTHILI L 11 08
1873 49 01 14 1i 5822 13600 GLACIER BAY 29 08
1673 49 01 14 11 5822 13600 GLACIER BAY 14 07
1873 49 01 14 11 5822 13600 GLACIER BAY 10 07
1873 49 01 14 28 5520 16030 POPOF ISLAND 19 07
1873 49 O11 14 34 5746 15312 TERROR BAY 16 08
1873 49 01 14 36 5747 15608 ISLAND LAKE 11 08
1673 49 O11 14 36 5732 15725 UGASHIK 250007
1873 49 1 14 39 5834 16146 CAPE PEIRCE 506
1873 49 1 14 39 5834 16146 CAPE PEIRCE 260 7
1673 49 01 14 41 5852 15703 KVICHAK RIVER 08
1673 49 01 14 43 5804 15304 PORT VITA 25) 07
1873 49 01 14 43 5804 15304 PORT VITA T2017
1673 49 01 14 52 5921 15737 BECHAROF LAKE 09 08
1873 eb a 14 67 6141 14148 RUSSELL GL 24 07
1873 49 01 14 67 6139 14218 SKOLAI VALLEY 11 07
1873 49 01 14 67 6140 14220 SKOLAI RIVER 20 06
1673 49 01 14 69 6155 14715 TAHNETA PASS 30 06
1673 49 01 14 69 6140 14900 MATANUSKA VY
1673 49 1 14 71 6155 15425 HEAD OF BIG Re 10 7
1873 49 1 14 73 6135 15915 RUSSIAN MISS. ne
1873 49 1 14 73 «©6137 15930 ANIAK 18 6
18673 49 1 14 73 6137 15930 ANIAK 18 6
222
960
959
959
959
959
959
959
959
959
959
959
959
959
959
959
959
959
959
960
S5i7
950
950
952
957
966
966
966
966
966
966
966
948
949
966
966
966
966
966
966
966
966
966
966
938
966
966
966
966
966
966
966
966
966
966
966
966
947
966
966
966
88
950
950
957
962
921
929
929
941
950
949
967
970
970
936
945
946
949
967
967
967
967
940
950
940
949
949
COLLECTOR NAME
oc DMOAODDAAVNADNANANAVN AVN ANANAVNAADNCHOHAADANVDNADAGDA<X<XTTAVDADODNDGODGVDOUYYSYOTFOTFCrHEPF SFE HTP YP KrrrPrrYrPPrPrrrKr.t
ZEZEETrFeDAAAZEOOVFTreVAZTTrzeseEeOE
SPwPteetezteazze DD PReNPESEEETERAEEZESA EE LES TS
Za NZ STE LEEZTELZELZE
=
SHACKLETTE
JOHNSON
JOHNSON
JOHNSON
JOHNSON
JOHNSON
JOHNSON
JOHNSON
JOHNSON
JOHNSON
JOHNSON
JOHNSON
JOHNSON
JOHNSON
JOHNSON
JOHNSON
JOHNSON
JOHNSON
VIERECK
CANTLON
SPETZMAN
SPETZMAN
HODGDON
SHETLER
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
SPETZMAN
SPETZMAN
SUDA
SUDA
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ANDERSON
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
AGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
BLACK
ARGUS
ARGUS
ARGUS
MURDOCK
THOMAS
THOMAS
Copy
SCOTTER
COOPER
COOPER
COOPER
COLE
CLARK
RAUP
STREUBEL
DICK
DICK
JONES
EYERDAM
EYERDAM
RAuP
SCOTT
SCOTT
SCOTT
HARMS
PALMER
DRURY
DRURY
DRURY
DRURY
COL NO
6503
585
451
648
98
236
4us
587
153
5a5
4Si
648
98
236
153
153
445
587
4596
57303
427B
4278
R4IT7
11097
58R5
5922
5BR4
5923
58a1
5883
58R8
1075
2635
26866
5430
5431
5432
5427
5428
5429
5394
5395
4356
5298
5293
5205
52R6
52Aa9
5296
52A8
5202
5291
5204
52Aa7
5290
57F
5867
5866
5a71
29
2134
2124
9988
3049
SeNe
SeNe
on7
GS eNeo
109
362
9257
SeNe
SeNe
140
1823
1714
1610
60°2
418
4144
1739
1493
1492
HFRS + NO
MICH
GWA
GWA
GWA
GWA
GWA
GWA
GWA
GWA
ALA
ALA
ALA
ALA
ALA
ALA
ALA
ALA
ALA
ALA
6H
NA 319188
NA 319188
6H
GWA
SWA
GWA
GWA
GWA
GWA
GWA
SWA
US 2032295
US 2032772
GWA
GWA
GWA
GWA
GWA
GWA
GWA
GWA
GWA
GWA
TSc
SWA
GWA
GWA
SWA
GWA
GWA
BWA
SWA
GWA
GWA
GWA
GWA
1sc
5WA
GWA
SWA
US
US 2312570
GH
9176
9172
9169
9196
9200
9195
16269
9202
9170
13239
255889
255890
32690
275156
QAO
CAN
MIN
MIN
MIN
Isc
Isc
LSC
ALA
GWA
GWA
tse
TSs¢
woe
ysc
SWA
GWA
GWA
ALA
ALA
CAN
CAN
CAN
CAN
256583
256402
219355
34730
146447
256404
256403
219354
34634
5219
Specimens Cited
TAXON SPC S V HYB PROV QUAND LAT LONG LOCALITY DATE COLLECTOR NAME COL NO HE&B + NO
18673 49 «#1 14-73 «6134 15934 ANIAK 18 6 949 WH DRURY 1599 CAN
1873 49° 1 14-73 «+6137 15930 ANIAK 18 6 949 WH DRURY 1508 CAN
1873 49 01 14 91 6352 16047 UNALAKLEET 11 08 925 L J PALMER 1247 ALA 2031
1873 49:01 14 92 6302 16318 PASTOLIK 19 07 931 C H ROUSE 30 GWA
1873 49 01 14 92 6302 16318 PASTOLIK 19 07 931 C H ROUSE 30 ALA 2040
1873 49:01 14 92 6302 16318 PASTOLIK 19 07 931 C H ROUSE 27 ALA 29318
1873 49:01 14 94 6430 16530 NOME 24 08 938 J P ANDERSON 5002 TSC 256535
1a73 4901 14 94 6430 16530 NOME 23 06 929 wR MILLER 112C GWA
1873 4901 14 94 6434 16533 NOME 02 07 968 R PEGAU 19868 GWA
1873 4901 14 94 6435 16538 NOME 21 08 966 R_ PEGAU wi0 SWA
1873 49:01 14 94 6435 16538 NOME 16 06 966 R- PEGAU W4 GWA
1673 4901 14 94 6435 16520 DEXTER CREEK 07 08 965 R PEGAU wi9 GWA
1873 4901 14 94 6435 16538 NOME 21 08 966 R- PEGAU Wi0 ALA 32704
1873 4901 14 95 6433 16302 GOLOVIN 17 06 938 J © ANDERSON 3424 ISC 256586
1873 4901 14 96 6402 16055 EGAVIK 21 08 929 wR MILLER 245C ALA 2021
1873 49 01 14 111 6516 16622 TELLER 24 08 938 J P ANDERSON 3576 ISC 256413
1873 49 01 14 111 6511 16533 DUCK CREEK 18 07 968 R- PEGAU 27368 GWA
1873 49 01 14 113 6655 16231 KOTZEBUE 11 08 966 G W ARGUS 5975 GWA
1873 49:01 14 113 6655 16231 KOTZEBUE 11 08 966 G W ARGUS 5974 WA
1873 49:01 14 113 6655 16231 KOTZEBUE 11 08 966 G W ARGUS 5978 GWA
1673 49 01 14 113 6655 16240 KOTZEBUE 09 07 966 S L WELSH 5757 SWA
1873 4901 14 113 6655 16240 KOTZEBUE 09 07 966 S L WELSH 5734 GWA
18673 49 01 14 114 6617 16153 CHORIS PEN 15 09 931 C H ROUSE 10 ALA 25439
1873 4901 14 114 6617 16153 CHORIS PEN 15 09 931 C H ROUSE 10 ALA 2050
1873 49 01 14 120 6709 14140 PORCUPINE R 12 08 961 F © DEAN 21 ALA
1873 4901 14 126 6706 15815 ONION PORTAGE 26 07 967 C SCHWEGER 154 ALA 34719
1673 4901 14 128 6744 16432 KIVALINA 20 06 960 A BUCKNELL 8 ALA 26587
1873 4901 14 129 6853 16613 CAPE LISBURNE 07 08 938 J P ANDERSON 4498 TSC 256555
1673 4901 14 129 6806 16545 OGOTORUK CREEK 11 08 966 G W ARGUS 5959 6WA
1873 49:01 14 129 6806 16545 OGOTORUK CREEK 11 08 966 G W ARGUS 5950 SWA
1873 4901 14 129 6815 16600 CAPE THOMPSON 10 07 960 L 4 BELSON 87B ALA 28687
1873 49:01 14 129 6808 16540 OGOTORUK CREEK 20 06 959 A W JOHNSON 41 GWA
1873 49 01 14 129 6808 16540 OGOTORUK CREEK 25 06 959 A W JOHNSON 168 GWA
1873 49:01 14 129 6808 16540 OGOTORUK CREEK 25 06 959 A W JOHNSON 170 GWA
1873 4901 14 129 6808 16540 OGOTORUK CREEK 05 08 959 A W JOHNSON 610 GWA
1873 4901 14 129 ©6808 16540 OGOTORUK CREEK 05 07 959 A W JOHNSON 303 GWA
1673 49 01 14 129 6808 16540 OGOTORUK CREEK 11 08 959 A W JOHNSON 643 GWA
1873 4901 14 129 6806 16545 OGOTORUK CREEK 05 07 959 A W JOHNSON 303 GWA
1873 490 1 14 129 6808 16540 OGOTORUK CREEK 11 08 959 A W JOHNSON 644 ALA 16270
1873 4901 14 129 6808 16540 OGOTORUK CREEK 20 06 959 A W JOHNSON 41 ALA 9197
1873 49: 01 14 129 6808 16540 OGOTORUK CREEK 05 07 959 A W JOHNSON 303 ALA 9194
1873 49 01 14 129 ©6808 16540 OGOTORUK CREEK 25 06 959 A W JOHNSON 168 ALA 9190
1673 49 01 14 129 ©6808 16540 OGOTORUK CREEK 25 06 959 A W JOHNSON 170 ALA 9189
1873 49 01 14 129 6808 16540 OGOTORUK CREEK 11 08 959 A W JOHNSON 645 ALA 162714
e739 01 14 129 ©6808 16540 OGOTORUK CREEK 05 08 959 A W JOHNSON 610 ALA 9192
1873 49 0 1 14 129 ©6808 16540 OGOTORUK CREEK 20 06 959 A Ww JOHNSON 41 ALA 26694
1873 4901 14 129 6808 16540 OGOTORUK CREEK 05 08 959 A w JOHNSON 610 ALA 26715
1673 49:01 14 129 ©6808 16540 OGOTORUK CREEK 13 06 961 K JONES 5 ALA 25076
18673 49 01 14 129 6816 16535 KUKPUK RIVER 04 08 963 H ® MELCHIOR 549 GWA
1873 4901 14% 129 0816 16535 KUKPUK RIVER 04 08 963 H P MELCHIOR 550 SWA
1673 49 01 14 129 6816 16535 KUKPUK RIVER 04 08 963 H P MELCHIOR 548 GWA
1873 4901 14 129 6816 16535 KUKPUK RIVER 16 07 963 H P MELCHIOR 544 GWA
1873 4901 14 129 ©6817 16532 ANGMAKROG MT 25 07 960 H ® MELCHIOR 168 ALA 17811
1873 4901 14 129 6845 16600 UKINYAK CREEK 30 07 960 L A VIERECK 44un& ALA 13230
1873 4901 14 129 6845 16600 UKINYAK CREEK 03 08 960 L A VIERECK 4499 ALA 13228
1473 4901 14 129 6815 16528 KUKPUK RIVER 22 08 960 L A VIERECK ALA 13237
1873 4901 14 129 6842 16615 CAPE DYER 22 07 960 L A VIERECK 4256 ALA 13229
1673 4901 14 136 6810 14530 ARCTIC VILLAGE 24 06 965 V L HARMS 3795 ALA 32567
1673 49 01 14 137 ©6822 14355 SHEENJEK R 23 06 956 B- KESSEL S66B ALA 5053
1873 4901 14 137 6840 14345 SHEENJEK R 1107968 KESSEL S169D ALA 3678
1873 49:01 14 141 6923 15210 UMIAT 29 07 966 Y SUDA 27166 GWA
1673 49 01 14 141 6923 15210 UMIAT 29 07 966 Y SUDA 27455 GWA
1873 4901 14 141 ©6923 15210 UMIAT 29 07 966 Y SUDA 27766 GWA
1673 49:01 14 141 6923 15210 UMIAT 29 07 966 Y SUDA 27266 GWA
18673 4901 14 142 6952 15350 IKPIKPUK 04 09 947 R © BLACK SeNe TSC 256553
1873 4901 14 142 6952 15350 IKPIKPUK 04 09 947 R © BLACK S.Ne TSC 256551
1873 4901 14 142 ©6948 15350 IKPIKPUK 30 08 947 R © BLACK TSC 256563
1673 49 01 14 142 6942 15450 IKPIKPUK R 10 07 959 0 wW GEIST A ALA 29310
1673 4901 14 143 6953 15708 MEADE RIVER 22 07 960 O W GEIST SeNe ALA 27726
1873 49 01 14 143 6956 15957 KAOLAK R 09 07 958 S_ SHUSHAN WIS
1873 4901 14 145 6946 16303 POINT LAY 06 08 938 J © ANDERSON 4469 TSC 256554
1873 4901 14 145 6902 16350 CAPE BEAUFORT 25 07 966 G W ARGUS 5554 GWA
1873 4901 14 145 ©6902 16350 CAPE BEAUFORT 24 07 966 G w ARGUS 5435 GWA
1673 4901 14 145 6902 16350 CAPE BEAUFORT 24 07 966 G W ARGUS 5520 GWA
1373 49 01 14° 145 6902 16350 CAPE BEAUFORT 24 07 966 G W ARGUS 5537 GWA
1873 49:01 14 145 6902 16350 CAPE BEAUFORT 24 07 966 G W ARGUS 5492 GWA
1673 49:01 14 145 6902 16350 CAPE BEAUFORT 24 07 966 G W ARGUS 5474 GWA
1873 4901 14 145 6902 16350 CAPE BEAUFORT 24 07 966 G W ARGUS 5476 GWA
1873 49 01 14 145 6902 16350 CAPE BEAUFORT 25 07 966 G W ARGUS 5555 GWA
1873 4901 14 145 6902 16350 CAPE BEAUFORT 27 07 966 G W ARGUS 5664 GWA
1873 49:01 14 145 6902 16350 CAPE BEAUFORT 28 07 966 G W ARGUS 5603 GWA
1873 4901 14 145 6902 16350 CAPE BEAUFORT 27 07 966 G W ARGUS 5662 GWA
18673 4901 14 145 ©6902 16350 CAPE BEAUFORT 25 07 966 G W ARGUS 5575 WA
1873 4901 14 145 6902 16350 CAPE BEAUFORT 27 07 966 G W ARGUS 5661 GWA
1673 4901 14 145 6902 16350 CAPE BEAUFORT 28 07 966 G W ARGUS 5698 GWA
1673 4901 14 145 6902 16350 CAPE BEAUFORT 24 07 966 G W ARGUS 5436 GWA
1873 49 01 14 145 6902 16350 CAPE BEAUFORT 24 07 966 G W ARGUS 5427 GWA
1873 4901 14 145 6902 16350 CAPE BEAUFORT 25 07 966 G W ARGUS 5559 GWA
1873 4901 14 145 6902 16350 CAPE BEAUFORT 24 07 966 G W ARGUS 5428 GWA
1873 49 01 14 145 6912 16201 KOKOLIK R 30 08 947 R © BLACK SeNe TSC 256689
1873. 49:01 14 145 6912 16201 KOKOLIX R 30 08 947 R © BLACK SeNe TSC 256590
223
Appendix B
TAXON SPC S V HYB PROV QUAD LAT LONG LOCALITY DATE
1873 89-01 14 146 7008 15945 AVALIK R 17 08
1873 49 01 14 146 7008 15945 AVALIK R 17 08
18735 49 01 14 147 7030 15730 MEADE RIVER LA Or.
1875 49 01 14 147 7030 15730 MEADE RIVER 14 07
1873 49 01 14 147 7030 15730 MEADE RIVER 17, O07
1873 79.0: J 14 147 7030 15730 MEADE RIVER 15 07
1873 49 01 144 147 7030 15730 MEADE RIVER i? O7
1873 49 O01 14 147 7030 15730 MEADE RIVER 14 07
1873 49 01 14 147 7030 15730 MEADE RIVER 14 07
1873 49 1 14 147 7030 15730 MEADE RePeO- iS: a7
1873 49 01 14 147 7030 15730 MEADE RIVER M7 Or
1873 oe ae | 14 147 7030 15730 MEADE RIVER 14 07
18735 wa,0 2 14° #147 7030 15730 MEADE RIVER 14 07
1873 49 01 14° #147 7030 15730 MEADE RIVER 14 07
1873 $9. 6..1 14 147 7030 15730 MEADE RIVER 15 07
18735 49 01 14 147 7030 15730 MEADE RIVER 14 07
1873 49 O01 14 147 7030 15730 MEADE RIVER 14 07
1873 49 01 14 147 7030 15730 MEADE RIVER 14 07
1673 a9.,0' 52 14 147 7030 15730 MEADE RIVER 14 07
1873 49 01 14 147 7031 15722 USUKTUK RIVER 07 07
1873 49 0 1 14 147 7028 15724 MEADE RIVER 08 08
1873 49 01 14 147 7031 15724 MEADE RIVER 17 08
1873 49 01 14 147 7031 15724 MEADE RIVER 17 08
1875 eS | 14 147 7030 15730 MEADE RIVER 14 07
1873 29) 0) 1 14 147 7030 15730 MEADE RIVER ate Oi
1875 49 01 14 147 7030 15730 MEADE RIVER atm, OW
1875 49 0 1 14 147 7030 15730 MEADE RIVER 14 07
1873 49 0 1 14 147 7030 15730 MEADE RIVER 17 07
1873 49 01 14 147 7030 15730 MEADE RIVER 15 07
1873 #904 14 147 7030 15730 MEADE RIVER 14 07
1873 $9- Od 14 147 7030 15730 MEADE RIVER 14 07
1873 49 01 14 147 7030 15730 MEADE RIVER 14 07
49 2 GLAUCA VARe ACUTIFOLIA
18753 49 2 2 1 5949 13638 HAINES RD MB2 O04 08
1873 49 2 2 1 5935 13627 THREEGUARDSMAN 04 08
18735 49 2 2 2 5935 13630 INSPECTOR CREE 12 7
1875 49 2 2 2 5941 13632 NADAHINI RIVER 13 7
1675 49 2 7 152 6820 13330 INUVIK 29h 26
1873 49 2 7 338 6300 12902 0 GRADY LAKE 25 07
18735 49 2 7 345 6225 11418 YELLOWKNIFE 23 08
1873 49 2 7 368 6118 12405 Se NAHANNI Re 26 6
1675 49 2 12 3 6924 14050 MT PAGE Die
1875 49 2 12 4 6812 13845 CROW FLATS 24 66
18673 49 2 12 4 6809 13949 SAM LAKE 15) a7
1873 49 2 12 4 6822 13910 SAM LAKE ay
1873 49 2 12 4 6826 13840 BARN MTS 30 6
1873 49 2 12 4 6840 13907 BABBAGE RIVER Ld
1873 49 2 12 4 6906 13807 KING POINT Siar
1673 49 02 12 5 6725 14100 RAMPART HOUSE 17 08
1873 49 02 12 5S ©6725 14100 RAMPART HOUSE 17 08
13875 49 0 2 12 5 6725 14100 RAMPART HOUSE 11 06
1673 49 0 2 12 15 6404 13925 DAWSON
1875 49 02 12 15 6408 13951 SIXTYML RD M12 26 06
1873 eo) 0-2 12 21 6350 13731 BARLOW 05 06
1875 $9. 0° 2 12 21 6352 13857 KING SOL DOME 03 07
1873 49 0 2 12 21 ©6325 13640 STEWART RIVER 29 07
1873 49 02 12 22 6335 13554 MAYO 06 08
18753 49 02 12 27 6205 13618 CARMACKS 21 08
1873 49 02 12 32 ©0159 14033 WHITE RIVER 21 07
14875 59:0 2 12 32 6159 14033 WHITE RIVER 21 07
1873 49 0 2 12 32 ©6120 13910 DUKE RIVER 21 08
1873 49 0 2 12 32 6120 13910 DUKE RIVER 21 08
1873 49 02 12 32 6120 13910 DUKE RIVER 21 08
1675 49 02 12 32 6102 138353 MT WALLACE 20 08
1673 49 0 2 12 32 6144 13832 PTARMIGAN HRT 13 07
1873 49 0 2 12 32 6146 13837 PTARMIGAN HRT 15 07
1673 49 0 2 12 32 ©6133 13840 KLUANE L 20 08
1675 49 02 12 32 6103 13831 KLUANE LAKE 12 07
1875 29.02 12 32 ©6103 13831 KLUANE LAKE 02 07
1873 49 0 2 12 32 6122 13859 BURWASH 02 07
1873 49 0 2 12 32 6115 13844 DESTRUCTION B 07 08
1673 49 0 2 12 32 6110 13845 KLUANE LAKE 30 06
1675 49 0 2 12 32 0129 13916 ALA HWY M1110 28 07
1873 49 02 12 34 6111 13512 LAKE LABERGE
18753 49 0 2 12 34 6111 13512 LAKE LABERGE
1875 49 02 12 34 6111 13512 LAKE LABERGE
1875 49 0 2 12 34 6111 13512 LAKE LABERGE
1675 49 02 12 35 6155 13228 LOWER LAPIE R 15 06
1673 49 0 2 12 35 ©155 13238 LOWER LAPIE R 15 06
1675 49 02 12 35 6154 13245 LAPIE R 27 06
1675 49 U 2 12 39 6059 13829 KLUANE LAKE 23 07
1673 49.0 2 12 39 6059 13829 KLUANE LAKE 24 07
1875 49 0 2 12 39 6057 13825 SLIMS RIVER 18 07
18673 49 0 2 12 40 6045 13615 MENDENHALL R 30 06
1675 49 02 12 40 0047 13735 PINE CREEK 10 06
1673 49 0 2 12 40 6047 13735 PINE CREEK 17 06
1673 49 0 2 12 4Q 6047 13735 PINE CREEK 17 06
02 12
1673 49 40 6047 13735 PINE CREEK 17 06
224
959
959
966
966
966
966
966
966
966
966
966
966
966
966
966
966
966
966
966
960
960
960
960
966
966
966
966
966
966
966
966
966
967
967
967
967
970
967
927
970
970
970
970
970
970
970
970
954
954
951
932
966
960
949
949
944
941
944
944
966
966
966
944
94R
948
944
944
944
948
94e
966
966
932
932
932
932
944
944
944
944
944
944
966
948
944
944
944
ZELLER EETE CAE Settee ae set
<~<<~<<<<<<=<XOOCOCDMNANANVNVDNAANAGVNANGDPDNANDANANGODGDVCO
VSS. OSCE ses scr == ps
©
WREST TTALTA TAC crceqce wv OCCNHNNNNNHCOCNODTEYNOOL TC
e£tetzvso
TIrIrI|9rTtcectcrrrp
_s'‘S= 4°00 1
COLLECTOR NAME
GEIST
GEIST
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
GEIST
GEIST
GEIST
GEIST
SUDA
SUDA
SUDA
SUDA
SUDA
SUDA
SUDA
SUDA
SUDA
VIERECK
VIERECK
ARGUS
ARGUS
WELSH
CODY
RAUP
SCOTTER
WELSH
RIGBY
WELSH
WELSH
WELSH
WELSH
WELSH
BUCKLEY
BUCKLEY
LOAN
BEAUCHAMP
WELSH
CALDER
CALDER
GILLETT
ANDERSON
COLE
ANDERSON
ANDERSON
ARGUS
ARGUS
ARGUS
RAUP
RAUP
RAUP
RAuUP
RAUP
RAUP
RAuUP
RAuUP
SUDA
WELSH
BEAUCHAMP
BEAUCHAMP
BEAUCHAMP
BEAUCHAMP
PORSILD
PORSILD
PORSILD
ANDERSON
ANDERSON
RAUP
ARGUS
RAUP
RAUP
RAUP
RAUP
COL NO
SeNe
SeNe
5313
5191
5310
5268
5314
5197
5201
S276A
SS it
5204
5207
52n3
5236
5193
5194
5198
5205
SeNe
SeNe
12
5
20266
21566
22066
19566
21666
21066
2166
19966
19866
8552
A531
6727
6810
10014
16477
503
12374
10271
55
10664
10510
10038
101964
10694
SeNe
SeNe
427
5584
25058A
3465
4195
9716
SNe
9313
9312
6006
6009
6011
128R8
13594
13677
128f8
12431
12185
13319
139014
16866
6055
ine
SeNe
9512
9511
10012
9378
9412
12595
5077
13067
117°7
11783
11795
HERB + NO
ALA 28138
ALA 27995
GWA
BWA
GWA
GWA
GWA
GWA
GWA
GWA
GWA
6WA
GWA
GWA
GWA
GWA
GWA
GWA
GWA
GWA
ALA 28950
ALA 27855
ALA 27849
BWA
GWA
BWA
GWA
GWA
GWA
GWA
GWA
GWA
GWA
GWA
6WA
GWA
OTF 1206
NAO 32665
GWA
GWA
OTF 1201
OTF 1186
OTF 1213
OTF 1203
OTF 1202
OTF 1187
OTF 1214
SWA
ALA 5023
BWA
ALA 523
GWA
GWA
TSC 256565
ISC 256564
TSC 256592
ISC 256582
TSC 256576
TSC 256577
BWA
GWA
GWA
GWA
ALA 19476
ALA 19475
ALA 19765
ALA 19759
ALA 20043
ALA 19471
ALA 19386
6WA
GWA
ALA 519
ALA 525
ALA 520
ALA 518
TSC 256678
1SC 256679
TSC 256680
ISC 256571
ISC 256570
ALA 19899
GWA
ALA 19469
ALA 19473
ALA 19476
ALA 20026
Specimens Cited
TAxON SPC
1873 49
1875 49
1873 49
1873 49
1873 49
1873 49
1873 49
1873 49
1873 49
1873 49
14673 49
1873 49
1873 49
1873 49
1873 49
1873 49
1873 49
1873 49
1873 49
1673 49
1873 49
1873 49
1873 49
1873 49
1673 49
1873 4g
1873 49
1873 49
1875 49
1873 49
1873 49
1873 49
1873 49
1873 49
1873 49
1873 49
1873 49
1673 49
14673 49
1873 49
1873 49
1873 49
1875 49
1873 49
1873 49
1473 49
1473 49
1873 49
1673 49
1873 49
1873 49
1873 49
1875 49
1673 49
1873 49
1673 49
1873 49
1673 4g
1873 49
1873 49
1873 49
1673 49
1873 49
1873 49
1673 49
1873 49
1873 49
1873 49
1673 49
1873 49
1373 49
1873 49
1673 49
1673 49
1873 49
1873 49
1873 49
1873 49
1873 49
1873 49
1873 49
1873 49
1873 49
1873 49
1873 49
1873 49
1873 49
1873 49
1473 49
1873 49
1673 a)
ooooooocoooeo°ecocoeococeoeoceoocoococoeoecocoecoecmoceceocoeoo0°co
MNVNNVNNMNNNNANANNNNNNNNNNANNNNNNNANANN NNN NA NNNNNNNANNNNNNNMNNNANNNNNNANNMNNNNNNNNNANANNNNNNNNNMNMNHNNNWN ND ff
coooooocoocoeooc*coooo°”oo
V HYB PROV QUAD LAT LONG
6047
6047
6047
6047
6045
6045
6043
6043
6023
6059
6034
6043
6010
6025
6022
6022
6022
6022
6005
6003
5842
5839
5845
5843
5842
5804
5804
5947
5947
6130
6130
6145
6146
6145
6151
6133
6146
6159
6157
6113
6113
6113
6157
6148
6157
6157
6158
6155
6155
6155
6155
6155
6155
6155
6153
6146
6134
6259
6225
6232
6233
6233
6230
6228
6231
6235
6233
6258
6258
6203
6231
6218
6218
6215
6216
6330
6233
6252
6252
6252
6255
6230
6253
6223
6223
6341
6320
6321
6341
6341
6322
13735
13735
13735
13741
13738
13615
13503
13503
13449
13510
13433
13503
13442
13335
13351
13351
13351
13351
13228
12840
15639
15637
15637
15700
15639
15304
15304
15648
15648
14300
14300
14630
14510
14630
14519
14525
14511
14653
14710
14953
149535
14953
14710
14737
14710
15436
15438
15430
15425
15425
15425
15425
15425
15425
15421
15808
15934
16318
15350
15337
15336
15337
15340
15350
15353
15338
15336
15520
15520
14631
14528
14518
14518
14525
14523
14530
14528
14128
14128
14128
14332
14330
14333
14321
14321
14215
14236
14235
14215
14213
14232
LOCALI
PINE CREE
PINE CREE
PINE CREE
BEAR CREE
ALSEK RIV
MENDENHALL R
WHITEHORS
WHITEHORS
MINTO
TY
K
K
K
K
ER
E
2
DAWSON RD M13
NW END MARSH L
WHITEHORS
CARCROSS
SQUANGA LAKE
LTL ATLIN
LTL ATLIN
LTL ATLIN
LTL ATLIN
ALA HWY M793
WATSON LA
KING SALM
NAKNEK
NAKNEK
NAKNEK
KING SALM
PORT VITA
PORT VITA
MULCHATNA
MULCHATNA
KENNICOTT
KENNICOTT
TAZLINA G
RICH HwY M87
TAZLINA G
=
L
L
L
L
KE
ON
ON
R
R
GL
GL
L
L
VY
VY
COPPER CENTER
RICH HwY M70
WILLOW MT
LTL NELCHINA R
GLENN HWY M128
ANCHORAGE
ANCHORAGE
ANCHORAGE
GLENN Hwy
GLENN HWY
GLENN Hwy
BIG RIVER
BIG RIVER
HEAD OF B
HEAD OF B
HEAD OF B
HEAD OF B
HEAD OF B
HEAD OF B
HEAD OF B
HEAD OF B
CANOE MTe
ANIAK
PASTOLIK
FAREWELL
FAREWELL
FAREWELL
FAREWELL
FAREWELL
FAREWELL
FAREWELL
FAREWELL
FAREWELL
KUSKOKWIM
KUSKOKWIM
M128
M109
M128
IG
IG
IG
IG
IG
IG
IG
IG
Le
Le
Le
MT.
Le
Le
Re
Re
Re
Re
Re
Re
Re
Re
Re
Re
GLENN HWY M152
RICH HwY M146
GAKONA
GAKONA
GULKANA
GULKANA
RAINBOW M
SOURDOUGH CR
M1247
M1247
M1247
CREEK
RD M29
L CAMP
ALA HWY
ALA HWY
ALA HWY
BARTELL
NABESNA
DEADMAN
TANACROSS
TANACROSS
MT FAIRPL
TETLIN JC
T
AY
ul
TAYLOR HWY M2
MT FAIRPL
AY
TAYLOR HWY M34
TAYLOR HWY M5
DATE
17
17
23
10
24
30
31
01
02
10
19
29
13
19
19
29
24
24
23
27
03
03
19
10
21
21
06
06
19
©O0OOWOWOMOD DN DW N NNN NNN NNN
ooo
ann
a
nO
oooo°ococo
NDONNNNN
07
944
944
944
948
944
966
944
932
932
960
960
943
943
966
943
943
943
943
966
966
965
949
949
952
952
946
939
965
965
957
957
957
967
957
967
967
967
968
944
947
947
948
947
967
965
950
950
950
950
950
950
950
950
950
950
949
949
931
949
949
949
949
949
949
949
949
949
949
949
946
935
944
944
957
932
965
968
944
944
944
944
962
968
944
944
948
963
963
960
963
963
COLLECTOR NAME
ee ee ee
a)
cuwrnrrnrrrrerree< ZZ ZEEKXAACOMWTIITIGOIICc
GCacceseSaCCCYVSCCCCUYUE OCCCCO EHEZEFTEZEEAZLZFOAATSFEZLAEZEAFZEZAZEZEYO MM
6.5.4.5. &
UreveeperprprPrPrrrnrraeagnrasIewrrwragrs SESESEPD
EvVvevewrrewoS TL eSerreEeEltehpheoT Sake Ss
fat Lael" a I tad fe
Our
RAUP
RAUP
RAUP
RAUP
RAUP
SUDA
ANDERSON
BEAUCHAMP
BEAUCHAMP
CALDER
CALDER
RAUP
RAUP
ARGUS
RAuUP
RAUP
RAUP
RAUP
WELSH
ARGUS
HARMS
RAUP
RAUP
SCHOFIELD
SCHOFIELD
EYERDAM
EYERDAM
HARMS
HARMS
VIERECK
VIERECK
VIERECK
VIERECK
VIERECK
VIERECK
VIERECK
VIERECK
WELSH
ANDERSON
DUTILLY
DUTILLY
LEPAGE
SCAMMAN
VIERECK
WELSH
DRURY
DRURY
DRURY
DRURY
DRURY
DRURY
DRURY
DRURY
DRURY
DRURY
DRURY
DRURY
ROUSE
DRURY
DRURY
DRURY
DRURY
DRURY
DRURY
DRURY
DRURY
DRURY
DRURY
DRURY
ANDERSON
ANDERSON
ANDERSON
ANDERSON
ARGUS
BEAUCHAMP
HARMS
WELSH
ANDERSON
ANDERSON
ANDERSON
ANDERSON
SHARROCK
WELSH
ANDERSON
ANDERSON
ANDERSON
BARTHOLOMEW
BARTHOLOMEW
CALDER
NAVA
NAVA
COL NO
11788
11786
11877
130A0
11896
16566
9696
24874
24923
11138
11459
5074
11275
11428
11470
HFRB
ALA
ALA
ALA
ALA
ALA
GWA
1sc
ALA
ALA
GWA
GWA
ALA
ALA
GWA
ALA
ALA
ALA
ALA
GWA
SWA
ALA
msc
msc
GWA
GWA
Isc
Isc
ALA
ALA
ALA
SWA
ALA
GWA
BWA
GWA
GWA
GWA
GWA
1sc
Isc
Tsc
™sc
GWA
GWA
Tsc
CAN
CAN
CAN
CAN
CAN
CAN
CAN
CAN
CAN
CAN
CAN
CAN
SWA
CAN
CAN
CAN
CAN
CAN
CAN
CAN
CAN
CAN
CAN
CAN
rsc
Tsc
1s¢
tse
SWA
ALA
ALA
GWA
msc
msc
TSsc
tsc
ALA
GWA
TSsc
rsc
Tsc
ALA
ALA
GWA
ALA
ALA
+ NO
19775
19480
19752
19470
19771
256575
524
521
19774
19767
19568
19755
19754
19761
32565
219352
219356
256591
256591
32561
32562
8412
8411
256599
256159
256556
256589
246046
256579
256596
256568
256569
522
32609
256578
256593
256683
256594
25404
256573
256572
256685
25903
25904
23943
23941
225
Appendix B
TAXON SPC S V
1873 49 0 2
1873 49 0 2
1873 49 02
1873 49 02
1673 49 0 2
1873 49 0 2
1873 49 02
1873 49 0 2
1873 49 02
1673 49 02
1873 49 0 2
1673 49 0 2
1873 49 0 2
1873 49 02
1873 49 0 2
1873 49 02
1873 49 02
1873 49 0 2
1873 49 02
1873 49 0 2
1873 49 02
1873 49 0 2
1873 49 0 2
1873 49 0 2
1873 49 0 2
1873 49 0 2
1873 49 02
1873 49 0 2
1873 49 0 2
1873 49 0 2
1873 49 02
1873 49 02
1873 49 0 2
1873 49 02
1873 49 0 2
1873 49 02
1873 49 0 2
1873 49 0 2
1873 49 02
1873 49 0 2
1873 49 02
1873 49 02
1873 49 0 2
1873 49 0 2
1873 49 0 2
1873 49 02
1673 49 02
1873 49 02
1873 49 0 2
1873 49 0 2
1873 49 0 2
1873 49 0 2
1873 49 02
1673 49 02
1673 49 0 2
1873 49 0 2
1873 49 0 2
1873 49 0 2
1873 49 0 2
1873 49 02
1673 49 0 2
1873 49 0 2
1873 49 0 2
1873 49 02
1873 49 0 2
1873 49 02
1873 49 02
1873 49 02
1873 49 0 2
1873 49 0 2
1873 49 02
1873 49 02
1873 49 0 2
1873 49 02
1873 49 02
1873 49 02
1873 49 02
1873 49 02
1873 49 02
1873 49 02
1873 49 02
1873 49 0 2
1873 49 02
1873 49 02
1673 49 02
1673 49 0 2
1873 49 02
1873 49 02
1873 49 02
18673 “gy 2
1873 49 0 2
226
HYB PROV QUAD LAT
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
0332
6341
6313
6318
6357
o3il
6310
6310
6310
6310
6315
6315
6340
6340
6340
6340
6340
6340
6340
6345
6345
6310
6448
6346
6334
6325
6325
6355
6351
6351
6351
6340
63406
6340
6340
6335
6325
6340
6340
6340
6335
6325
6344
6335
6351
6335
6243
6244
6335
60331
6335
6325
6244
6352
6332
6332
6332
6330
6330
6328
6325
6332
6332
6330
6330
6328
6327
6332
6232
6232
6330
6332
6232
6232
6330
6327
6324
0327
6325
6327
6324
6324
6324
6327
6326
6324
6324
6330
6330
6301
6352
LONG
14352
14215
14542
14542
14547
14533
14530
14530
14530
14530
14530
14540
14551
14645
14645
14645
14645
14645
14645
14433
14433
14541
14545
14545
14551
14545
14545
14548
14858
14858
14858
14930
14930
14930
14930
14935
14900
14930
14930
14930
14935
14900
14855
14940
14858
14935
14912
14855
14935
14956
14935
14900
14855
14902
15058
15058
15058
15002
15002
15010
15020
15058
15058
15002
15002
15010
15052
15058
15001
15001
15003
15058
15001
15001
15003
15050
15025
15045
15050
15045
15025
15025
15020
15050
15020
15020
15020
15002
15002
15532
16047
LOCALITY
ALA HWY M1349
MT FAIRPLAY
RICH HwY M202
RICH HWY M207
RICH HwY M249
ISABELLA PASS
PHELAN CREEK
PHELAN CREEK
PHELAN CREEK
PHELAN CREEK
PHELAN CREEK
MCCALLUM CREEK
DONNELLY DOME
MT HAYES
MT HAYES
MT HAYES
MT HAYES
MT HAYES
MT HAYES
ALA HWY M1380
ALA HWY M1380
FIELDING LAKE
DONNELLY DOME
DONNELLY DOME
RICH HwY M230
DELTA RIVER
BLACK RAPIDS G
BIG DELTA
HEALY
HEALY
HEALY
TEKLANIKA R
TEKLANIKA R
TEKLANIKA R
TEKLANIKA R
IGLOO CREEK
CANTWELL
TEKLANIKA R
TEKLANIKA R
TEKLANIKA R
IGLOO CREEK
CANTWELL
MCKINLEY PARK
SABLE PASS
HEALY
IGLOO CREEK
MCK PK RD M7
MCKINLEY PARK
IGLOO CREEK
POLYCHROME PS
IGLOO CREEK
CANTWELL
MCKINLEY PARK
HEALY
KANTISHNA
KANTISHNA
KANTISHNA
TOKLAT R
TOKLAT R
HIGHWAY PASS
CAMP EIELSON
KANTISHNA
KANTISHANA
TOKLAT R
TOKLAT R
HIGHWAY PASS
MCKINLEY BAR
KANTISHNA
MCK PK RD
MCK PK RD
TOKLAT
KANTISHNA
MCK PK RD
MCK PK RD
TOKLAT
WONDER LAKE
MULDROw GL
MCKINLEY RIVER
MCKINLEY RIVER
MCKINLEY RIVER
MULDROW GL
MULDROW GL
THOROFARE R
WONDER LAKE
THOROFARE R
THOROFARE R
THOROFARE R
TOKLAT R
TOKLAT R
APPEL MTe
UNALAKLEET
M35
M35
M35
M35
DATE
08
08
07
O44
951
935
946
948
935
957
957
957
957
957
957
951
933
941
941
941
941
941
944
944
961
967
967
967
966
957
965
939
939
939
956
956
956
956
956
956
956
956
956
956
956
947
954
950
959
950
960
939
939
939
924
940
967
956
956
956
956
956
956
956
956
956
956
956
956
967
939
939
939
939
939
939
. 939
939
939
939
958
958
958
958
958
956
956
956
956
956
965
965
950
938
COLLECTOR NAME cOL NO
H M RAUP 12743
Iz SCAMMAN 6254
J P ANDERSON 2129
J P ANDERSON 10190
J © ANDERSON 10860
J P ANDERSON 2702
G6 w ARGUS 1148
G W ARGUS 1149
G w ARGUS 1147
G6 w ARGUS 1145
G w ARGUS 1144
G6 W ARGUS 1032
w J CODY 5739
L J PALMER 456
L J PALMER 630
L J PALMER 456
L J PALMER 459
L J PALMER 458
L J PALMER 643
H M RAUP 12658
H “ RAUP 12760
R SPOONER Pas
L A VIERECK 8332
L A VIERECK R304
L A VIERECK 8204
L A VIERECK 8017
L A VIERECK 2133
S t WELSH 4336
J P ANDERSON 5710
J © ANDERSON 5710
J P ANDERSON 5762
G W ARGUS 613
G w ARGUS 612
G Ww ARGUS 615
G W ARGUS 617
G w ARGUS 691
G Ww ARGUS 598
G w ARGUS 613
G w ARGUS 612
G Ww ARGUS 615
G w ARGUS 691
G w ARGUS 598
DUTILLY 20131
G “ FROHNE 54252
D HANSON 56
A MURIE 8
A MURIE 13
A MURIE 11
A NELSON 3764
A NELSON 3779
A NELSON 3764
L J PALMER 1943
L J PALMER 4el
L A VIERECK R297
G W ARGUS 649
G w ARGUS 651
G w ARGUS 650
G w ARGUS 683
G Ww ARGUS 6R7
G w ARGUS 628
G w ARGUS 642
G w ARGUS 650
G w ARGUS 651
G w ARGUS 683
G W ARGUS 683
G w ARGUS 628
J FOOTE B427
A NELSON 3822
A NELSON 3688
A NELSON 3691
A NELSON 4094
A NELSON 3822
A NELSON 3688
A NELSON 3691
A NELSON 4094
A NELSON 3876
A NELSON 4272
L A VIERECK 3029
L A VIERECK 3119
L A VIERECK 3157
L A VIERECK 3193
L A VIERECK 3179
L A VIERECK 1095
L A VIERECK 1713
L A VIERECK 1775
L A VIERECK 1095
L #& VIERECK 109B8A
S Lt WELSH 4B4S
S L WELSH 4B61
Ww H DRURY 4669
J © ANDERSON 3266
HERB
ALA
GWA
Tsc
msc
rsc
rsc
SWA
GWA
GWA
GWA
GWA
GWA
GWA
ALA
ALA
ALA
ALA
ALA
ALA
ALA
ALA
ALA
GWA
GWA
GWA
GWA
GWA
TSC
msc
rsc
Isc
GWA
GWA
GWA
GWA
GWA
GWA
ALA
ALA
ALA
ALA
ALA
1sc
ALA
Isc
GWA
SWA
GWA
iGle
ALA
ALA
ALA
ALA
GWA
GWA
GWA
GWA
GWA
SWA
GWA
GWA
ALA
ALA
ALA
ALA
ALA
GWA
Isc
Isc
Tsc
Isc
ALA
ALA
ALA
ALA
ALA
ALA
GWA
GWA
GWA
GWA
GWA
GWA
ALA
ALA
ALA
ALA
msc
Tsc
CAN
1sc
+ NO
19756
256597
256580
256686
256584
32150
5190
5635
5625
5626
5193
19757
19764
19074
246879
27551
256414
256415
4562
4760
4566
4907
4908
256557
21850
256567
256408
534
531
5198
5632
4739
4962
4578
22465
4734
256412
256411
256410
256409
527
526
528
529
530
515
11605
11619
11617
3501
247169
247201
256581
Specimens Cited
TAXON SPC
1873 49
1673 49
1873 49
1873 49
1873 49
1873 49
1673 49
1873 49
1873 49
1873 49
1873 ug
1873 49
1873 49
1873 49
1873 49
1873 49
1873 49
1873 49
1873 49
1873 49
1873 49
1873 49
1673 49
1873 49
1873 49
1875 4g
1873 49
1873 49
1873 49
1873 49
1673 49
1873 4g
1873 49
1873 49
1873 49
1873 ug
1873 49
1873 49
1873 49
1873 49
1673 49
1873 49
1873 49
1873 49
1673 ug
1873 49
1873 49
1873 49
1873 49
1873 49
1873 49
1873 49
1873 49
1873 49
1873 49
1873 49
1873 49
1873 49
1873 49
1873 49
1873 49
1673 49
1873 49
1873 49
1873 49
1873 49
1873 49
1873 49
1873 4g
1673 49
1873 4g
1873 ag
1873 4g
1873 49
1873 49
1873 49
1873 49
1873 49
1873 49
1873 ag
1873 ag
1873 a)
1873 4g
1673 43
1873 ag
1873 4g
1873 49
1673 49
1873 4g
1873 ag
1873 49
ooooocoocoocoocooce°cococcocoocococcocoocococo°ocococococcococcococococooco°cococoooeoococococcoeceoeccocoococeceeceococoocoococeocoececoco0coocoo CCC cCeCceeCcCcCcoOCcCco
NNNNNNANNANNAA NA NNN N NNN NNNNANNNNANNNANNNNNWNANNNNNNANNNNNNNNNNANNNNANNNNMNAN NNN NNN NNNNNMNNNWN WYNN WNW NW WP
V HYB PROV QUAN LAT
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
91
91
92
92
92
92
92
92
92
92
92
92
92
94
94
95
95
a7
97
97
97
98
98
98
98
98
98
98
98
100
100
100
100
100
~ 100
100
100
100
100
100
100
100
100
100
100
100
100
100
100
100
100
100
100
100
100
100
100
100
100
100
100
100
100
100
100
100
100
100
100
100
100
100
100
100
102
102
102
102
102
102
102
102
102
102
102
102
104
104
104
104
104
6352
6352
6329
6331
6329
6329
6335
6335
6335
6335
6335
6335
6302
6430
6430
6458
6458
6445
6420
6420
6420
6445
6445
6445
6445
6445
6445
6445
6445
6447
6453
6453
6453
6452
6452
6452
6452
6452
6454
6454
6454
6454
6454
6454
6454
6454
6454
6454
6454
6454
6453
6453
6453
6453
6453
6453
6453
6452
6452
6454
6453
6453
6452
6451
6451
6451
6451
6451
6451
6451
6451
6443
6453
6452
6410
6410
6410
6410
6450
6450
6450
6450
6450
6450
6450
6405
6550
6532
6532
6532
6532
LONG
16047
16047
16202
16217
16202
16202
16230
16230
16230
16230
16230
16230
16318
16530
16525
16310
16310
15657
15843
15843
15843
15530
15530
15530
15530
15530
15530
15530
15530
14757
14750
14750
14750
14750
14750
14750
14750
14750
14750
14751
14751
14751
14751
14751
14751
14751
14751
14751
14750
14750
14752
14752
14752
14752
14752
14750
14750
14750
14750
14750
14752
14752
14750
14752
14743
14743
14743
14743
14743
14743
14743
14806
14750
14750
14147
14147
14147
14147
14340
14340
14340
14340
14340
14340
14340
14147
14404
14513
14513
14513
14513
LOCALITY
UNALAKLEET
UNALAKLEET
ST MICHAEL
STEBBINS
ST MICHAEL
ST MICHAEL
STUART
STUART
STUART
STUART
STUART
STUART
PASTOLIK
NOME
NOME
ETCHEPUK R
ETCHEPJK R
GALENA
KALTAG
KALTAG
KALTAG
RUBY
RUBY
RUBY
RUBY
RUBY
RUBY
RUBY
RUBY
R
ISLAND
ISLAND
ISLAND
ISLAND
ISLAND
ISLAND
CHENA 3LUFFS
COLLEGE
COLLEGE
COLLEGE
COLLEGE
COLLEGE
COLLEGE
COLLEGE
COLLEGE
COLLEGE
GOLDSTREAM
GOLDSTREAM
GOLDSTREAM
GOLDSTREAM
GOLDSTREAM
GOLDSTREAM
GOLDSTREAM
GOLDSTREAM
GOLDSTREAM
GOLDSTREAM
GOLDSTREAM
SMITH LAKE
SMITH LAKE
SMITH LAKE
SMITH LAKE
SMITH LAKE
COLLEGE
COLLEGE
COLLEGE
COLLEGE
COLLEGE
SMITH LAKE
SMITH LAKE
COLLEGE
CR
cR
CR
CR
CR
CR
CR
CR
CR
CR
cR
UNIV EXP FARM
FAIRBANKS
FAIRBANKS
FAIRBANKS
FAIRBANKS
FAIRBANKS
FAIRBANKS
FAIRBANKS
FAIRBANKS
COLLEGE
COLLEGE
FRANKLIN
FRANKLIN
FRANKLIN
FRANKLIN
CHARLIE
CHARLIE
CHARLIE
CHARLIE
CHARLIE
CHARLIE
CHARLIE
CHICKEN
CIRCLE
MILLER
MILLER
MILLER
MILLER
RIVER
RIVER
RIVER
RIVER
RIVER
RIVER
RIVER
HOUSE
HOUSE
HOUSE
HOUSE
DATE
29
10
19
02
14
14
29
20
29
29
29
29
19
16
23
02
02
18
04
COLLECTOR NAME
MMMMOCKMDONDODODADADOCCCCHKrOOKOTOTTTTTKTCKEDDDADADAAVADAAVAVDAVDD ANYDVD AVNANAVADANAANAADNVDGVIAIVNDDAAVAADVDAVIAININAANNAANAANAIAMABDAZENINNVAVNANANANNNNACCCCH
Se ee eae ar
n
VUFPzECCLCLEETCvVVIvVerrvCcecccccartM CECT TET ECEACaACEEACALECEAZESEAREREECEALELCAEAZEAECOCTIOCITELTIICICITFT
ANDERSON
PALMER
ANDERSON
ANDERSON
ROUSE
ROUSE
ROUSE
ROUSE
ROUSE
ROUSE
ROUSE
ROUSE
ROUSE
HELLER
MILLER
PEGAU
PEGAU
LITTLE»
ROUSE
ROUSE
ROUSE
ROUSE
ROUSE
ROUSE
ROUSE
ROUSE
ROUSE
ROUSE
ROUSE
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
HARMS
OKEDARA
PALMER
PALMER
PALMER
PALMER
PALMER
PALMER
PALMER
VIERECK
VIERECK
VIERECK
ANDERSON
ANDERSON
ANDERSON
ANDERSON
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
HARMS
ANDERSON
SCAMMAN
SCAMMAN
SCAMMAN
SCAMMAN
JR.
COL NO
50A1
1211
3449
3733
31
31
4
49
4
49
25
25
30
1072
112C
wi3
wi3
19505
42
42
48
48
52
46
48
52
32
4B
46
363
1008
1009
1005
342
1019
1196
1020
5107
436
1174
1166
1163
1164
1169
1168
1165
1167
1171
5089
5088
781
311
287
298
207
1009
1005
342
1019
436
7Al
287
398
R7
154
193
160
185
186
180
178
B1i01
R299
uBNT
7142
7300
7142
7300
866
876
B61
878
866
861
878
3070
2505
3493A
733
2042
2043
HERS + NO
Isc 256559
ALA 2028
TSC 256530
ISC 256595
ALA 2042
ALA 25449
ALA 2039
ALA 25437
ALA 25441
ALA 2074
ALA 25463
ALA 2051
ALA 25438
ALA 27068
ALA 2729
GWA
ALA 32701
GWA
ALA 25450
ALA 21488
ALA 2019
GWA
GWA
GWA
ALA 25434
ALA 25451
ALA 2015
ALA 2018
ALA 2045
GWA
GWA
GWA
GWA
GWA
GWA
GWA
GWA
SWA
GWA
GWA
SWA
GWA
5WA
GWA
GWA
GWA
GWA
GWA
5WA
5WA
65WA
GWA
GWA
GWA
GWA
ALA 8408
ALA 16289
ALA 4511
ALA 16290
ALA 4523
ALA 4699
ALA 4518
ALA 32660
ALA 27172
GWA
ALA 5225
ALA 5273
ALA 5278
ALA 5279
ALA 5277
ALA 5276
GWA
GWA
GWA
ISC 256407
ISc 256406
ALA i |
ALA 532
GWA
GWA
GWA
GWA
ALA 4897
ALA 4847
ALA 4898
ALA 32613
Isc 256585
SWA
GWA
GWA
GWA
227
Appendix B
5 GLAUCA VAR» VILLOSA
TAXON SPC
1873 49
1873 a7
1873 49
1873 49
1873 4s
1873 49
1873 ve)
1873 49
1873 49
1873 49
1873 a
1873 43
1873 Ph)
1873 ute)
1873 4g
1873 49
1873 as
1873 aa)
1873 49
1873 49
1873 49
1873 aS
1873 4g
1873 49
1873 49
1873 Gg
1873 43
1873 4g
1873 4S
1873 ag
1873 ad
1873 49
1873 49
1873 4d
1873 49
1873 49
1873 49
1873 49
1873 49
1873 49
1873 49
1873 ug
1873 49
1873 49
1873 ug
1873 49
1873 4g
1873 49
1873 49
1873 49
1873 ag
1873 4g
1873 4g
1873 ug
1873 49
1873 49
1873 29
1673 ug
1873 4g
1873 49
1673 49
1873 ug
1873 49
1673 49
1873 4g
1873 49
1873 49
1873 4g
1873 4g
1873 ae)
1873 49
1873 +)
1873 4g
1873 49
1873 ay
1673 49
18673 49
1873 4G
1873 ag
1873 49
a3
1873 ae
1873 49
1873 49
1873 49
1873 4g
1873 49
1873 4g
228
oooo0oocoocoococoocococoecocoeceocooecoeoccoecoecoocoeocoecoeoecoececococeocoecocooceocococecooceoecoccoocoococeoccocoocooocoococecoeococococoococoocooco°ocoo
NNN NNMNNNNANNNNANNNNNNNANNANNNNANAN NNN NNNNNNDANN NNN NANNNNNNNNNANA NNN NNNNNN NNN NWN HN NWN PW AW WW fo
V HYB PROV QUAD LAT
Onunnnnun
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
NNNNNNPE
104
104
104
104
104
104
104
104
104
105
105
105
105
113
113
113
114
118
119
119
120
120
120
120
121
121
121
121
121
121
121
121
121
121
421
121
121
121
121
121
121
421
121
121
121
121
121
121
121
124
124
124
124
124
124
124
124
124
124
124
124
128
136
136
136
136
136
136
136
137
141
141
141
141
141
141
141
144
141
141
wo
FNNMNNNN
i
6532
6532
6532
6524
6532
6532
6532
6532
6532
6505
6520
6550
6532
6654
6604
6654
6658
6630
6635
6635
6644
6644
6644
6644
6710
6710
6717
6710
6710
6710
6710
6710
6710
6717
6715
6710
6717
6717
6710
6710
6710
6710
6710
6712
6708
6716
6717
6723
6723
6725
6725
6725
6725
6725
6725
6725
6725
6725
6725
6725
6725
6759
6803
68035
6803
68035
6803
6803
6803
6839
6923
6923
6923
6923
6922
6922
6922
6922
6922
6922
9213
5941
5941
5941
5935
5941
5822
LONG
145135
14513
14513
14554
14513
14513
14513
14513
14513
14750
14800
14750
14833
16235
16242
16235
16026
14742
14510
14510
14334
14334
14334
14334
14155
14155
14135
14140
14140
14140
14140
14155
14155
14135
14155
14155
14135
14135
14140
14140
14140
14140
14140
14138
14208
14138
14140
14350
14350
15007
15007
15006
15006
15007
15007
15007
15007
15006
15006
15006
15006
16433
14500
14500
14500
14500
14500
14500
14500
14100
15210
15210
15210
15210
15210
15210
15210
15210
15210
15210
11712
13632
13632
13632
13650
13632
12955
LOCALITY
MILLER HOUSE
MILLER HOUSE
MILLER HOUSE
TWELVEMILE CR
MILLER HOUSE
MILLER HOUSE
MILLER HOUSE
MILLER HOUSE
MILLER HOUSE
PEDRO DOME
LIVENGOOD
LIVENGOOD
LIVENGOOD
KOTZEBUE
DEERING
KOTZEBUE
KIANA
NELSON LAKE
FORT YUKON
FORT YUKON
FISH HOOK
FISH HOOK
FISH HOOK
FISH HOOK
PORCUPINE
PORCUPINE R
PORCUPINE R
OLD RAMPART
OLD RAMPART
OLD RAMPART
OLD RAMPART
PORCUPINE
PORCUPINE
PORCUPINE
PORCUPINE
PORCUPINE
PORCUPINE
PORCUPINE
OLD RAMPART
OLD RAMPART
OLD RAMPART
OLD RAMPART
OLD RAMPART
PORCUPINE R
PORCUPINE R
PORCUPINE R
PORCUPINE R
SMALL LAKE
SMALL LAKE
WISEMAN
WISEMAN
WISEMAN
WISEMAN
WISEMAN
WISEMAN
WISEMAN
WISEMAN
WISEMAN
WISEMAN
WISEMAN
WISEMAN
KIVELINA
OLD JOHN
OLD JOHN
OLD JOHN
OLD JOHN
OLD JOHN
OLD JOHN LAKE
OLD JOAN LAKE
FIRTH RIVER
UMIAT
CR
CR
CR
CR
R
DAADAADAD
LAKE
LAKE
LAKE
LAKE
LAKE
UMIAT
UMIAT
UMIAT
UMIAT
UMIAT
UMIAT
UMIAT
UMIAT
UMIAT
SUNWAPTA PASS.
NADAHINI RIVER
NADAHINI RIVER
NADAHINI RIVER
INSPECTOR CREE
NADAHINI RIVER
DEASE L VILLAG
DATE
07
06
07
06
06
06
06
06
06
07
08
06
06
08
08
08
06
06
06
07
DBNNNNN®
940
945
940
957
953
953
953
953
953
956
957
959
940
938
938
951
937
933
965
965
957
957.
957
957
951
967
957
957
957
957
954
951
951
954
957
957
957
957
S577
957
957
957
957
961
961
961
961
957
957
939
931
939
939
962
962
962
962
937
940
940
940
938
957
957
S5ir
957
957
954
954
961
966
966
966
966
964
964
964
964
964
964
96B
967
967
967
967
967
969
COLLECTOR NAME
DDDODDD<<<<MYNNUNNNUNYCMMMMAADADAcCCCCHUNAUNAMANCCCCCCCCCCCCCCCCCCCYNYNYUYNEKCOOMCCMADDYNYNUNNNYMMM
AANDAADAD
D
v=EezqndDNGdDN|D
vO
pl yaar alta) ayo) ie om ce eel ee oe lee ee ell a lo ia melo 3 (>
ett tze
SCAMMAN
SCAMMAN
SCAMMAN
SHETLER
SMITH
SMITH
SMITH
SMITH
SMITH
ARGUS
ARGUS
O: FARRELL
SCAMMAN
ANDERSON
ANDERSON
SCAMMAN
CLARK
GEIST
HARMS
HARMS
SHETLER
SHETLER
SHETLER
SHETLER
BUCKLEY
BUCKLEY
BUCKLEY
BUCKLEY
BUCKLEY
BUCKLEY
BUCKLEY
BUCKLEY
BUCKLEY
BUCKLEY
BUCKLEY
BUCKLEY
BUCKLEY
BUCKLEY
BUCKLEY
BUCKLEY
BUCKLEY
BUCKLEY
BUCKLEY
DEAN
DEAN
DEAN
DEAN
SHETLER
SHETLER
ANDERSON
ANDERSON
ANDERSON
ANDERSON
BROCKMAN
BROCKMAN
BROCKMAN
BROCKMAN
SCAMMAN
SCAMMAN
SCAMMAN
SCAMMAN
ANDERSON
SHETLER
SHETLER
SHETLER
SHETLER
SHETLER
SMITH
SMITH
LITTLE?»
SUDA
SUDA
SUDA
SUDA
WEST
WEST
WEST
WEST
WEST
WEST
JR.
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
RIGBY
COL NO 4E8B + NO
2042
34O3A
2043
AFI61
17698
1769A
1769A
17708
17698
569
1193
27
1717
4706
4BN7
6458
SeNe
3821
3820
Z1i4AF
GUSAF
357AF
353AF
SeNe
189
114
182
179
153
SeNe
SeNe
SeNe
SeNe
190
189
114
114
153
153
179
1e2
177
SeNe
20
22
al
6627AF
BULAF
SWA
GWA
GWA
GWA
GWA
GWA
ALA 10649
ALA 10641
ALA 10650
SWA
SWA
ALA 9631
GWA
TSC 256598
ISC 2565868
GWA
ALA 10188
ALA 29715
ALA 32597
ALA 32596
ALA 3868
ALA 3948
ALA 3887
ALA 3885
GWA
GWA
SWA
GWA
SWA
6WA
ALA 5625
ALA 267575
ALA 5039
ALA 5626
ALA 5680
ALA 5037
ALA 5986
ALA 26769
ALA 5024
ALA 26760
ALA 5677
ALA 5085
ALA 5091
ALA 24078
ALA 24059
ALA 24064
ALA 24060
ALA 4087
ALA 4193
ALA 27556
ALA 27502
TSC 256544
ISC 256558
ALA 28512
ALA 28437
ALA 28483
ALA 28466
GWA
GWA
SWA
SWA
TSC 256560
GWA
SWA
ALA 4363
ALA 4249
ALA 4364
SWA
ALA 10737
GWA
GWA
SASK
SASK
SASK
GWA
GWA
GWA
GWA
GWA
GWA
SWA
SWA
SWA
SWA
SWA
SWA
ITF 1142
Specimens Cited
TAXON SPC S
1873 49:05 2
1873 49 05 2
1873 49 05 2
1873 49 05 2
1873 4905 2
1873 49 05 2
1873 49 05 2
1673 49 05 2
1873 49 05 2
1673 49:0 5 2
1673 49:05 2
1873 495 2
1873 4905 2
1873 49 05 2
1873 49:05 2
1873 4905 2
1873 4905 2
1873 49 05 2
1873 49 05 2
1873 49— 5 2
1873 49 05 12
1873 49 05 12
1873 49:05 12
1873 49 05 12
1873 49:05 12
1873 49 05 12
1875 49 05 12
1673 4905 12
1873 4905 12
1875 49 0 5 12
18735 49 05 12
1673 49 05 Le
1873 4905 12
ings CS 12
1873 49:05 12
1873 49 05 12
1873 49 0 5 12
1673 49 05 12
1873 4905 12
1673 49 05 12
1873 49 0 5 12
1873 49 05 12
1873. 49:05 12
1473 49 05 12
1873 49:0 5 12
1875 49 05 12
1873 49 05 12
1873 49:05 12
1673 49 05 12
1873 49 0 5 12
1875 49:0 5 12
1673 49 05 12
1673 49:0 5 12
1873 49 05 12
1873 49 05 12
1873 49 05 12
1573 49 05 12
1873 49 05 12
1873 49 05 ie
50 ARCTOPHILA
1873 50 3
1873 50 7
1873 50 Te
1873 50 7
1873 =650 U
1873 50 7
1873 50 7
1873 50 Tf
1873 + 50 7
1673 50 12
1873 50 12
1873 50 14
1873 50 14
1873 50 14
1873 50 14
1873 50 14
1873 50 14
1873 50 14
1873 50 14
1873 50 14
1873 50 14
1873 50 14
1873 50 14
152
V HYB PROV QUAD LAT
5840
5840
5840
5840
5448
5448
5831
5831
5831
5847
5847
5739
5705
5705
5705
5714
5714
5723
5723
5320
6409
6229
6120
6120
6120
6120
6120
6120
61035
6103
61035
6103
6140
6140
6143
6045
6045
6051
6051
6051
6043
6030
6048
6010
6025
6025
6013
6022
6022
6022
6002
6002
60135
6015
6015
6013
60135
6013
6013
5842
6927
6900
6750
6050
6050
6050
6050
6050
6912
6315
6810
6822
6832
6832
6839
6839
6926
7010
6950
7005
7005
7005
LONG
12425
12425
12425
12425
12500
12500
12434
12434
12434
12500
12500
12844
12235
12235
12235
12243
12243
12248
12248
12245
13953
13024
13910
13910
13910
13910
13910
13910
13831
13831
13831
13831
13602
13602
13304
13615
13615
13747
13747
13747
13503
13415
13545
13441
13335
13335
13252
13351
13351
13351
12835
12835
12840
12840
12840
12840
12840
12840
12840
9403
13302
13440
11505
09425
09425
09425
09425
09425
13830
13005
14530
14355
14355
14355
14100
14100
14347
14650
14220
14335
14335
14335
LOCALITY
N TETSA RIVER
N TETSA RIVER
N TETSA RIVER
N TETSA RIVER
MACDONALD CR
MACDONALD CR
SUMMIT PASS
SUMMIT PASS
SUMMIT PASS
MACDONALD CR
MACDONALD CR
STIKINE MT
BEATTON RIVER
BEATTON RIVER
BEATTON RIVER
SIKANNI RIVER
SIKANNI RIVER
BUCKINGHORSE R
BUCKINGHORSE R
PRINCE GEORGE.
SIXTYML RD M12
OTTER LAKE
DUKE RIVER
DUKE RIVER
DUKE RIVER
DUKE RIVER
DUKE RIVER
DUKE RIVER
KLUANE LAKE
KLUANE L
KLUANE L
KLUANE L
TWIN LAKES
TWIN LAKES
LAPIE LAKE
MENDENHALL R
MENDENHALL R
BEAR CREEK
BEAR CREEK
BEAR CREEK
WHITEHORSE
ALA HWY M8B2
TAKHINI R BR
CARCROSS
SQUANGA LAKE
SQUANGA LAKE
TESLIN LAKE
LTL ATLIN L
LTL ATLINL
LTL ATLIN L
LIARD RIVER
LIARD RIVER
WATSON LAKE
WATSON LAKE
WATSON LAKE
WATSON LAKE
WATSON LAKE
WATSON LAKE
WATSON LAKE
FARNWORTH Le
TUKTOYAKTUK
RICHAROS IS
COPPERMINE
MH MCCONNELL
MH MCCONNELL
MH MCCONNELL
MH MCCONNELL
MH MCCONNELL
KING POINT
CANOL RD M 284
ARCTIC VILLAGE
OLD WOMAN CR
LOBO LAKE
LOB0 LAKE
FIRTH RIVER
FIRTH RIVER
JAGO LAKE
BULLEN
NUVAGAPAK PT
BARTER ISLAND
BARTER ISLAND
BARTER ISLAND
DDDAD
DATE
14
20
06
06
06
06
06
06
07
07
07
06
06
8
06
06
06
07
07
09
09
6
06
08
08
08
08
08
08
08
07
07
07
07
06
6
06
06
06-
06
06
06
07
06
07
07
06
06
06
08
08
08
08
08
966
966
966
966
966
966
943
943
943
968
968
969
943
943
943
943
943
943
943
967
966
960
966
COLLECTOR NAVE
<<~<<<<<TITIIIICOHOUMUMNUNCTIAEITYVGAVPUYNUNAIETIEIAMVDAVGVGVOVONCHNOIICATIETATIEIAYVNTIIVDAMQANGD®D
DPOODODODCMMBAWAW<e PPA KAKAKRAC HP KPO
S$SseseztetetteervprzesesKFsesesarreseseseztTetezeEe
S45 SS. 2 Se Oe ee ee ee
ane ee > we
cr
ZZEz£E
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
RAUP
RAuUP
RAUP
WELSH
WELSH
IGBY
RAUP
RAUP
RAUP
RAUP
RAUP
RAuUP
RAUP
ARGUS
WELSH
CALDER
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
RAUP
RAUP
RAuUP
RAuUP
WELSH
WELSH
PORSILD
ARGUS
ARGUS
RAUP
RAUP
RAUP
ANDERSON
WELSH
WELSH
WELSH
ARGUS
ARGUS
CALDER
RAuP
RAUP
RAUP
RAUP
RAUP
SUDA
SUDA
SUDA
SUDA
SUDA
SUDA
SUDA
ARGUS
PORSILD
PORSILD
LARSEN
MACINNES
MACINNES
MACINNES
MACINNES
MACINNES
PORSILD
PORSIL)
HARMS
KESSEL
KESSEL
KESSEL
LITTLE?
LITTLE»
CANTLON
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
JR
JR
COL NO
4975
4972
4974
4973
4978
4979
10671
10473
10672
7324
7326
265
10259
10257
10028
10409
10410
11620
11621
674
5582
27778
608
6007
6010
6012
6004
6005
12184
12217
12430
12183
5549
5549
9304
5080
50P1
11810
11813
11812
9607
5502
7764
7733
5075
5073
25742
11387
11457
11455
in9R2
10987
5019
11866
12966
13066
12766
17866
11766
142/58
HERB + NO
GWA
GWA
GWA
BWA
BWA
GWA
ALA
ALA
ALA
6WA
GWA
OTF
ALA
ALA
ALA
ALA
ALA
ALA
ALA
6WA
BWA
6WA
GWA
GWA
GWA
GWA
GWA
GWA
ALA
ALA
ALA
ALA
BWA
6WA
TSC
SWA
GWA
ALA
ALA
ALA
TSC
GWA
GWA
BWA
BWA
5WA
5WA
ALA
ALA
ALA
ALA
ALA
GWA
GWA
GWA
65WA
GWA
GWA
BWA
19477
19569
19566
1142
19901
19900
19902
19770
19768
19906
19758
19748
19773
19753
19763
256682
19882
19876
19875
256574
19762
19766
19760
19567
19769
GWA
GwA
SH
SWA
Uwo
wo
Uuwo
UwOo
uwWwd
SH
Isc
ALA
4LaA
ALA
GWA
GWA
GWA
US 2386024
GWA
SWA
GWA
SWA
GWA
256385
32594
5050
5047
229
Appendix B
TAXON SPC S V HYB PROV QUAD LAT
57
1873
1873
1873
1873
1873
1873
1873
1873
1873
1873
1873
1873
1873
1873
1873
1873
1873
1873
1873
1873
1873
1873
1873
1873
1873
1873
1873
1873
1873
1873
1873
1873
1873
1873
1873
1873
1873
1873
1873
1873
1873
1873
1873
1873
1873
1873
1873
1873
1873
1873
1873
1873
1873
1873
1873
1873
1873
1873
1873
1873
1873
1873
1873
1873
66
1673
1475
167535
1873
1873
16753
1875
16735
1875
1875
1873
1875
1673
1875
16753
1873
1875
1873
1673
230
HOOKERIANA
LASIANDRA
NNANN NNN N PR
388
22
5210
5747
5952
5952
5952
5915
§915
5915
5915
5915
5915
5915
5915
5915
5915
5932
5932
5952
5952
5952
5952
$952
5952
5955
5952
5933
5952
3952
5952
5955
5952
5952
5952
5952
5955
5955
5952
5952
5952
5940
5940
6950
6950
6950
5957
5957
5957
6950
6950
5916
5917
5916
5917
5925
5925
5925
5925
5925
6039
6039
6005
6005
6005
6005
5908
5936
5850
5402
5330
5320
6251
6251
6056
6404
6404
6404
6404
6336
6155
6145
6155
6054
6054
LONG
13105
15247
13945
13945
13945
13830
13830
13830
13830
13830
13830
13830
13830
13830
13830
13940
13940
13945
13945
13945
13945
13945
13945
13945
13945
13944
13945
13945
13945
13945
13945
13945
13945
13945
13945
13945
13945
13945
13945
14000
14000
13935
13935
13935
13932
13932
13932
13935
13935
13830
13909
13830
13909
14620
14620
14620
14620
14620
14446
14446
13930
13930
13930
13930
11126
12643
12235
12401
12830
12245
12123
12123
12318
13925
13925
13925
13925
13553
13238
13258
13238
13258
13258
LOCALITY
U VICTORIA
KODIAK
YAKUTAT BAY
YAKUTAT BAY
YAKUTAT BAY
TANIS LAKE
AZIMUTH PEAKe
TANIS
TANIS
TANIS
TANIS
TANIS
TANIS
TANIS LAKE
TANIS LAKE
YAKUTATe
YAKUTATe
YAKUTAT
YAKUTAT
YAKUTAT
YAKUTAT
YAKUTAT
YAKUTAT
YAKUTAT
YAKUTAT
YAKUTAT.
YAKUTAT
YAKUTAT
YAKUTAT
YAKUTAT
YAKUTAT
YAKUTAT
YAKUTAT
YAKUTAT
YAKUTAT
YAKUTAT
YAKUTAT
YAKUTAT
YAKUTAT
YAKUTAT
YAKUTAT
YAKUTAT
YAKUTAT
YAKUTAT
LAKE
LAKE
LAKE
LAKE
LAKE
LAKE
BA
BA
BA
BA
BA
BA
BA
BA
BA
BA
BA
BA
BA
BA
BA
BA
BA
BA
BA
BA
BA
BA
BA
BA
BA
BA
HAENKE ISLAND
HAENKE ISLAND
HAENKE ISLAND
YAKUTAT BA
YAKUTAT BA
TANIS MESA
MTH ITALIO
TANIS MESA
MTH ITALIO
MIDDLETON
MIDDLETON
MIDDLETON
MIDDLETON
MIDDLETON
CHILDS GL
CHILDS GL
HUBBARD GL
HUBBARD GL
HUBBARD GL
HUBBARO GL
SLAVE R
COAL R
FORT NELSO
VANDERHOOF
SKEENA RIVERe
PRINCE GEORGE>
FORT SIMPS
FORT SIMPS
NAHANNI BUTTE
DAWSON
DAWSON
DAWSON
DAWSON
MAYO
LAPIE R
ROSE LAPIE
LAPIE R
NISUTLIN R
NISUTLIN R
Y
Y
Y
Ye
m
Ye
Ye
Ye
Ye
Ye
Ye
Ye
Ye
Y
Y
Ye
Ye
Ye
Y
Y
Y
Y
uf
\f
uf
Y
Y
Y
R
R
Is
Is
Is
Is
IS
N
ON
ON
R
DATE
5
Ld
oO
14
14
14
19
20
19
19
19
19
19
19
19
19
23
23
14
14
14
14
14
14
15
16
17
14
14
14
15
14
14
14
14
15
15
14
14
14
21
21
19
19
19
22
21
21
22
22
10
11
11
29
23
23
7
o
o
DDDHDHDHDDHDHDHDDDDDAMHADDAHADAHADAHHRBARBRDOWNHNSN
964
916
967
967
967
967
967
967
967
967
967
967
967
967
967
967
967
967
967
967
967
967
967
967
967
967
967
967
967
967
967
967
967
967
967
967
967
967
967
899
899
899
899
899
899
899
899
899
899
965
965
965
965
956
956
956
956
956
931
931
899
899
899
899
927
960
949
919
967
967
939
939
961
914
914
902
916
949
944
944
944
944
944
COLLECTOR NAME
MUMMNMNZSECCCCCHEE CTT UAUMNAUARAUNRANNTEAADANVDADADANAGVAAVAANVDANAVDADAVNDNDANGVAVNANGVNDANDNDADNANGVGDGVDMOOAOAIOC
PPPrPrrcocrrzETrIQOgcecct
QS
ZSEZETCZFETSTLLTELTTTLTATETZEEAZTTATZTLETSTTLEZTEZAEZEE
PKROrRPrP<cccccccc
<<cc
Csesteteervs
<
AANA N >
CALDER
CHURCH
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
BREWER
BREWER
COVILLE
COVILLE
COVILLE
COVILLE
COVILLE
COVILLE
COVILLE
COVILLE
VIERECK
VIERECK
VIERECK
VIERECK
THOMAS
THOMAS
THOMAS
THOMAS
THOMAS
SETCHELL
SETCHELL
COVILLE
COVILLE
COVILLE
COVILLE
RAUP
CALDER
GILLETT
MACOUN
ARGUS
ARGUS
RAUP
RAUP
CODY
EASTWOOD
EASTWOOD
MACOUN
MALTE
CALDER
PORSILD
PORSILD
PORSILD
PORSILD
PORSILD
COL NO HERB + NO
DAO
US 1072423
GWA
GWA
GWA
GWA
GWA
GWA
GWA
GWA
GWA
GWA
GWA
GWA
GWA
GWA
GWA
GWA
GWA
GWA
GWA
GWA
GWA
GWA
GWA
GWA
GWA
GWA
GWA
GWA
GWA
GWA
GWA
GWA
GWA
SWA
GWA
GWA
GWA
US 378309
YS 378308
US 373422
US 373416
US 373423
US 373420
US 373421
US 373415
US 373413
NY
GWA
GWA
FSLC 337
FSLC 335
CAN 248171
US 2313024
US 2313033
US 2313005
NA 328886
NA 318443
US 373414
US 373418
US 373419
US 1437349
ALA 14482
ALA 17428
TSC 256566
TSC 256705
GWA
GWA
ALA 16401
ALA 16406
GWA
CAN 45302
CAN 45330
CAN 54403
CAN 122153
ISC 256701
CAN 45328
CAN 45327
ISC 256702
CAN 45329
TSC 256700
Specimens Cited
TAXON SPC S V HYB PROV QUAD LAT
1873
1873
1873
1873
1873
1873
1873
1873
1873
1873
1875
1873
1873
1873
1873
1873
1873
1873
1673
1873
1873
1873
1873
1873
1873
1873
1873
1873
1873
1873
1875
1873
1873
1873
1873
1873
1873
1873
1873
1873
1873
1873
1673
1873
71
1873
1873
18735
1873
1875
1873
1873
1673
18735
18735
1875
1873
186735
18735
1673
1873
1875
1873
1673
1873
1873
1873
1873
1873
1873
1673
1873
18735
1873
18753
1673
18735
1873
1873
1873
1875
18753
1873
1875
1873
1873
12
12
44
44
6003
60035
60035
6003
60035
6003
6002
6002
6002
5822
5822
5907
5928
5928
5919
6132
6132
6212
6451
6451
6451
6452
6454
6451
6451
6450
6452
6455
6454
6439
o445
6443
6450
6452
6454
6454
6455
6425
6454
6454
6549
6557
6507
6507
5936
5936
5936
5327
5327
5327
5327
5327
5327
5327
5327
5327
5327
5103
5925
5923
5925
5923
5925
5925
5602
4954
6517
6424
6152
6048
5935
5935
5427
5408
5355
5345
5355
5345
5322
5312
5312
5312
5312
$251
5246
LONG
12855
12855
12855
12855
12855
12855
12835
12835
12835
13600
13600
13520
13605
13605
13444
14914
14914
15946
14743
14743
14743
14750
14751
14743
14743
14725
14750
14745
14751
14908
14809
14809
14915
14751
14749
14749
14749
14650
14625
14625
14403
14408
14735
14735
11307
11307
11307
11335
11333
11335
11333
11333
113335
11333
11333
113335
11333
11405
12610
12609
12609
12609
12609
12610
12243
09904
12651
12450
12122
115435
10925
10925
10826
10826
10605
10512
10525
10512
10400
10455
10455
10455
10455
10404
10401
LOCALITY
UPPER
UPPER
UPPER
UPPER
UPPER LIARD
UPPER LIARD
WATSON LAKE
WATSON LAKE
WATSON LAKE
GLACIER BAY
GLACIER BAY
HAINESe MIe 6
MOSQUITO LAKE
MOSQUITO LAKE
HAINES
MATENUSKA
MATENUSKA
HOLY CROSS
FAIRBANKS
FAIRBANKS
FAIRBANKS
COLLEGE
GOLDSTREAM CR
FAIRBANKS
FAIRBANKS
FAIRBANKS
COLLEGE
GOLDSTREAM CR
GOLDSTREAM CR
NENANA
SAM CHARLEY IS
SAM CHARLEY IS
MINTO
COLLEGE
COLLEGE
COLLEGE
GOLDSTREAM CR
HARDING L
CHENA RIVER
CHENA RIVER
CIRCLE
CIRCLE
CHATANIKA R
CHATANIKA R
LIARD
LIARD
LIARD
LIARD
DBADADAD
MOOSE LAKE
MOOSE LAKE
MOOSE LAKE
EDMONTON
EDMONTON
EDMONTON
EDMONTON
EDMONTON
EDMONTON
EDMONTON
EDMONTON
EDMONTON
EDMONTON
CALGARY
LIARD 4OT
LIARD HOT
LIARD HOT
LIARD HOT
LIARD HOT
LIARD HOT
PEACE
SIDNEY
NORMAN WELLS
KEELE RIVER
FT SIMPSON IS
HAY RIVER
CARSWELL LAKE
CARSWELL LAKE
WATERHEN RIVER
MEADOW LAKE
WASKESIU LAKE
CANDLE LAKE
CANDLE LAKE
CANDLE LAKE
NIPAWIN
PRINCE ALBERT
PRINCE ALBERT
PRINCE ALBERT
PRINCE ALBERT
TISDALE
GOLBURN
SPR
SPR
SPR
SPR
SPR
SPR
966
966
966
966
966
966
943
943
943
935
935
967
967
967
967
931
931
925
940
956
956
957
957
956
956
956
957
966
957
965
966
966
965
967
967
967
966
951
966
966
963
965
957
957
929
929
929
940
939
941
940
941
939
939
941
940
939
897
966
943
960
960
943
966
932
906
953
951
955
951
962
962
949
966
938
964
965
964
947
93a
939
938
896
939
939
COLLECTOR NAME
VOTFTMDOMOCOTOTFOKOCOTOTTKOMDMDMDDOANOAVDOCTrCCTFADDEZTIIVDAV DOD
PPCZEEEPOOVVDECHPOOPZPECI<KXTPrPoacCcoaqaoooaooooaortirTt
ESstcztezezteteE
PPPPPrPrPrPrPrPEEZEZLEEZETVUCVVPZ EE
ZSEfrvtt
je pe ae lie ai ii SS ie a ie a — i, Hi ~
= SomMM ITs
UVUvcetaezavVIEC Ea Ye >
a
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
RAUP
RAUP
RAUP
COOPER
COOPER
ARGUS
ARGUS
ARGUS
VIERECK
ANDERSON
ANDERSON
PALMER
ANDERSON
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
VIERECK
VIERECK
VIERECK
VIERECK
VIERECK
VIERECK
VIERECK
VIERECK
SCAMMAN
ARGUS
ARGUS
SPETZMAN
VIERECK
ARGUS
ARGUS
RAUP
RAuUP
RAUP
TURNER
TURNER
TURNER
TURNER
TURNER
TURNER
TURNER
TURNER
TURNER
TURNER
MACOUN
ARGUS
CLARKE
PORSILD
PORSILD
RAUP
SUDA
RAuUP
MACOUN
CODY
LINDSEY
CODY
LINDSEY
ARGUS
ARGUS
BREITUNG
HUDSON
FRASER
ARGUS
ARGUS
ARGUS
BREITUNG
FRASER
FRASER
FRASER
MACOUN
BREITUNG
BREITUNG
COL NO
5041
5030
5034
5040
5039
5033
10969
10968
10968
87
44
6750
6682
6677
8593
869
869
1257
6088
888
432
1095
1026
888
432
488
1095
5091
1026
7687
8034
8039
7647
8303
8193
8301
8027
64e4
5094
5095
966
7706
1184
1154
HERB + NO
GWA
GWA
GWA
GWA
GWA
GWA
ALA
ALA
GWA
F 824048
824029
19514
19379
F
GWA
GWA
GWA
GWA
ALA
Isc
ALA
sc
ALA
ALA
ALA
ALA
GWA
GWA
GWA
GWA
GWA
GWA
GWA
GWA
GWA
GWA
GWA
GWA
GWA
SWA
SWA
GWA
GWA
CAN
FSLC
ALA
GWA
27350
256703
2034
256704
4B25
4757
6803
6723
6733
49155
49154
49156
CAN
CAN
CAN
5AO
SAO
DAO
NAO
DAO
QAO
AO
NAO
SASK
SASK
CAN
33709
33707
94426
SASK 34000
CAN 50169
CAN 263928
CAN 263934
CAN 278796
SASK 33997
CAN
CAN
SAO
CAN 215891
NAO
CAN
SWA
NAO
SAO
SASK
SASK
GWA
SASK
SASK
HAO
NAO
SASK
SASK
CAN
DAO
DAO
70267
215892
31611
33715
28796
33983
33710
33713
13671
231
Appendix B
TaxON SPC S V HYB PROV QUAD
1873
1873
1873
1873
1673
1873
1873
1875
1873
1873
1873
1873
1873
1873
1873
1873
1873
18735
1873
1873
1873
1875
1873
1873
1873
1875
1873
1873
1873
18753
1873
1873
1673
1873
1873
1873
1873
1873
1873
1873
1873
1673
1873
1673
1873
1873
1873
1873
1873
1873
1873
1873
1873
1873
1873
1873
1873
1873
1873
1873
1673
1673
1873
1873
1873
1873
1873
wo
1873
1873
18753
77
16753
1875
1873
1873
1673
1873
1875
1875
1873
1875
1673
1873
1873
232
71 il
71 11
pes 11
71 Bt
71 ll
71 ll
71 11
71 11
71 11
71 Nt
71 ll
ce) ll
71 il
fe 118
pe 11
71 re
71 11
ae ll
71 11
71 il
71 il
71 11
71 11
71 11
71 11
71 il
al 11
71 ll
71 11
71 1i
71 11
71 11
ie 11
71 11
71 12
71 12
7fi 12
71 12
71 a
71 12
71 a2
71 12
71 12
71 12
71 12
71 12
71 12
71 12
71 12
71 12
71 12
71 12
71 12
71 12
71 12
71 12
71 12
71 12
71 12
71 12
71 14
71 14
71 14
71 14
71 14
Les 14
71 14
MACCALLIANA
73 2
73 11
73 12
MONTICOLA
Lat A 2
77 2
77 7
77 7
reef 12
77 12
77 12
LACS 12
ae 12
77 12
77 12
77 12
77 12
34
34
34
34
34
34
35
35
35
35
35
35
35
35
35
35
35
35
35
35
35
35
35
35
35
35
35
35
35
35
35
35
35
25
44
LAT
5246
5246
5246
5246
5246
5242
5237
5237
5237
$237
5237
5237
$237
5237
5237
5237
5237
5237
5237
Sest
5237
5237
5237
S257
5237
5237
5237
5237
9237
5237
5237
peat
5237
Sear
6158
6158
6003
6003
60035
6005
6003
6003
6005
60035
6005
6003
6000
6000
6000
6005
6005
6005
6002
6002
6005
6002
6003
6005
6003
60035
6448
6333
6323
6448
6400
6400
6400
9925
5450
6003
9925
5822
6156
6152
6404
6334
6335
6205
6155
6155
6045
6045
6047
LONG
10401
10401
10401
10401
10401
10400
10356
10356
10356
10356
10356
10356
10356
10356
10356
10356
10356
10356
10356
10356
10356
10356
10356
10356
10356
10356
10356
10356
10356
10356
10356
10356
10356
10356
13240
13240
12840
12840
12840
12840
12840
12840
12840
12840
12840
12840
12833
12833
128335
12843
128435
12843
12835
12835
12843
12835
12840
12840
12840
12840
14755
14330
14330
14755
14530
14530
14530
12610
10330
12840
12610
12955
12711
12122
13925
13945
13554
13618
13238
13238
13615
13615
13735
LOCALITY
GOLBURN
GOLBURN
GOLBURN
GOLBURN
GOLBURN
SYLVANIA
MCKAGUE
MCKAGUE
MCKAGUE
MCKAGUE
MCKAGUE
MCKAGUE
MCKAGUE
MCKAGUE
MCKAGUE
MCKAGUE
MCKAGUE
MCKAGUE
MCKAGUE
MCKAGUE
MCKAGUE
MCKAGUE
MCKAGUE
MCKAGUE
MCKAGUE
MCKAGUE
MCKAGUE
MCKAGUE
MCKAGUE
MCKAGUE
MCKAGUE
MCKAGUE
MCKAGUE
MCKAGUE
PELLY R VALLEY
PELLY R VALLEY
WATSON
WATSON
WATSON
WATSON
WATSON
WATSON
WATSON
WATSON
WATSON
WATSON
WATSON
WATSON
WATSON
WATSON
WATSON
WATSON
WATSON
WATSON
WATSON
WATSON
WATSON
WATSON
WATSON
WATSON
COLLEGE
LAKE
LAKE
LAKE
LAKE
LAKE
LAKE
LAKE
LAKE
LAKE
LAKE
LAKE
LAKE
LAKE
LAKE
LAKE
LAKE
LAKE
LAKE
LAKE
LAKE
LAKE
LAKE
LAKE
ALA HWY M 1237
ALA HWY M 1237
COLLEGE
CLEARWATER R
CLEARWATER R
CLEARWATER R
LIARD HOT SP
DESCHAMBAULT L
WATSON LAKE
LIARD HOT SPR
DEASE L VILLAG
RABBITKETTLE H
FT+« SIMPSON
DAWSON
OGILVIE
MAYO
CARMACKS
LOWER LAPIE R
LOWER LAPIE R
MENDENHALL R
MENDENHALL R
PINE CR
CREEK
DATE
06
06
08
05
08
06
05
05
06
08
08
08
08
08
08
05
06
06
06
90
940
939
939
939
938
939
939
939
939
939
939
939
939
939
939
939
939
939
939
936
939
939
939
939
939
939
939
939
93e
939
939
939
939
944
944
966
966
966
966
966
966
966
966
966
966
960
960
960
943
943
943
943
943
943
943
966
966
966
966
956
966
966
956
956
956
956
966
964
966
966
969
970
955
914
914
944
914
944
944
966
966
944
COLLECTOR NAME
OOOOCrTrOD<<<<KXTITIIIIIICCCOHONADADADVNGAVNDGDVD OPES FPF P PEF YFKFKFrPPPKHFPPPKrPrPrrPrPPrPrPPrYrPrreErrYrrrp
=ZzettSgoOve
DOD
=t
ITAOPYPrrcrr=zaDnAgD
s=zeezmnn
cCcecccac ccc Ke ccc oo c Con yvcC cc Ce
Lal a Sy <
6G
NuNccca
SeceogestsesecerPvrxr>zt ee zezTeztzeeE
BREITUNG
BREITUNG
BREITUNG
BREITUNG
BREITUNG
FRASER
BREITUNG
BREITUNG
BREITUNG
BREITUNG
BREITUNG
BREITUNG
BREITUNG
BREITUNG
BREITUNG
BREITUNG
BREITUNG
BREITUNG
BREITUNG
BREITUNG
BREITUNG
BREITUNG
BREITUNG
BREITUNG
BREITUNG
BREITUNS
BREITUNG
BREITUNS
BREITJNG
BREITUNG
BREITUNG
BREITUNG
BREITUNG
BREITUNG
PORSILD
PORSILD
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
CALDER
CALDER
CALDER
RAUP
RAUP
RAuP
RAuUP
RAUP
RAUP
RAUP
SUDA
SUDA
SUDA
SUDA
ARGUS
VIERECK
VIERECK
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
RIGBY
SCOTTER
CODY
EASTWOOD
EASTWOOD
ANDERSON
EASTWOOD
PORSILD
PORSILD
ARGUS
ARGUS
RAuUP
cOL ‘IO
533
§32
458
76
456
SeNe
int
108
173
394
HERB + NO
ay Ye)
DAO
NAO
DAO
CAN
NAO
DAO
3A0
DAO
NAO
BY: Ne)
NAO
DAO
DAO
NAO
NAO
DAO
9A0
HAO
Bye)
NAO
TAO
JA0
NAO
NAO
NAO
NAO
NAO
NAO
DAO
DAO
CAN 49026
CAN 49027
CAN 49058
TSC. 256562
CAN 49149
GWA
GWA
SWA
GWA
GWA
GWA
GWA
SASK
SASK
GWA
GWA
BWA
DAO
ALA
ALA
ALA
ALA
ALA
CAN
CAN
GWA
GWA
SASK 33986
SASK 33991
BWA
GWA
GWA
ALA
ALA
ALA
GWA
34002
35968
19482
19403
19483
19481
19404
278797
278795
6738
4896
22495
GWA
GWA
GWA
SWA
OTF
GWA
GWA
US 1011863
US 1011868
TSC 256716
US 1011872
ISC 256714
US 2052012
GWA
GWA
ALA
1148
19742
Specimens Cited
TAXON SPC S V HYd PROV BUA LAT LONG LOCALITY DATE
1675 77 12 40 0047 13735 PINE CR 28 06
1575 77 12 41 ©043 13503 WHITEHORSE 01 06
1873 77 12 41 6030 13415 MARSH LAKE 24 06
1873 77 le 42 6022 13351 LTL ATLIN L 19 08
18735 77 12 42 ©022 13351 LTL ATLINL 19 08
1875 77 14 68 6148 14512 WILLOW CREEK 01 08
1675 77 14 83 ©0218 14518 GAKONA 18 06
le735 77 14 83 6215 14525 GULKANA 19 06
1873 77 14 83 ©0215 14525 GULKANA 19 06
1875 77 14 85 6307 14240 TETLIN LAKE 05 08
14673 77 14 87 6308 14927 SUMMIT 24 07
16735 77 14 87 6340 14935 TEKLANIKA R
1875 77 14° 100 6451 14743 FAIRBANKS 10 05
16735 77 14 #100 6453 14750 COLLEGE 13 06
1675 77 14 100 0453 14750 COLLEGE 13 06
1875 77 14 #100 6453 14750 COLLEGE 15 08
1873 77 14° #100 6453 14750 COLLEGE 13 06
1873 77 14 #100 6453 14750 COLLEGE 15 08
1373 77 14 100 6453 14750 COLLEGE 13 06
1675 77 14 100 6454 14750 COLLEGE 07 07
1373 77 14 100 6453 14750 COLLEGE 13 06
1873 77 14 100 6453 14750 COLLEGE 15 08
1875 77 14° 100 6453 14750 COLLEGE 13 06
1873 77 14 100 6453 14750 COLLEGE 15 08
1873 77 14 100 6451 14752 UNIV EXP FARM 14 06
1875 77 14 #100 6433 14703 SALCHA SLOUGH 18 06
18735 77 14 100 6433 14703 SALCHA SLOUGH 23 06
1675 77 14° 100 6451 14743 FAIRBANKS 06
1875 77 14 100 6431 14703 SALCHA R 22 06
1875 77 14 100 6431 14703 SALCHA R 20 05
1873 77 14° 100 64468 14744 FAIRBANKS 19 05
1873 77 14 #100 ©6448 14744 FAIRBANKS 19) 05
1573 oe 14 100 6452 14752 UNIV EXP FARM 17 06
1875 77 14 101 6355 14452 TANANA R 25 07
14873 77 14 101 6412 14506 TANANA R 06 08
1873 77 14 101 6412 14506 TANANA R 22 06
1673 77 14 101 6410 14550 TANANA R 30 06
1873 77 14 103 ©6525 14340 YUKON & 06 08
1873 77 14° 104 6545 14430 EAGLE SUMMIT 15 06
1673 77 14° 120 0644 14344 BLACK 2 11 07
76 MYRTILLIFOLIA
1873 78 2 2. 5941 13632 NADAHINI RIVER 13 7
18735 78 2 5 5956 13156 ALA HWY M762 1°05
1873 78 2 5 5955 13150 SWIFT RIVER 30 06
1875 78 2 16 5831 12434 SUMMIT PASS 14 07
1675 78 2 17 5850 12235 FORT NELSON 20 05
1873 78 2 24 5705 12235 BEATTON R 26 06
18673 76 2 24 9705 12235 BEATTON R 26 06
187535 78 2 24 5705 12235 BEATTON R 13 06
1873 78 2 24 5705 12235 BEATTON R 26 06
18753 78 2 30 5603 12340 WICKED RIVER 16 07
1873 78 3 9 5838 9349 TWIN Le HILL 28 7
18753 78 7 133 ©0917 13354 E CHAN MACK R_ 01 07
1873 78 7 133 ©6915 13404 E CHAN MACK R_ 02 07
1675 78 7 133 6915 13404 E CHAN MACK R_ 02 07
1875 78 7 241 6517 12651 NORMAN WELLS 22 il
18753 78 7 276 6454 12534 FORT NORMAN ot:
1873 78 7 307 6©338 12837 JUNE LAKE ol a7
1873 78 7 308 6330 12647 HAYHOOK LAKE 6 86
1873 78 7 339 ©6241 12750 GRIZZLY BEAR L 10 8
1875 78 7 339 ©6205 12735 BRINTNELL L 23 06
1873 78 7 339 6205 12735 BRINTNELL L 23 06
1873 78 7 341 6242 12337 MT FLETT 1. 8
1875 78 7 347 6237 11130 TALTHEILEI NAR 09 07
18735 78 7 367 6157 12713 RABBITKETTLE L 10 8
18673 78 7 368 6137 12544 Se NAHANNI Re 29 6
13875 78 7 369 6152 12320 LTL DOCTOR Le o 8
1675 78 7 369 6158 12325 CLI LAKE ay 38
1875 78 7 421 6158 12815 FLAT RIVER 2:8
1875 78 11 1 5948 10833 TAZIN RIVER 10 07
1873 78 12 15 6404 13925 DAWSON 27 05
1873 78 12 22 6345 13554 MAYO 06 08
1875 78 12 30 ©6249 13043 CANOL RD M245 21 08
1675 78 12 32 6159 14033 WHITE RIVER 21 07
1873 78 12 32 6122 13859 BURWASH LDG 13 07
1875 78 12 32 6118 13530 DAWSON HWY 436 25 06
1873 78 12 35 6140 13302 CANOL RD M105 10 06
1875 78 12 35 6155 13228 CANOL RD M132 17 06
1673 78 le 35 6155 13228 CANOL RD M132 17 06
1875 78 12 40 6045 13615 MENDENHALL R 30 06
18735 78 12 40 6047 13735 PINE CREEK 25 06
1873 78 12 40 6047 13735 PINE CREEK 17 06
1875 78 12 40 6047 13735 PINE CREEK 17 06
1873 78 12 40 6047 13735 PINE CREEK 17 06
1875 78 12 40 6047 13735 PINE CREEK 17 06
1875 78 12 40 6047 13735 PINE CREEK 17 06
1873 78 12 41 6043 13503 WHITEHORSE 31 05
1875 78 12 41 6043 13503 WHITEHORSE 01 06
944
944
966
943
943
967
944
957
957
963
967
940
957
957
957
957
957
957
966
957
O57
957
957
956
922
922
933
966
966
965
965
964
965
966
966
964
964
951
957
967
960
968
943
949
943
943
943
943
932
958
957
557
957
953
940
967
967
967
939
939
961
927
967
970
961
961
967
914
914
944
944
944
944
966
944
944
944
966
948
944
944
944
944
944
949
944
COLLECTOR NAME
WCFrTrrrrrrererrroogmgOgOagagDOaODDOODOOCETrTrADOCETrILTYYPL
PWCTITIIIO SP rPPvICrCcCrPnAZEGAZLTLELILESZEPzZeESZBAzEOIIIIICIMCO®e
NESSCSCESCSCEMANS Suv
=
a 2. Ce Sir 3
ZAC LE LA SELLA VS
@arrmpvpaPprarrrrvF? yr RrPPGL
Zanes eeegsrrre
©
r
Zqcaetac £C ££ £LGAG (er
RAUP
PORSILD
WELSH
RAuUP
RAUP
VIERECK
ANDERSON
ARGUS
ARGUS
SCHNEIDER
VIERECK
VIERECK
ANDERSON
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
MURIE
MURIE
PALMER
VIERECK
VIERECK
VIERECK
VIERECK
VIERECK
VIERECK
VIERECK
VIERECK
VIERECK
VIERECK
BUCKLEY
SHETLER
ARGUS
CALDER
WELSH
RAUP
GILLETT
RAUP
RAUP
RAUP
RAUP
RAUP
ARGUS
CODY
CODY
CODY
CODY
DUTILLY
CODY
CODY
CODY
RAuUP
RAUP
CODY
RAuUP
CODY
SCOTTER
CODY
CODY
SPICER
HARPER
EASTWOOD
ANDERSON
PORSILD
ANDERSON
RAUP
WELSH
PORSILD
PORSILD
PORSILD
ARGUS
RAUP
RAUP
RAUP
RAuP
RAUP
RAuP
GILLETT
PORSILD
cOL NO
120°8
9157
5504
11389
11389
RUST
HE23 + NO
SASK 8683
'IS 2051866
GWA
ALA 19743
SASK 8479
6WA
TSC 256715
ALA 8416
GWA
ALA 264619
GWA
FSLC 291
TSC 256717
ALA 6710
ALA 6714
SLA 6730
ALA 6736
ALA 6712
GWA
SWA
SWA
SwA
GWA
GWA
GWA
JS 1119995
US 1119981
ALA 5271
SWA
SWA
FSLC
FSLC
FSLC 313
SWA
GWA
GWA
FSLC 302
FSLC 287
SWA
ALA 3967
5WA
GWA
SWA
ALA 19841
TSC 256141
ALA 19840
ALA 19839
ALA 19865
SWA
F 719718
CWA
DAO 32696
SAO 32694
NAO 32695
NAO
NAO
DAO
NAO
AO
ALA 14776
ALA 14777
AO
ALA 14605
5A0
GWA
NAO
DAO
SAO
GWA
A
Isc 2567390
SH
TSC 256738
ALA 19772
GWA
TSC 256138
TSC 256139
6H
GWA
ALA 19478
ALA 19866
ALA 19859
ALA 19843
GWA
GWA
Isc 256142
6H
233
Appendix B
TAXON SPC S V HYB PROV QUAD
1873
1873
1873
1873
1873
1873
1873
1873
1873
1873
1873
1873
1673
1873
1873
1873
1673
1873
1873
1873
1873
1473
1673
1873
1873
1673
1673
1873
1873
1873
1873
18673
1873
1873
1873
1873
1873
1673
1873
1873
1473
1873
1873
1873
1873
1673
1873
1873
1873
1673
1873
1873
1873
1873
1873
1873
1873
1873
1873
1873
1873
1873
1673
1873
1873
1873
60
1873
1873
1873
83
1873
1873
1873
1673
1873
1873
1673
1673
1873
1873
1873
1673
1873
1873
234
78 12
78 12
78 12
78 12
78 12
78 l2
78 12
78 12
78 12
78 12
78 ie
78 12
78 12
78 12
78 12
78 14
78 14
78 14
78 14
78 14
78 14
78 14
78 14
78 14
78 14
78 14
78 14
78 14
78 14
78 14
78 14
78 14
78 14
78 14
78 14
78 14
78 14
78 14
78 14
78 14
78 14
78 14
78 14
78 14
78 14
78 14
78 14
78 14
78 14
78 14
78 14
78 14
78 14
78 14
78 14
78 14
78 14
78 14
78 14
78 14
78 14
78 14
78 14
78 14
78 14
78 14
NUMMULARIA
60 14
80 14
60 14
1 OVALIFOLIA VAR»
63 01 12
83 01 14
8301 14
83 01 14
63 O1 14
8301 14
63 O01 14
83 1 14
83 1 14
63 01 14
63 01 14
83 01 14
63 01 14
63 01 14
LAT LONG
42 6009 13245
42 6009 13245
42 6009 13245
42 ©6054 13258
42 6022 13351
42 6022 13351
42 ©6006 13227
42 6016 13255
43 6005 13040
43 6001 13111
44 6003 12840
44 6003 12840
44 6003 12840
44 ©6002 12900
44 6007 12844
68 0148 14512
69 6156 14710
69 6156 14710
69 6156 14710
69 6156 14710
69 6156 14710
80 6233 15336
83 6225 14451
83 6225 14451
83 6208 14528
85 ©0217 14516
83 6206 14542
83 6206 14542
83 6206 14542
85 ©6317 14232
85 ©3530 14220
85 6355 14209
86 6350 14450
86 63555 14420
86 6355 14452
87 6352 14858
87 0350 14925
87 6344 14855
87 ©6335 14845
89 6315 15520
100 6448 14750
100 6452 14750
100 6452 14752
100 6451 14752
100 6452 14750
100 0452 14750
100 6452 14750
100 6452 14752
100 6452 14750
100 6452 14750
100 6452 14752
100 6450 14750
100 6452 14750
100 6448 14750
100 6448 14750
100 6448 14750
100 6448 14750
100 6448 14750
100 6448 14750
100 6448 14750
101 6402 14534
101 6405 14515
102 6410 14136
102 6405 14147
119 6603 14417
120 6644 14334
38 5710 17015
38 $710 17015
38 5710 17015
OVALIFOLIA
2 6930 13920
16 5212 17406
18 5207 17430
22 5310 16845
22 5310 16845
24 5408 16546
24 5408 16546
25 9459 16228
25 5459 16228
27 5540 15950
26 5520 16030
28 5507 16144
28 95543 16011
30 5618 15824
LOCALITY
TESLIN
TESLIN
TESLIN
NISUTLIN R CP
ie Aen NYC
ETC AGEIN C
ALA HWY M793
TESLIN LAKE
RANCHERIA
ALA HWY M733
WATSON LAKE
WATSON LAKE
WATSON LAKE
ALA HWY M658
WATSON LAKE
WILLOW CREEK
GLENN HWY M128
GLENN HWY M128
GLENN HWY M128
GLENN HWY M128
GLENN HWY M128
FAREWELL LAKE
TOKSLA HWY M20
TOKSLA HWY M20
RICH Hwy Ml22
GAKONA
GLENNALLEN
GLENNALLEN
GLENNALLEN
ALA HWY M1247
TAYLOR HWY M19
TAYLOR HwY MSO
GERSTLE RIVER
LOWER BERRY CR
HEALY LAKE
HEALY
SAVAGE R CAMP
MCKINLEY PARK
CANTWELL
NIXON FORK
FAIRBANKS
COLLEGE
SMITH LAKE
UNIV EXPL FARM
COLLEGE
COLLEGE
COLLEGE
SMITH LAKE
COLLEGE
COLLEGE
SMITH LAKE
COLLEGE
COLLEGE
FAIRBANKS
FAIRBANKS
FAIRBANKS
FAIRBANKS
FAIRBANKS
FAIRBANKS
FAIRBANKS
DELTA JUNCT
GERSTLE RIVER
FRANKLIN
CHICKEN
YUKON RIVER
BLACK RIVER
ST PAUL
ST PAUL
ST PAUL
ISLAND
ISLAND
ISLAND
FIRTH RIVER
NAZAN 3AY
ATKA ISLAND
UMNAK ISLAND
UMNAK ISLAND
AKUTAN
AKUTAN
COLD BAY
COLD BAY
STEPOVAK BAY
SAND POINT
DOLGOI ISLAND
STEPOVAK BAY
CHIGNIK
956
956
956
944
943
943
966
968
948
960
946
966
966
968
968
967
944
944
944
947
947
949
944
944
944
944
967
967
965
944
965
963
957
957
965
939
928
959
964
950
940
957
956
956
957
957
957
965
965
965
965
952
952
933
931
931
933
963
963
963
951
963
941
964
965
957
941
941
925
953
949
944
962
962
933
934
971
899
899
931
899
945
COLLECTOR NAME
WFreeCcCrerrrrrrroao~wmwce cece OOQDOODOCEZEFOXCFFFCHKCHWFFOCCCEMMCECCOCTMNUNOMODCCCYHNTLESPADD|A
mre
KFOTNnOSSEKHKOZTZEOIM
UG ee Se OS OTe ee ee,
GMUrsaSPuvIVwv=xs
Q0rv->
| eeeil pest oil roel ac, RC a AS AK ilo 1 iyi
=r USTcy ye SPCC Cc
ARGUS
ARGUS
ARGUS
PORSILD
RAUP
RAUP
WELSH
WELSH
ANDERSON
CALDER
ANDERSON
ARGUS
ARGUS
WELSH
WELSH
VIERECK
ANDERSON
ANDERSON
ANDERSON
SCAMMAN
SCAMMAN
DRURY
ANDERSON
ANDERSON
ANDERSON
ANDERSON
VIERECK
VIERECK
WELSH
ANDERSON
HARMS
NAVA
SPETZMAN
SPETZMAN
VIERECK
ANDERSON
MEXTA
PETRIDES
VIERECK
DRURY
ANDERSON
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
HARMS
HARMS
HARMS
HARMS
KESSEL
KESSEL
PALMER
PALMYER
PALMER
PALMER
VIERECK
VIERECK
VIERECK
CopY
VIERECK
ANDERSON
HARMS
VIERECK
SHETLER
COLE
J COLE
JOHNSTON
MCEWEN
MILLER
YORK
JOHNSON
JOHNSON
GEIST
NORBERG
WILLIAMS
WILLIAMS
PALACHE
COVILLE
MASON
PALACHE
NORBERG
cOL NO
272
272
273
10782
11390
11390
5478
7616
109428
24853
9948
5063
5043
7474
7yuug
8455
R4ULD
RYSO
BY49
4524
4524
une
8710
8710
RYO
B63
Rene
R210
4306
HERB
ALA
GWA
GWA
6H
ALA
GWA
GWA
GWA
TSC
GWA
TSC
GWA
GWA
GWA
GWA
GWA
GWA
TSC
rsc
GWA
SASK
CAN
GWA
TSC
TSsc
TSC
GWA
SWA
Tsc
TSsc
ALA
ALA
ALA
ALA
FSLC
SiG
ALA
ALA
PSUC
CAN
ysc
ALA
GWA
GWA
GWA
GWA
GWA
ALA
ALA
ALA
ALA
ALA
ALA
ALA
ALA
ALA
ALA
FSLC
site
ESLC
GWA
FSLC
ALA
ALA
ESEC
ALA
msc
WIS
NA
CAN
MICH
1Ssc
wIs
wIS
ALA
1sc
+ NO
4821
19863
256732
256731
256741
256729
34212
256728
256727
256743
246052
256739
32590
23942
6846
6845
256725
10204
27163
296
256745
6824
32630
32635
32634
32586
22863
22864
5269
5971
5974
5272
22
23
536
32650
3965
256642
319618
226062
256237
29713
253240
375470
373507
375471
Specimens Cited
TaxON SPC
1873 83
1873 83
1673 83
1873 83
1873 83
1873 83
1873 83
1673 83
1873 83
18735 83
1873 83
1673 83
1873 83
1873 83
1673 83
1873 83
1873 83
1873 83
1873 83
1873 83
1873 83
1873 83
1873 83
1673 83
1873 83
18735 83
1673 83
1873 83
1873 83
1873 83
1873 83
1875 83
1873 83
1673 83
1873 63
1873 83
1673 83
1873 83
1875 83
1873 83
1873 83
1673 83
1473 83
1673 83
1873 83
13873 83
1873 83
1673 63
1873 83
1873 83
18673 83
1873 83
1873 63
15873 83
1873 83
1873 B83
1673 83
1873 83
1873 83
1673 83
1673 63
1573 63
1873 83
1673 83
1873 63
1873 63
1673 83
1673 83
1873 83
1673 83
1673 83
1673 83
1673 83
1873 63
1873 63
1673 83
1873 83
1873 83
1673 83
1673 83
1873 83
1873 83
1673 63
1573 83
1673 83
1873 83
1573 63
1873 83
1373 83
1673 83
1673 83
oooooococococcoecocoeococco°coococoococoococccoccacacocceo
oooocoeocoocoooococeococoeccoedce°dooc0cecocooao0cocoe0cd0ao
oooooo0cococoo
Cl eel al al eel cll el el el el cl ll ll ll el el cll ll el cl cl cl cc a a ee nr eae
V HYB PROV QUAD LAT
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
35
35
35
36
36
38
41
42
55
57
62
62
62
62
92
93
a5
129
129
129
129
129
129
129
130
130
130
138
138
1338
138
138
138
139
139
139
145
145
145
146
147
147
147
147
147
147
148
14%
149
149
151
151
151
i51
151
151
151
151
151
151
151
151
151
151
5705
5705
5741
5734
5748
5710
5843
5838
5949
6004
6026
6028
6026
6028
6330
6319
6342
6342
6342
6342
6342
6342
6342
6305
6430
6430
6430
6430
6442
0458
6512
6512
6512
6758
6758
o82l
68el1
6853
6806
6821
6810
6810
6810
6810
6810
6810
6821
6819
6845
6821
6821
6355
0855
6855
695U
6950
6950
6950
6953
6946
7000
6917
6924
6946
6946
60946
7020
7040
7040
7040
7040
704uU
7040
7047
7057
7027
7027
7010
7010
7010
7010
7010
7010
7010
7010
7010
7010
7010
7010
7010
7010
LONG
15425
15425
15529
15602
15632
17015
15652
15552
16608
16716
15028
15028
15028
15028
16200
17127
17029
17029
17029
17029
17029
17029
17029
16849
16530
16530
16530
16530
16202
16805
16645
16645
16645
16432
16432
16627
16627
16615
16545
16627
16540
16540
16540
16540
16540
16540
16627
16640
16600
16627
16627
16437
16437
16436
14220
14220
14220
14220
14215
14137
14454
14600
14608
163035
16305
16304
16152
15655
15655
15655
15655
15655
15655
15502
15540
15007
15007
14650
14650
14650
14650
14650
14650
14650
14650
14650
14650
14650
14650
14650
14650
LOCALITY
OLGA BAY
OLGA BAY
PUALE BAY
KANATAK
BECHAROF LAKE
ST PAUL ISLAND
SAVONOSKI
NAKNEK LAKE
DUCHIKTHLUK R
DOOKSOOK LG
SKILAK LAKE
SKILAK LAKE
SKILAK LAKE
SKILAK LAKE
WHALE ISLAND
Sw CAPE
SAVOONGA
SAVOONGA
SAVOONGA
SAVOONGA
SAVOONGSA
SAVOONGA
SAVOONGA
PUNUK ISLAND
NOME
NOME
NOME
NOME
MOSES POINT
KING ISLAND
PORT CLARENCE
PORT CLARENCE
PORT CLARENCE
KIVALINA
KIVALINA
POINT HOPE
POINT OPE
CAPE LISBURNE
OGOTORUK CREEK
POINT HOPE
OGOTORUK CREEK
OGOTORJK CREEK
OGOTORUK CREEK
OGOTORJK CREEK
OGOTORJK CREEK
OGOTORJK CREEK
POINT HOPE
POINT HOPE
UKINYI«K CREEK
POINT HOPE
POINT HOPE
PITMAGEA R
PITMAGEA R
CAPE SABINE
NUVAGAPAK PT
NUVAGAPAK PT
NUVAGAPAK PT
NUVAGAPAK PT
NUVAGAPAK LG
ICY REEF
COLLINSON PT
CANNING RIVER
CANNING RIVER
POINT LAY
POINT LAY
POINT LAY REEF
ICY CAPE
MEADE RIVER
MEADE RIVER
MEADE RIVER
MEADE RIVER
MEADE RIVER
MEADE RIVER
HALF MOON 3
OARLOCK IS
COLVILLE DELTA
COLVILLE DELTA
BULLEN
BULLEN
BULLEN
BULLEN
BULLEN
BULLEN
BULLEN
BULLEN
BULLEN
BULLEN
BULLEN
BULLEN
BULLEN
BULLEN
DATE
940
940
904
933
949
941
919
918
965
965
951
951
951
951
931
933
933
933
933
933
933
933
933
934
938
954
954
926
950
938
901
901
901
960
960
938
938
938
966
956
959
959
959
959
959
959
951
960
960
966
966
B57
957
959
966
966
966
966
965
959
914
947
947
938
947
951
931
958
952
952
952
952
952
962
951
901
901
966
966
966
966
966
966
966
966
966
966
966
966
966
966
COLLECTOR NAME
MPOODADODANOANAMDANONANVONOND UNAANAHOMDODAONDDDIADCLEANANVTOAANONCCHUNFTCTMPE PHP SEE YP PEDCCCEPPNNNCWDEPNANDACOCDODOOCOOOOICCCCOOVPFAOCOMM
> Pr p
VECEEZTELCEE
aa fi bi ek
Z£=eet
LOOFF
LOOFF
PIPER
GEIST
RAUP
COLE
MILLER
HAGELBARGER
BOS
BOS
CALDER
CALDER
CALDER
CALDER
MASON
GEIST
GEIST
GEIST
GEIST
GEIST
GEIST
GEIST
GEIST
GEIST
ANDERSON
HELLER
HELLER
PORSILD
SIGAFOOS
ANDERSON
WALPOLE
WALPOLE
WALPOLE
BUCKNELL
BUCKNELL
ANDERSON
ANDERSON
ANDERSON
ARGUS
BERRY
JOHNSON
JOHNSON
JOHNSON
JOHNSON
JOHNSON
JOHNSON
SCAMMAN
VIERECK
VIERECK
WELSH
WELSH
CANTLON
CANTLON
SHETLER
AGUS
ARGUS
ARGUS
ARGUS
HENRY
LEWIS
JOHANSEN
MCGREGOR
SPETZMAN
ANDERSON
BLACK
CHAMBERS
MASON
WARD
WARD
WARD
WARD
WARD
WARD
CHESE MORE
WIGGINS
SCHRADER
SCHRADER
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
COL NO
1275
1276
4596
190
247
4988
4ORT
4988
4ORT
6059
1n9
19%
112
105
111
102
113
31788
1084
1084
1330
3601
36078
1826
1672
1672
36
36
4602
3787
4496
5945
SeNe
68
2n7
Tes
723
68
on7
SeNe
4g22
4420
5R30
5830
57149
57209
3332
5277
HERS + NO
A
A
US 421036
ALA 29714
US 2176361
ISC 256238
YS 1072627
US 1072871
ALA 32289
ALA 32288
NAO 32783
NAO 32782
US 2231326
US 2331325
6H
ALA 29782
ALA 29387
ALA 29379
ALA 29414
ALA 29415
ALA 29385
ALA 29380
ALA 29386
ALA 29044
ISC 253233
ALA 24328
ALA 27065
GH
5WA
TSC 255908
NS 378940
US 2440700
US 378784
ALA 26594
ALA 26594
TSC 253223
TSC 256226
TSC 2562731
GWA
SWA
ALA 9205
ALA 9199
ALA 9198
GWA
SWA
GWA
SWA
ALA 13222
ALA 13238
GWA
TSC 253604
GH
6H
“MICH
GWA
GWA
SWA
SWA
ALA 30384
CAN 296361
CAN 50113
“ICH
TSC 256230
TSC 256227
TSC 256229
GH
US 1789440
64
6H
'IS 2312307
JS 2312331
YS 2312305
US 2312306
ALA 29361
GH
US 377533
US 377532
GWA
GWA
SWA
SWA
5WA
GWA
SwA
GWA
GWA
GwA
SWA
GWA
GWA
SWA
235
Appendix B
TAXON SPC S V HYB PROV wUAD LAT LONG
1675 63 01 14 #151 7010 14650
1875 63 01 14 151 7010 14650
1675 63 01 14 151 7010 14650
1673 83 01 14 151 7010 14650
1873 63 0 1 14 151 7010 14650
1875 68301 14 151 7010 14650
1873 83 01 14 151 7010 14650
1873 63 0 1 14 151 7010 14650
1673 83 01 14 151 7010 14650
1573 63 0 1 144 151 7010 14650
1673 83 01 14 151 7010 14650
1673 83 01 14 151 701v0 14650
1873 63 01 14 151 7010 14650
1873 83 01 14 151 7010 14650
1873 63 01 14 151 7010 14650
1873 63 01 14 151 7010 14650
1873 63 01 14 151 7010 14650
14673 83 0 1 144 151 7010 14650
138735 83 0 1 14 151 7010 14650
1875 63 01 14 151 7010 14650
1873 63 0 1 14 151 7010 14650
1873 83 1 14 151 7010 14650
1675 8301 14 151 7001 14510
1673 83 01 Pa IS2 7002 14510
1873 83 0 1 14 #151 7001 14428
1873 83 01 14 152 7008 14315
1675 83 0 1 14 (153 7125 15628
1875 83 01 14 #153 7057 15540
1873 83 0 1 14 “853 7kO8 15705
63 2 OVALIFOLIA VARe ARCTOLITORALIS
18735 83 0 2 7 133 6929 13515
1873 63 02 7 4335 6929 13525
1673 83 02 ie 2 ©6900 13730
18735 85 0 2 14 91 6352 160047
1673 63 0 2 14 91 ©6352 16047
1873 83 02 14 94 6430 16525
1875 83 0 2 14 94 6430 16525
16735 83 0 2 14 94 6428 16517
1873 83 02 14 94 6430 16525
1873 83 02 14 95 6426 16500
1873 63 0 2 14 95 6426 16500
1873 83 02 14 111 6516 16622
1675 83 0 2 14 111 6516 16622
1873 83 02 14 #111 6512 16645
18735 63 02 14 113 ©6604 16242
1673 83 02 14 #113 6654 16235
1873 683 0 2 14 113 6654 16235
1873 63 0 2 14 113 6655 16240
1673 83 0 2 14 113 6655 16240
1873 63 02 14 113 6655 16240
1673 63 0 2 14 #113 6655 16240
1873 63 02 14° 113 6655 16240
1675 63 02 14° #126 6758 16432
1673 83 0 2 14° #129 6806 16545
1873 83 0 2 14 129 6806 16545
18673 63 0 2 14 129 6©806 16545
1673 83 0e2 14 #129 6806 16545
1873 63 0 2 144 #129 6806 16545
1873 83 02 14 129 6806 16545
18735 83 2 14 129 6806 16545
1673 83 02 14 129 6806 16545
1673 83 02 14 129 6806 16545
1673 63 0 2 144 129 6820 16651
1873 83 0 2 14 130 6855 16430
1673 83 0 2 14° #145 6©946 16303
18735 83 02 14° #145 6946 16303
16735 63 0 2 14° #145 6946 16303
83 3 OVALIFOLIA VAR> GLACIALIS
1873 683 0 3 14 139 6959 14454
1873 683 0 3 14 i39 6959 14454
1673 63 0 3 14 147 7040 15655
1873 83 0 3 14% 147 7040 15655
1673 83 0 3 14 #148 7032 15548
1673 83 0 3 14 "153 97 t25 15650
1873 63 0 3 14 153 7125 15630
1873 83 0 3 14 153 7125 15630
1873 83 0 3 1% 153 7125 15650
1873 683 0 3 14 153 ‘7125 15630
1673 83 0 3 1% 153 7125 15630
18735 63 0 3 14 153 7125 15630
1873 83 0 3 14 153 7125 156350
1873 83 0 3 14° 153 7125 15650
1673 683 0 3 149° 453 =+7125°15650
1873 63 0 3 14 153 7125 15630
1673 83 0 3 14 153 7125 15630
236
LOCALITY
BULLEN
BULLEN
BULLEN
BULLEN
BULLEN
BULLEN
BULLEN
BULLEN
BULLEN
BULLEN
BULLEN
BULLEN
BULLEN
BULLEN
BULLEN
BULLEN
BULLEN
BULLEN
BULLEN
BULLEN
BULLEN
BULLEN
KONGANEVIK PT
KONGANEVIK PT
ANDERSON POINT
MARTIN POINT
POINT 3ARROW
DEASE INLET
BARROW
KENDALL ISLAND
KENDALL ISLAND
SHINGLE POINT
UNALAKLEET
UNALAKLEET
NOME
NOME
FT DAVIS
NOME
CAPE NOME
CAPE NOME
TELLER
TELLER
PORT CLARENCE
DEERING
KOTZEBUE
KOTZEBUE
KOTZEBUE
KOTZEBUE
KOTZEBUE
KOTZEBJE
KOTZEBJE
KIVELINA
OGOTORUK CR
OGOTORUK CR
OGOTORUK CR
OGOTCRUK CR
OGOTORUK CR
OGOTORUK CR
OGOTORUK CRe
OGOTORUK CR
OGOTORUJK CR
POINT HOPE
PITMEGEA R
POINT LAY
POINT LAY
POINT LAY
CAMDEN BAY
CAMDEN BAY
MEADE RIVER
MEADE RIVER
TOPAGORUK R
POINT BARROW
POINT BARROW
POINT BARROw
POINT BARROW
POINT 3ARROW
POINT 3ARROW
POINT BARROW
POINT S8ARROW
POINT BARROW
POINT 3ARROW
POINT BARROW
POINT BARROW
DATE
08
08
06
08
08
08
08
08
0A
08
08
08
08
oA
08
08
08
08
08
08
08
8
07
07
06
O74
06 §
08
07
966
966
COLLECTOR NAME
ATH MNNAVAVDAANAVANAVAAVADDVNANANADNDNDDADANANADNONODD
“et
NMCUMAQDAAADANVDAGODCUHUMMACCC TMMMMMABDOTIYSMM
DAQMDODODDOADODOAANANOHKHODONnN
ZSEetecetttzteztzeriete
At
vuad
ZEZCESCZAZE VS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
AGUS
ARGUS
ARGUS
JOHANSEN
JOHANSEN
SPETZMAN
JOHANSEN
MURDOCK
WIGGINS
WIGGINS
MCEWEN
MCEWEN
PORSILD
JOHNSTON
JOHNSTON
JONES
KELLOGG
PEGAU
SCAMMAN
LEPAGE
LEPAGE
SCAMMAN
SCAMMAN
WALPOLE
ANDERSON
ANDERSON
ANDERSON
RAUSCH
SCAMMAN
SCAMMAN
WELSH
WELSH
ANDERSON
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
SCAMMAN
SHETLER
ANDERSON
GOODMAN
GOODMAN
JOHANSEN
JOHANSEN
WARD
WARD
WIGGINS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
cOL NO
5702
5741
5746
5740
5745
5789
5774
5791
5707
5777
5796
5700
5793
57A1
5772
5728
5724
5775
5a11
5776
5743
5780
P28
ROA
3579A
1368
A5
128N2
12500
HERB + NO
mwa
WA
CWA
GWA
GWA
BWA
SWA
GWA
5WA
GWA
SWA
GWA
BWA
GWA
AWA
aWA
GWA
TWA
GWA
GWA
GWA
GWA
CAN 50119
US 1243596
CAN 296360
CAN 50115
6H
US 2312904
GH
c
CAN 219336
CAN 219337
CAN 50110
NY
US 2034617
ISC 146465
NS 1366547
GWA
6H
DAO 32785
US 2068316
GH
GH
US 378785
TSC 256232
TSC 256243
YS 2440684
US 2349384
SWA
GWA
SWA
GWA
ISc 256242
GWA
GWA
GWA
GWA
GWA
GWA
SWA
GWA
GWA
SY
5H
TSC 256228
5H
US 1602441
CAN 50116
CAN 50117
US 2312329
US 2312330
US 2263953
GWA
GWA
GWA
SWA
SWA
GWA
GWA
SWA
SWA
GWA
GWA
SWA
Specimens
TAaxON SPC
1873 83
1673 83
1673 83
1673 83
14673 83
1673 83
1673 83
1473 83
1873 83
1673 83
1873 63
1873 83
1673 83
1573 83
1673 63
1673 83
1873 83
1573 83
1873 63
1873 63
1673 83
1873 83
1873 63
1873 83
1873 83
1873 83
1473 83
1873 83
1673 83
1873 83
1573 83
1873 83
1873 83
1873 83
1873 83
1873 83
1873 83
1873 83
1873 83
1673 83
1873 83
83 4
1873 83
1873 83
1873 83
1873 83
1873 83
1873 83
1873 83
1673 83
1873 83
1875 83
1873 83
1873 83
13873 83
1873 83
1873 83
1673 83
1873 83
1873 63
1873 83
1873 83
1873 83
1873 83
18735 83
1873 83
1873 63
1873 83
1873 83
1873 83
1873 83
1873 83
1873 83
1673 83
1873 83
1873 83
1673 83
1673 83
18673 83
1873 83
1673 83
1873 83
1873 83
1873 83
1873 83
1873 83
1873 83
1873 83
Cited
S V HYB PROV WA) LAT LONG LOCALITY
0 3 14 155 7125 15630 POINT 3ARROwW
0 3 14 153 7125 15630 POINT 3ARROw
0 3 14 153 7125 15630 POINT BARROW
03 14 153 7125 15630 POINT BARROW
0 3 14 153 7125 15630 POINT 3ARROW
0 3 14 153 7125 15630 POINT 3ARROW
0 3 14 153 7125 15630 POINT BARROW
03 14 153 7125 15630 POINT BARROw
0 3 14 153 7125 15630 POINT BARROW
03 14 153 7125 15630 POINT 3ARROW
0 3 14 153 7125 15630 POINT 3ARROW
0 3 14 153 7125 15630 POINT 3ARROW
0 3 14 153 7125 15630 POINT 3ARROW
0 3 14 153 7125 15630 POINT 3ARROW
0 3 144% 153 7125 15630 POINT 3ARROW
3 14 153 7125 15630 PT+«BARROW
0 3 144 153 7119 15643 BARROW
0 3 14 153 7119 15643 BARROW
0 3 14 #153 7119 15643 BARROW
03 14 153 7119 15643 BARROW
0 3 14 #153 7120 15640 BARROW
03 14 153 7125 15630 POINT 3ARROW
0 3 14 153 7125 15630 POINT BARROW
0 3 14 153 7125 15630 POINT 3ARROW
03 14 153 7125 15630 POINT BARROW
0.3 14 153 7119 15643 BARROW
0 3 14 #153 7119 15643 BARROW
03 14 153 7119 15643 BARROW
0.3 14% 153 7119 15643 BARROW
0 3 14 153 7119 15643 BARROW
a 14 #153 7119 15643 BARROW
a 3 14% 153 7124 15629 NUWUK
0 3 14 153 7124 15629 NUWUK
0 3 14 153 7124 15629 NUWUK
0 3 14 #153 7124 15629 NUWUK
0 3 14 153 7119 15643 BARROW
0.3 14 1535 7123 15629 NUWUK
0 3 14% 153 7123 15629 NUWUK
0 3 144 #153 7124 15629 NUWUK
03 144 153 7124 15629 NUWUK
0 3 14 #153 7123 15629 NUWUK
OVALIFOLIA VAR»* CYCLOPHYLLA
04 14 15 51468 17947ERAT ISLAND
04 14 15 5132 17900E AMCHITKA IS
04 14 17 5205 17608 GR SITKAN IS
04 14 17 5205 17608 GR SITKAN IS
04 14 17 5156 17644 ADAK ISLAND
04 14 17 5156 17644 ADAK ISLAND
04 14 21 5256 16835 NIKOLSKI
04 14 23 5335 16650 UNALASKA
04 14 23 5352 16632 UNALASKA
04 14 27 5505 15929 HALL ISLAND
04 14 27 9505 15929 HALL ISLAND
04 14 27 5505 15929-HALL ISLAND
04 14 27 5538 15940 FOX BAY
04 14 29 5520 16248 IZEMBEK LAGOON
04 14 38 5710 17015 ST PAUL IS
04 14 36 5710 17015 ST PAUL IS
04 14 38 5710 17015 ST PAUL IS
04 14 38 5700 17000 SEAL ISLAND
04 14 38 5710 17015 ST PAUL IS
04 14 38 5710 17015 ST PAUL IS
04 14 38 5710 17015 ST PAUL IS
04 14 38 5710 17015 ST PAUL IS
04 14 38 5710 17015 ST PAUL IS
04 14 38 5710 17015 ST PAUL IS
04 14 38. 5710 17015, ST PAUL. IS
04 14 38 5635 16935 ST GEORGE IS
04 14 38 5710 17015 ST PAUL IS
04 14 38 5710 17015 ST PAUL IS
04 14 38 5710 17015 ST PAUL IS
04 14 38 5710 17015 ST PAUL IS
04 14 38 5710 17015 ST PAUL IS
4 14 38 5707 17016 ST PAUL ISLAND
c 4 14 38 5710 17015 ST PAUL IS
04 14 38 5710 17015 ST PAUL IS
04 14 38 5710 17015 ST PAUL IS
04 14 38 5710 17015 ST PAUL IS
04 14 38 5710 17015 ST PAUL IS
04% 14 38 5710 17015 ST PAUL IS
04 14 38 5710 17015 ST PAUL IS
04 14 38 95710 17015 ST PAUL IS
04 14 56 6024 17238 ST MATTHEW IS
04 14 56 ©6024 17238 ST MATTHEW IS
04 14 56 6024 17238 ST MATTHEW IS
04 14 56 0024 17238 ST MATTHEW IS
04 14 56 ©024% 17238 ST MATTHEW IS
04 14 56 6024 17238 ST MATTHEW IS
DATE
08
08
08
08
07
07
07
07
07
07
07
07
07
07
07
8
07
07
07
07
07
06
06
08
07
07
07
07
07
07
08
08
08
08
06
06
06
08
08
06
966
966
966
966
966
966
966
966
966
966
966
966
966
966
966
966
947
947
947
947
961
88
88
947
947
95Nn
950
950
950
950
950
950
950
950
950
950
952
952
950
950
952
932
967
950
950
950
949
962
926
926
899
899
899
913
925
938
899
899
895
941
899
963
899
899
899
932
924
897
914
914
914
914
897
897
914
890
890
890
893
879
912
899
899
899
966
966
966
COLLECTOR NAME cOL “0
G Ww ARGUS 59R2
G “ ARGUS 5923
G W ARGUS 59A0
G w ARGUS 5992
G w ARGUS 5153
G w ARGUS 5150
G W ARGUS 5144
G W ARGUS 5122
G w ARGUS 5142
G w ARGUS 5146
G w ARGUS 5143
G Ww ARGUS 5133
G Ww ARGUS 5127
G W ARGUS 5154
G6 W ARGUS 5126
G w ARGUS 5981
R = BLACK 4715F
R © BLACK 4714F
R F BLACK 4716F
R © BLACK
K HOLMEN 611°47
MURDOCK 45
MURDOCK 29
P © SCHOLANDFR
P SCHOLANDER S13
J THOMAS 2135
J THOMAS 2136
J THOMAS 2136
J THOMAS 2135
J THOMAS 2156
J THOMAS 2153
I t WIGGINS 12573
I L WIGGINS 12573
I tL WIGGINS 12573
I Lt WIGGINS 12573
I L WIGGINS 12413
I L WIGGINS 1-842
I L WIGGINS 12843
I L WIGGINS 19573
I Lt WIGGINS 12573
I L WIGGINS 1pau3s
— HULTEN 5005
Ww “ KLEIN P3AL
T BANK JURe 4033
T BANK JURe 4038
T BANK JURe 4225
L JORDAL 2700
M JOHNSON ORD
G HALEY
G HALLEY
F COvVILLE 10R4
F v COVILLE 2085
F v COVILLE 2085
R © GRIGGS
O MURIE
J © ANDERSON 4059
w H BREWER 329
WH BREWER 325
BRYANT
L J COLE
@ J iGole
P COLINVAUX 1
F v COVILLE 1837
F v COVILLE 1837
F v COVILLE 1837
—E HULTEN
E JOHNSTON
T KINCAID
J MACOUN
J _MACOUN
J = MACOUN
J MACOUN
J “ MACOUN 16645
J “ MACOUN
J = MACOUN 93815
W PALMER 556
Ww PALYER 424
W PALMER 353
C 4 TOWNSEND
R WHITE
A G WHITNEY
F v COVILLE 2087
S COVILLE 2087
F v COVILLE 20R7
VL HARMS 5554
VL HARMS 5326
VL HARMS 5357
HERB + NO
GWA
GWA
GWA
BWA
OWA
BWA
GWA
SWA
GWA
GWA
GWA
nWA
BWA
GWA
GWA
GWA
TSC
1SC
1SC
1Sc
"AO
US
GH
1SC 256627
US 2031640
CAN 248983
US 2312572
64
US 2312571
GH
64
ALA 6551
CAN 248984
TSC 224815
“ICH
GH
YS 2312909
US 2312910
NS 2396195
6H
G4
256626
256623
256625
256624
32733
19207
373511
373513
373512
US 1072233
GH
ysc
US
US
US
tsc
US
GH
A
US
US
“IY
US
US
CAN
CAN
CAN
CAN
NY
US
GH
US
US
YS
US
GH
us
US
A
US
ALA
ALA
ALA
256561
378317
378315
26251
256503
378343
373508
373509
1289510
376364
46451
46452
46450
46449
338561
328007
327983
327984
219257
423250
373515
373514
32673
32662
32664
237
Appendix B
TaxON SPC S
1873
1873
1873
1873
1873
1873
1873
1873
1873
1873
18673
1873
1873
1873
1873
1673
1873
1873
1873
1673
87
1873
1873
1873
1873
1873
1873
1873
1673
1673
1873
1873
1673
1873
1873
1873
1873
1873
1673
1473
1873
1873
1873
1873
1673
1673
1873
1873
1473
1873
1873
1675
1873
1873
1875
1473
14673
1873
1873
1873
1873
1873
1873
15673
1873
1673
1873
1873
1673
1673
1473
1673
1873
1873
1673
1873
1473
1873
18673
1673
1673
1873
1673
1473
14673
1873
1673
1675
238
83
83
83
83
83
83
83
83
83
83
83
83
63
83
83
63
83
83
83
83
oooooococooccoocoococococo0ondo
5 ee ee ee
BARCLAYI
87
87
87
87
87
87
87
87
87
87
87
87
87
87
87
87
87
87
87
87
87
87
87
87
87
87
87
87
87
87
87
87
67
67
67
87
87
87
67
87
67
87
87
87
87
67
87
87
67
87
87
87
87
87
87
87
87
87
87
87
67
87
87
67
87
87
87
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
V HYB PROV QUAD LAT
6024
6024
6024
6024
6024
6024
6024
6024
6024
6024
6342
6342
6342
6342
6330
6330
6330
6459
6545
6536
5251
5935
5935
5935
5935
5935
5935
5935
5935
5935
5935
5935
5935
5935
5935
5935
5935
5935
5935
5935
5935
5935
5935
5935
5941
5831
5831
5320
5320
5320
4926
6331
6205
6257
6140
6120
6130
6145
0058
0057
6004
6004
6004
0043
6043
6005
6005
0003
6002
5555
5626
5648
56468
5637
5637
5637
5859
3859
5859
5859
5858
5859
5858
5858
5857
5855
5855
LOCALITY
MATTHEW IS
MATTHEW
MATTHEW IS
MATTHEW IS
MATTHEW IS
MATTHEW IS
MATTHEW IS
MATTHEW IS
MATTHEW IS
ST MATTHEW IS
SAVOONGA
SAVOONGA
SAVOONGA
SAVOONGA
ST LAWRENCE IS
ST LAWRENCE IS
ST LAWRENCE IS
KING ISLAND
LTL DIOMEDE IS
CA PRINCE WALE
MEDICINE L
THREE SUARDSME
THREE GUARDSME
THREE GUARDSME
THREE GUARDSME
THREE GUARDSME
THREE GUARDOSME
THREE SUARDSME
THREE SUAROSME
THREE SUARDSME
INSPECTOR CREE
INSPECTOR CREE
INSPECTOR CREE
INSPECTOR CREE
INSPECTOR CREE
INSPECTOR CREE
INSPECTOR CREE
INSPECTOR CREE
INSPECTOR CREE
INSPECTOR CREE
INSPECTOR CREE
INSPECTOR CREE
INSPECTOR CREE
MT GLAVE
NADAHINI RIVER
SUMMIT PASS
SUMMIT PASS
PRINCE GEORGEe
> PRINCE GEORGE.
PRINCE GEORGE.
APEX MT
JUNE LAKE
BRINTNELL LAKE
ITSI RANGE
CANOL 2D MI 77
ROSE R
UPPER ROSE R
ROSE RIVER
KLUANE L
KLUANE L
HAINES RD M95
HAINES RD M 95
> HAINES RD M 95
WHITEHORSE
WHITEHORSE
RANCHERIA
RANCHERIA
WATSON LAKE
WATSON LAKE
HYDER
WRANGELLe MI
PETERS3URG-~
PETERSSURGe
PETERS3URG.
PETERS3URGe MI
PETERSSURGe MI
MUIR INLET? GL
MUIR INLET?
MUIR INLET?
MUIR INLET?
MUIR INLET?
MUIR INLET?
MUIR INLET»
MUIR INLET?
MUIR INLET?
MUIR INLET»
MUIR INLET?
DATE
Oo
oo
DBNOWDN NNN NN AN N NN N NN N NN WN NNN ANNAN ANANSI
co °
~w Oo
|
oooo0oocoocco°9e oo
DDDNWDNDNNNNNNNFEFNODDBDADMDNNAODDNNAINN
966
966
966
966
966
966
966
966
957
957
933
533
933
931
931
931
931
968
926
924
941
967
967
967
967
967
967
967
967
967
967
967
967
967
967
967
967
967
967
967
967
967
967
967
967
943
943
967
967
967
953
967
939
960
944
944
944
944
944
944
94R
948
94e8
944
944
966
966
966
94%
939
967
967
967
967
967
967
967
967
967
967
967
967
967
967
967
967
967
COLLECTOR NAME
rTrepwmrrrTlde909#coooeccccccce
DOOMDADAANDANDAGDONANONGNOAODCIOOOPSPCLTILTIC PrP PrPPCTIEcagaaLIAONADNDDAANDAVNADDAVDAVDANANVDAADVDDGA AM
£229 0-6 eee
a |
= >
GOAN NH 1 >
HARMS
HARMS
HARMS
HARMS
HARMS
HARMS
HARMS
HARMS
KLEIN
KLEIN
GEIST
GEIST
GEIST
GEIST
MASON
MASON
MASON
PEGAU
PORSILD
RYNNING
SCAMMAN
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
RAUP
RAUP
ARGUS
ARGUS
ARGUS
CALDER
CODY
RAUP
CALDER
PORSILD
PORSILD
PORSILD
PORSILD
ANDERSON
RAuUP
RAUP
RAUP
RAUP
ANDERSON
PORSILD
ARGUS
ARGUS
ARGUS
RAuUP
ANDERSON
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
AGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
COL NO
5575
5576
5649
5336
5357
5575
5649
5336
133
129
110
099
194
6101
6191
4168
1676B
1070
2521
6758
6759
6752
6753
6757
6769
6768
6766
6755
6714
6701
6702
6703
6704
6705
6700
6715
6499
6729
6738
6737
6732
6783
6808
1N663
10666
6780
6981
6976
10763
17300
a259
27719
10273
10075
i1nyag
10561
9404
129523
13116
3115
SiS
a6N5
1N662
6019
6016
5055
10966
Suns
69°43
69°35
6°36 !
6233
6830
6°29
6512 W
6507
6°18
65904
6455
6482
6450
644s
6431
6775
6RASC
HERB +
NO
32674
32675
32678
ALA
ALA
ALA
GWA
GWA
SWA
GWA
GWA
ALA
ALA
ALA
ALA
ALA
NY
US
GH
NY
GWA
US 1789736
ALA 2029
6668
6667
29417
2938?
29416
1789435
ALA 14755
GWA
GWA
GWA
GWA
GWA
GWA
SWA
SWA
GWA
BWA
GWA
GWA
5WA
5WA
GWA
SWA
GWA
5WA
BWA
5WA
CWA
GWA
BWA
GWA
ALA
ALA
GWA
GWA
GWA
ALA
NAO
ALA
GWA
CAN
TSC
TSC
CAN
TSC
ALA
ALA
ALA
CWA
TSC
TSC
SwA
GWA
GWA
19871
19873
11A10
58179
14777
4302
255982
255976
48299
2559s
19874
19867
19868
19868
256713
256711
19872
256015
Specimens Cited
TAxON SPC S V HYB PROV QUAD LAT LONG LOCALITY DATE
1673 87 14 10 5855 13603 MUIR INLETe GL 28 6
1873 87 14 10 5855 13603 MUIR INLET* GL 28 6
1873 87 14 10 5855 13603 MUIR INLET? GL 28 6
1873 87 14 11 5825 13432 MENDENHALL SL 07 06
1875 87 14 11 95818 13424 JUNEAU 08 08
1573 87 14 11 5825 13432 MENDENHALL GL 31 05
1873 87 14 11 5825 13432 MENDENHALL GL 07 06
1873 87 14 11 5825 15432 MENDENHALL GL 24 06
1873 87 14 1i 5823 13440 JUNEAU. 13216
1873 8&7 14 11 5823 13440 JUNEAU 13° 6
1373 87 14 11 5823 13440 JUNEAU. 136
1873 87 14 1i 5822 13436 MENDENHALL RIV 13 6
ls73 87 14 11 5826 13435 MENDENHALL GLA 10 7
1673 87 14 11 5831 13445 EAGLE RIVERe 110 7
1573 87 14 11 5826 13435 MENDENHALL GLA 10 7
14673 87 14 li 5826 13435 MENDENHALL GLA 10 7
1873 87 14 11 5820 13436 DOUGLAS. 07> 7,
1873 87 14 11 3818 15423 MT ROBERTS TR 09 7
1873 87 14 11 5818 134235 MT ROBERTS TR 09 7
14673 87 14 11 5820 13436 DOUGLAS ISLAND 07 7
1873 87 14 11 5820 13436 DOUGLAS ISLAND 07 7
1873 87 14 11 5820 13436 DOUGLAS ISLAND 07 7
1673 87 14 11 5825 13540 GUSTAVUS. 04 07
1873 87 14 11 5825 13540 GUSTAVUS. 04 07
1873 87 14 11 5825 13540 GUSTAVJS. 04 07
1673 387 14 11 5825 13540 GUSTAVJS. 04 07
1573 87 14 11 9825 13540 GUSTAVUS. 04 07
16735 87 14 11 5825 13540 GUSTAVUS. 04 07
1875 87 14 1i 5825 13540 GUSTAVUS. 04 07
1675 87 14 TY (5831 VWS445 EACLE RIVER. FT 107
1873 87 14 11 5839 13413 TAKU B NUNATAK 10 08
1875 87 14 11 5853 13420 JUNEAU 22 08
1873 87 14 11 5818 13424 JUNEAU 07 07
1573 87 14 11 5820 13433 MENDENHALL GL 12 06
1673 87 14 11 5826 13440 MONTANA CREEK 07 08
18735 87 14 11 58631 13445 MTH EAGLE R 08 08
1873 87 14 11 5831 13445 MTH EAGLE R 07 08
1475 87 14 11 3825 13432 MENOENHALL GL 07 08
1673 87 14 11 5825 13432 MENDENHALL GL 07 08
1673 87 14 11 5820 13433 MENDENHALL GL 12 06
1873 87 14 23 5352 16632 UNALASKA 22 07
18673 87 14 23 5352 16632 UNALASKA 18 06
1873 87 14 23 535% 16631 DUTCH HARBOUR 07 07
138735 8&7 14 23 5354 16631 DUTCH HARBOUR 30 06
1873 87 14 23- 5354 16631 DUTCH HARBOUR 29 06
18673 87 14 23 5352 16632 UNALASKA 27 07
1873 87 14 23 5402 16603 AKUTAN PASS 19 06
1875 87 14 23 5352 16632 UNALASKA 2027
187535 87 14 23 5352 16631 ILIULIJK LAKE 01 07
18735 87 14 23 53552 16631 ILIULIUK LAKE 01 07
1873 87 14 24 5408 16546 AKUTAN
1873 87 14 28 5520 16030 POPOF ISLAND 9017
1473 87 14 29 9515 16230 COLD BAY 28 08
1873 87 14 30 5618 15824 CHIGNIK 20 07
1873 87 14 32 5957 15410 ALITAK 05
1873 87 14 34 5713 15315 SHEEP ISLAND 06
1873 87 14 34 $717 15331 THREE SAINTS B 05 08
18735 87 14 34 5717 15333 THREE SAINTS B 27 06
1873 87 14 34 5750 15228 KODIAK 14-08
1673 87 14 34 S747 15247 KODIAK 28 07
1873 87 14 35 5705 15425 OLGA BAY
1873 87 14 41 5841 15700 KING SALMON 23 07
1873 87 14 42 5835 15550 DUMPLING MT 29 05
1873 87 14 42 5835 15550 DUMPLING MT 29 05
1875 87 14 42 5841 15515 GROSNENOR LAKE 22 07
1873 87 14 42 5817 15509 BROKEN MT 06 08
1873 87 14 42 5817 15509 BROKEN MT 06 08
1873 87 14 43 5804 15304 RASPBERRY IS 11 06
18735 87 14 45 5804 15304 RASPBERRY IS 04 07
1873 87 14 43 5804 15304 RASPBERRY IS 10 09
1873 87 14 43 5804 15304 RASPBERRY IS 27 06
1673 87 14 43 5802 15245 AFOGNAK ISLAND 15 06
1873 87 14 45 5907 13520 HAINESe MIe 6 13 7
1873 87 14 45 5907 13520 HAINESe MIe 6 13 7
1873 87 14 45 5928 13605 MOSQUITO LAKE 45 7
1873 87 14 45 5928 13605 MOSQUITO LAKE 45 7
1875 87 14 45 5925 13603 KLEHINI RIVER 12 7
1673 87 14 45 5928 13605 MOSQUITO LAKE 45 7
1875 87 14 45 5928 13605 MOSQUITO LAKE 45 7
1873 87 14 45 5928 13605 MOSQUITO LAKE 45 7
1873 87 14 45 5928 13602 MOSQUITO LAKEe 11 7
1873 87 14 45 5914 13529 HAINES 04 08
1875 87 14 46 5915 13830 TANIS LAKEe 22. "6
1873 87 14 46 5915 13830 TANIS LAKE 19 6
18753 87 14 46 5915 13830 TANIS LAKEe 2e6
1873 87 14 46 5915 13830 TANIS LAKE 19 6
1873 87 14 46 5915 13830 TANIS LAKE iS 6
1873 87 14 46 5915 13830 TANIS LAKE-s 22 46
1873 87 14 46 5915 13830 TANIS LAKE 196
13873 87 14 46 5915 13830 TANIS LAKE WE pies
1873 87 14 46 5915 13830 TANIS LAKE 2. 6
967
967
967
940
O17
925
940
917
967
967
967
967
967
967
967
967
967
967
967
967
967
967
967
967
967
967
967
967
967
967
966
958
966
965
967
967
967
967
967
965
93R
932
907
907
907
940
914
971
899
899
933
941
958
941
937
963
963
963
963
904
965
967
967
954
954
954
945
945
946
939
940
967
967
967
967
967
967
967
967
967
967
967
967
967
967
967
967
967
967
967
COLLECTOR NAME
=
ARGUS
ARGUS
ARGUS
ANDERSON
ANDERSON
ANDERSON
ANDERSON
ANDERSON
ARGUS
ARGUS
ARGUS
w ARGUS
w ARGUS
Ww ARGUS
w ARGUS
w ARGUS
WwW ARGUS
Ww ARGUS
ARGUS
Ww ARGUS
ve ARGUS
w ARGUS
w ARGUS
w ARGUS
Ww ARGUS
w ARGUS
w ARGUS
w A26US
W ARGUS
w ARGUS
BESCHEL
HEUSSER
ROBUCK
VIERECK
VIERECK
VIERECK
VIERECK
VIERECK
VIERECK
VIERECK
ANDERSON
EYERDAM
VAN DYKE
VAN DYKE
VAN DYKE
== S00. 0.0. 0 sa
Ls os > > > >
+
(= We )
MACOUN
WILLIAMS
JEPSON
JEPSON
W GEIST
J COLE
J HARBO
J COLE
LOOFF
GORDON
H NYBAKKEN
H NYBAKKEN
H NYBAKKEN
PIPER
LOOFF
L HARMS
ESTABROOK
ESTABROOK
SCHALLER
SCHALLER
SCHALLER
EYERDAM
EYERDAM
EYERDAM
EYERDAM
GABRIELSON
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
VIERECK
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
GG GG
DOOANADNANANVND ACT ANAND AANDOADAD OH EZTEEDONODQOOCMOWDWVDAMTUHNFTOCZZTSECHMPPZCFFOFTFTFTTFENDADDNDDADAANVDADANDGVADNDANDGMVAGNADAOADONCCCCCAOD
Z£2€2 2.2. 4:E:8 15 > 2322252125 5155 = n
GABRIELSON
COL NO HF28 + NO
6277
6366
6359
6226
413
2A16
6226
4ns
6103
6102
6104
6106
6664
6669
6663
6657
6600
6641
6641
6598
6599
6596
6590
6587
6585
6584
6576
6571
6570
6668
15493
CWA
GWA
SWA
ALA
1sc
Isc
msc
TSsc
SWA
GWA
GWA
GWA
GWA
GWA
5WA
GWA
GWA
SWA
GWA
SWA
5WA
GWA
SWA
SWA
SWA
GWA
GWA
SWA
GWA
GWA
GWA
GWA
16 FSLC
7623 GWA
R677 GWA
R6ER7T GWA
R625 SWA
R640 GWA
R639 GWA
7623 FSLC
4enS SC
8RB GH
29 «66H
7 Ge
196 SH
MA
500
85662
25598n
256931
85663
493
340
256019
2912
a5 US
87 US
883670
883671
29710
256006
24753
256005
17 ALA
170 “A
2746
1036
2697
1058
46n2 A
12th
4451
1°5
124
SeNe
SeNe
SeNe
3708
3775
52R2
2061 A
32564
34692
34691
2932
2980
2979
256158
256156
256155
6748
6749
6678
6678
6687
6686
6681
6680
6673
BSR6
6339
6200
6338
6244
6243
6310
6202
6201
62°73
239
Appendix B
TaxON SPC S V HYB PROV QUAD LAT LONG LOCALITY DATE
1873 87 14 46 5915 13830 TANIS LAKE ee oO
1475 87 14 46 5915 13830 TANIS LAKE el 6
1873 87 14 46 5915 13830 TANIS LAKEe 22 6
18735 87 14 Go 5915 13830 TANIS LAKE Lo 6
16735 67 14 46 5915 13830 TANIS LAKE 19°46
1875 67 14 46 5955 153945 YAKUTAT BAY. 1576
1873 87 14 46 30955 13945 YAKUTAT BAY. 15-6
1873 87 14 46 5952 13945 YAKUTAT BAY. to 246
1875 87 14 46 5915 13830 TANIS LAKE Loic
1873 87 14 46 5952 13945 YAKUTAT BAY. lbs ‘6
1673 87 14 46 5952 13945 YAKUTAT BAY. 1% 6
1473 87 14 4o 5952 13945 YAKUTAT BAY. 14% #6
18753 87 14 46 5915 13830 TANIS LAKE 19 06
1875 87 14 46 5932 13940 YAKUTAT. 25 6
1873 87 14 46 5932 13940 YAKUTAT. 25, 6
1875 87 14 46 5933 13930 SITUK RIVER. 23 6
1673 87 14 46 5952 13945 YAKUTAT BAY 14° 6
1873 87 14 46 5952 13945 YAKUTAT BAYe 15° 6
1873 87 14 46 5952 13945 YAKUTAT BAY. Te 6
1873 87 14 4e 5915 13830 TANIS LAKE L9-"6
1873 87 14 46 5935 13940 SITUK RIVER 10 06
1573 87 14 46 5935 13940 SITUK RIVER 09 06
1873 87 14 46 5935 13940 SITUK RIVER 12 06
1873 87 14 4o 5935 13940 SITUK RIVER 12 06
1673 87 14 46 9935 13940 SITUK RIVER 09 06
1873 87 14 46 5935 13940 SITUK RIVER 09 06
158735 87 14 46 5935 13940 SITUK RIVER 09 06
1875 87 14 46 5935 13940 SITUK RIVER 10 06
1873 87 14 46 5922 13913 DANGEROUS R 11 06
1873 87 14 46 5930 15940 YAKUTAT 09 06
1875 87 14 4o 5924 13900 HARLEQUIN L 10 06
1875 87 14 46 5924 13900 HARLEQJIN L 10 06
1675 87 14 46 5922 13913 DANGEROUS R 11 06
1675 87 14 46 5924 13900 HARLEQUIN L 10 06
1873 87 14 46 5924 13900 HARLEQUIN L 10 06
1873 87 14 46 5930 13940 YAKUTAT 09 06
1873 87 14 46 5922 13913 DANGEROUS R 11 06
1673 87 14 48 5926 14620 MIDDLETON IS 09 06
1873 87 14 50 5959 15145 STERL AWY M1359 13 06
18753 87 14 50 5940 15128 HOMER 13 06
1873 87 14 Su 994U 15128 HOMER 13 06
1873 87 14 57 ©6037 16515 NELSON ISLAND 02 07
1673 87 14 60 6015 15859 NUSHAGAK 15 06
1873 87 14 62 0023 15118 KASILOF 09 08
1673 87 14 62 6030 15100 SOLDOTNA 07 07
1673 87 14 62 6025 15020 SKILAK LAKE
1673 87 14 63 6029 14920 MOOSE PASS 01 06
18735 87 14 63 6006 14926 SEWARD 31 05
1873 87 14 63 6029 14950 COOPERS LANDG 21 06
18735 67 14 63 0029 14950 COOPERS LANDG 21 06
1873 87 14 63 6029 14920 MOOSE PASS 01 06
1873 87 14 63 6029 14920 MOOSE PASS 01 06
16735 87 14 63 ©6029 14920 MOOSE PASS 01 06
1673 87 14 63 6029 14920 MOOSE PASS 01 06
1873 87 14 63 6029 14920 MOOSE PASS 01 06
18675 87 14 65 6029 14920 MOOSE PASS 01 06
1673 87 14 63 6029 14920 MOOSE PASS 01 06
1673 87 14 63 6006 14926 SEWARD 31 05
1675 87 14 63 6008 14925 SEWARD 06 06
1673 87 14 63 ©6038 14835 TEBENKOF GL 27 08
1873 87 14 63 6056 14710 COLUMBIA BAY 05 09
18735 87 14 63 6006 14926 SEWARD 27 05
1873 87 14 63 6045 14914 SEWARD HWY “65 29 06
16735 87 14 63 ©6029 14920 MOOSE PASS 20 06
1673 87 14 63 ©6029 14920 MOOSE PASS 20 06
1873 87 14 63 06055 14940 HOPE 18 06
1873 87 14 63 6003 14755 LATOCHE ISALND 15 07
1875 87 14 63 6049 14857 PORTAGE GL RD 11 06
1873 87 14 63 6049 14857 PORTAGE GL RD 11 06
1673 87 Be 63 6045 14914 SEWARD HWY “65 11 06
1875 87 14 63 6045 14914 SEWARD HWY M65 11 06
1873 87 14 63 6032 14931 SEWARD HWY W38 12 06
1873 87 14 63 6046 14924 SEWARD HWY M55 01 07
1673 87 14 63 6050 14858 PORTAGE 07 07
1673 87 14 64 6033 14545 CORDOVA 12 06
1673 87 14 64 6033 14545 CORDOVA 10 06
1873 87 14 68 6107 14616 VALDEZ 05 07
1873 87 14 68 6107 14616 VALDEZ 05 07
1873 87 14 68 6107 14616 VALDEZ 03 07
1873 87 14 68 6100 14500 COPPER R FLATS
1673 87 14 68 6100 14500 COPPER R FLATS
1875 87 14 68 6100 14500 COPPER R FLATS
1875 87 14 68 6100 14500 COPPER R FLATS
18675 87 14 68 6100 14500 COPPER R FLATS
1875 87 14 68 6100 14500 COPPER R FLATS
1875 87 14 68 6112 14547 WORTHINGTON GL 01 O08
1673 87 14 68 6112 14547 WORTHINGTON GL 01 08
1873 87 14 68 6108 14547 THOMPSON PASS 01 08
1875 87 14 68 6112 14615 VALDEZ 02 08
1873 67 14 68 6108 14610 VALDEZ GLACIER 09 08
1873 87 14 68 6108 14610 VALDEZ GLACIER 09 08
240
967
967
967
967
967
967
967
967
967
967
967
967
167
967
967
967
967
967
967
967
966
966
966
966
966
966
966
966
965
965
965
965
965
965
965
965
965
956
967
967
967
946
935
949
933
939
941
941
941
941
941
940
941
941
941
941
94)
941
951
935
935
945
967
939
939
962
957
967
967
967
967
967
966
965
928
940
935
935
931
960
960
960
960
960
960
967
967
967
967
957
957
COLLECTOR NAME
AaArrrrPprrrrr=e=eccmxununrrrrrirrrcZtzeccccccceccecccccrriccrrrerrrrrrrrr<ee cece ec KC KC MNADADDANIDAVAAVADANAVDANDANADANADNGDAD
Ww
a aPrarrwvrbrrserrerrrny rere
<= >
PrVIVUVVVVVVVVYG
pee
=
RPPr?rp»>
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
HARMS
HARMS
HARMS
HARMS
HARMS
HARMS
HARMS
HARMS
VIERECK
VIERECK
VIERECK
VIERECK
VIERECK
VIERECK
VIERECK
VIERECK
VIERECK
THOMAS
VIERECK
VIERECK
VIERECK
STEWART
MERTIE JR.-
LUTZ
PALMER
SHARPLES
ANDERSON
ANDERSON
ANDERSON
ANDERSON
ANDERSON
ANDERSON
ANDERSON
ANDERSON
ANDERSON
ANDERSON
ANDERSON
ANDERSON
CALDER
COOPER
COOPER
EYERDAM
HARMS
NELSON
NELSON
ROWINSKI
SHACKLETT
VIERECK
VIERECK
VIERECK
VIERECK
VIERECK
WELSH
WELSH
MEXIA
SCAMMAN
ANDERSON
ANDERSON
SETCHELL
SHEETS
SHEETS
SHEETS
SHEETS
SHEETS
SHEETS
VIERECK
VIERECK
VIERECK
VIERECK
VIERECK
VIERECK
COL NO HERA + NO
6769
62°91
6337
6237
6242
6178
6182
6127
6199
6189
6118
6119
6198
6240
63242
6349
6159
6168
6163
6205
5i71A
5140
5217
52235
5141
5146
5159
52ny4
7616
75°6
7606
76°7
7616
7607
7696
75°6
7619
8564
R264
R273
R272
231
158
116
8
6468
6466
6929
6931
6772
6478
6498
6467
6468
6469
6470
6466
5100
2Al
373
3673
6063
502
3471
4396A
8226
8228B
R235
B254
8253
5670
4548
2005
1598
1868
1866
257
SeNe
SeNe
SeNe
GS eNe
SeNe
8477
AYUAL
8488
8751
2226
8394
SWA
SWA
GwA
GWA
SWA
GWA
GWA
GWA
SWA
GWA
SWA
GWA
SWA
GWA
GWA
GWA
GWA
GWA
SWA
SWA
ALA 32575
ALA 32580
ALA 32570
ALA 32569
ALA 32579
ALA 32578
ALA 32577
ALA 32573
GWA
SWA
GWA
GWA
FSLC 331
ESEC 339
FSEC 328
FSLC 338
FSLC 344
US 2313000
GWA
GWA
GWA
NY
US 1632402
NA 326898
NA 326908
TSC 256014
ALA 499
ALA 501
TSC 256030
TSC 256153
TSC 256150
¥SC 256152
TSC 256025
ISC 256027
ISC 256028
TSC 256029
TSC 256151
ISC 256023
GWA
F 824012
F 824040
ISC 256157
ALA 34635
ALA
Isc 256021
ALA 25672
US 2387513
BWA
SWA
GWA
SWA
BWA
GWA
ISC 246902
ALA 10199
ALA 14752
ISC 256018
ISC 256026
NA 326924
ALA 25131
ALA 25134
ALA 25154
ALA 25124
ALA 25126
ALA 25132
GWA
GWA
GWA
GWA
GWA
ALA
Specimens Cited
TaxON SPC S V HYB PROV QUAD LAT LONG LOCALITY DATE
18735 87 14 69 615U 14706 GLENN HWY M128 13 06
1873 87 14 69 6140 14900 MATANUSKA VY 06 07
1873 87 14 69 6150 14706 GLENN HwY M128 13 06
1873 87 14 69 0230 14900 TALKEETNA MTS 12 07
14675 87 14 69 6230 14900 TALKEETNA MTS 12 07
1675 87 14 69 6230 14900 TALKEETNA WTS 28 06
1873 87 14 69 ©0230 14900 TALKEETNA MTS 12 07
1873 87 14 69 0230 14900 TALKEETNA WTS 12 07
18735 87 14 69 0230 14900 TALKEETNA MTS 28 06
1873 87 14 69 6105 14811 HARRIMAN FIORD 21 08
1875 87 14 69 6105 14811 HARRIMAN FIORD 18 08
1873 87 14 69 6113 14953 ANCHORAGE 29 06
1875 87 14 69 ©0124 14929 BIRCHWOOD FLAT 01 06
18753 87 14 69 ©6104 14950 ANCHORAGE 10 07
1873 87 14 69 o140 14900 MATANUSKA VY
1873 87 14 69 6140 14900 MATANUSKA VY
1873 87 14 69 6140 14900 MATANUSKA VY
1673 67 14 69 0140 14900 MATANUSKA VY
14673 87 14 69 6140 14900 MATANUSKA VY
1673 87 14 69 6149 14825 CHICKALOON CR 11 06
1875 8&7 14 69 6100 14702 COLUMBIA GL 02 08
1675 87 14 69 6146 14915 PALMER 15 07
18735 87 14 69 6113 14953 ANCHORAGE 15 06
1873 67 14 69 6147 14907 LTL SUSITNA CN 13 06
1873 87 14 69 0147 14907 LTL SUSITNA CN 13 06
1673 87 14 69 6113 14953 ANCHORAGE 15 06
16735 87 14 71 6152 15433 HEAD OF BIG Re 4& 7
1873 87 14 71 6152 15435 HEAD OF BIG Re 2 7
1873 87 14 71 6152 15433 HEAD OF BIG Re 2 7
1873 87 14 79 6217 15612 KUSKOKWIM Re 1 Oh 37,
1873 87 14 80 6228 15350 FAREWELL 12 8
1673 87 14 81 6226 15122 CHELATNA LAKE 14% 06
1873 87 14 81 6257 15222 KUSKOKWIM R 29 07
1875 87 14 81 6225 15125 CHELATNA LAKE 21 06
1873 87 14 81 6257 15222 KUSKOKWIM R 10 08
18735 87 14 81 6257 15222 KUSKOKWIM R 05 08
1873 87 14 81 6257 15222 KUSKOKWIM R 21 08
1873 87 14 83 ©6215 14525 GULKANA 19 06
1873 87 14 83 ©6215 14525 GULKANA 19 06
1875 87 14 83 6215 14525 GULKANA 19 06
1475 87 14 83 6215 14525 GULKANA 19 06
1875 87 14 83 6215 14525 GULKANA 19 06
1873 87 14 83 6215 14525 GULKANA 19 06
16735 87 14 83 6215 14525 GULKANA 19 06
18735 87 14 83 ©6215 14525 GULKANA 19 06
1873 87 14 85 6307 14315 BIG TOK R BR 24 06
1875 87 14 85 6307 14315 BIG TOK R BR 24 06
186735 87 14 85 6307 14315 BIG TOK R 8R 24 06
1675 87 14 85 6307 14315 BIG TOK R BR 24 06
1873 87 14 85 63507 14315 BIG TOK R BR 24 06
1873 87 14 85 6312 14330 TANACROSS 19 07
1873 87 14 8o 6315 14540 MCCALLUM CR 19 06
1875 87 14 86 6347 14547 DONNELLY DOME 29 06
1873 87 14 86 6315 14540 MCCALLUM CREEK 19 06
1873 87 14 86 6315 14540 MCCALLUM CR 19 06
1873 87 14 86 6347 14547 DONNELLY DOME 29 06
16753 87 14 86 6315 14540 MCCALLUM CREEK 19 06
1673 87 14 86 6347 14547 DONNELLY DOME 08 06
1873 87 14 86 ©6347 14547 DONNELLY DOME O01 07
1873 87 14 86 ©6320 14535 RAINBOW MT 30 06
1875 87 14 86 6342 14553 RICH HWY M237 25 07
1673 87 14 86 6305 14536 DENALI HWY M5 26 07
1873 87 14 87 6350 14925 SAVAGE RIVER 18 06
1873 87 14 87 ©308 14927 SUMMIT 27 07
1873 87 14 88 6525 15050 WONDER LAKE 26 07
1875 87 14 88 ©6326 15033 MCK PK RD M67 20 06
1675 87 14 88 6326 15031 MCK PK RD M74 29 07
18735 87 14 88 6324 15025 MULDROW GL 13 07
1873 87 14 88 0325 15050 WONDER LAKE 08 08
1875 87 14 88 6325 15020 THOROFARE R 04 07
1875 87 14 88 6325 15050 WONDER LAKE 29 07
16753 87 14 88 6325 15020 THOROFARE R 04 07
1875 87 14 8&8 6325 15050 WONDER LAKE 08 08
1873 87 14 88 6324 15025 MULDROW GL 13 07
1873 87 14 88 6324 15020 THOROFARE R 14 06
1873 87 14 88 6324 15025 THOROFARE R 10 07
1873 87 14 88 6325 15050 WONDER LAKE 04 08
1873 87 14 86 6325 15050 WONDER LAKE 01 07
18753 87 14 88 6325 15050 WONDER LAKE 30 06
1675 87 14 88 6325 15035 MULDROW GL 17 06
1873 87 14 88 6325 15035 MULDROW GL 22 07
1875 67 14 88 6325 15035 MULDROW GL 22 06
1875 87 14 88 6325 15035 MULDROW GL 28 07
1873 87 14 88 6325 15035 MULDROW GL 22 06
1875 87 14 88 6325 15050 WONDER LAKE 26 06
1675 87 14 88 6325 15050 WONDER LAKE 07 07
1873 87 14 8&8 6325 15050 WONDER LAKE 28 06
1873 87 14 86 6325 15050 WONDER LAKE 29 07
1873 87 14 88 ©6325 15035 MULOROW GL 22 06
1873 87 14 88 6325 15050 WONDER LAKE 04 08
1873 87 14 88 . 6325 15050 WONDER LAKE 01 07
944
931
944
931
931
941
931
931
941
935
935
948
948
94A
940
940
940
940
940
967
957
968
965
965
965
965
950
950
950
949
949
961
961
956
961
961
961
957
957
957
957
957
957
957
957
944
944
944
944
944
957
957
957
957
957
957
957
965
964
966
967
967
928
967
956
960
963
956
958
956
956
956
958
956
958
958
958
958
958
958
958
958
958
958
958
958
958
956
958
958
958
COLLECTOR NAME
Preeerererererrerer reer eee er rrr ea ePoar x<rTr noe Ke OODDADODACFTCCCCOCHAHDODDDODOCFFFFKTMEFSFFENNNWNVWVTCTCOCTCOCeOMMMEZcececcececc
vuggvug07mn0
aa
| oie ey
cCwa
>
CEFR SV Sos LeEES LE FE et ke
rr-ecrzt ae ecCecztePVweevvucztz Ee
>>
=>
RBDerwr V?rRrr VPrPYVPKRTR FFP PYF PYPrPF PrP?
ANDERSON
ANDERSON
ANDERSON
ANDERSON
ANDERSON
ANDERSON
ANDERSON
ANDERSON
ANDERSON
COOPER
COOPER
LEPAGE
LEPAGE
LEPAGE
PALMER
PALMER
PALMER
PALMER
PALME
VIERECK
VIERECK
WELSH
WELSH
WELSH
WELSH
WELSH
DRURY
DRURY
DRURY
DRURY
DRURY
LITTLE»
VIERECK
VIERECK
VIERECK
VIERECK
VIERECK
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ANDERSON
ANDERSON
ANDERSON
ANDERSON
ANDERSON
SPETZMAN
ARGUS
ARGUS
ARGUS
AGUS
ARGUS
ARGUS
HARMS
HARMS
PARKER
VIERECK
VIERECK
MEXIA
VIERECK
ARGUS
MURIE
RICHEY
VIERECK
VIERECK
VIERECK
VIERECK
VIERECK
VIERECK
VIERECK
VIERECK
VIERECK
VIERECK
VIERECK
VIERECK
VIERECK
VIERECK
VIERECK
VIERECK
VIERECK
VIERECK
VIERECK
VIERECK
VIERECK
VIERECK
VIERECK
VIERECK
cOL NO
RYLT
BAS
RYUT
10%9
1027
70n2
1037
1039
7004
1R7
148
23371
23041
23,119
216
31
19
193
343
R217
2334
Q237
4146
4ORT
4103
4146
3979
3953
3986
2119 ¢
2078
19424
5184
1024
5304
5260
53A1
1046
1039
1045
1044
1039
1045
1046
1044
R714
R715
R714
B715
R713
722
1033
1057
1031
1033
1057
1031
3543
2BNG
RM?0
R338
R363
2017
RuND
657
8
1309
3295
10°4
1583
1094
3295
1309
3094
3003
3267
3128
3120
3010
3244
3062
3266
3060
30R9
3165
3194
1583
3060
3267
3128
HF 28
+ NO
256016
256009
27307
27315
256022
256020
256017
256154
R24014
B24039
255973
255975
255974
5980
5161
5167
5165
5162
30023
247227
247374
246119
8418
8415
8401
8418
8417
256008
256010
256007
6838
6709
6728
6739
32639
32646
32688
10196
26102
11622
8382
11623
11613
241
Appendix B
TaxON SPC S V H¥cG PROV BUA)
1873
18735
1873
15873
1s73
89
1873
1873
1873
ls673
1573
1873
1873
1873
lo73
1873
ls735
15873
1873
13873
1873
1675
1673
1673
1673
1873
1673
lo75
1673
16735
1673
1873
1473
1873
1673
1673
1873
18/3
1873
1873
1673
18735
1873
1s73
1873
1873
1873
18673
1673
1673
1673
1873
1673
1673
1673
15673
1673
1673
1873
1473
1873
90
1673
1673
1673
1873
lo73
1873
1873
1673
1673
1873
1673
1473
1873
1573
1873
1873
1873
1673
1673
1473
1673
1573
1673
242
14
14
14
14
14
87
87
67
87
87
CHAMISSONIS
89 7
89 FA
89 z
89 it
89 7
69 Le
69 14
89 14
89 a4
89 14
89 14
89 14
89 i4
89 14
89 44
89 14
89 14
89 14
89 14
89 14
69 14
89 14
89 i4
&9 14
69 14
89 14
89 14
69 14
69 14
89 14
89 14
89 14
89 14
89 14
89 14
89 14
89 14
89 14
89 14
89 14
89 14
89 14
89 14
89 14
89 14
89 14
89 14
89 14
89 14
89 14
89 14
89 14
89 14
89 14
89 14
BARRATTIANA
90
90
90
90
90
9U
90
90
90
90
90
90
90
90
90
90
90
90
90
90
90
90
90
NSN NSN NN NNN NNN NNN NDR
83
88
88
101
101
LAT
0325
o325
0325
6425
o415
68135
6800
0757
6800
6800
6835
5255
5255
6321
6318
6320
6320
0320
6320
o32l
6430
0430
0430
o43U
6437
0437
6430
o430
6420
o420
6456
0456
6524
6524
0524
0524
6524
6524
6524
0524
6525
6516
6512
6512
6725
6725
6810
6810
0810
6810
6810
6810
6810
0805
6847
oB22
6805
6805
6917
6935
5317
5241
5720
5648
6833
6511
6430
6310
6305
6340
6241
6205
0205
6205
6205
6205
6205
o12u
6252
6205
6102
6146
6149
LONG
15050
15050
15035
14650
14606
13554
13540
15627
13600
13600
13804
LOCALITY
wONDER LAKE
WONDER LAKE
MULDROW GL
HARDING L
SHAW CR FLATS
CANOE Le
RICHARISON MTS
FISH CRe
RICHARISON MTS
RICHARDSON MTS
SAM Le 22MI NE
17255EATTU ISLAND
17255EATTU TSLAND
16924
10842
17136
17136
17136
17136
17134
16530
10530
16590
16530
16542
16542
16530
16530
16500
16500
15442
15442
14558
14558
14558
14558
14558
14558
14558
14558
14525
16622
16645
16645
15007
15007
16540
16540
16540
16540
16540
16540
16540
16532
15950
15438
15030
15050
14600
14445
11755
11802
12355
12240
12829
12328
12810
13008
12850
12850
12750
12735
12735
12735
12735
12735
12735
12420
13015
128<c5
13832
13837
13835
MT TOM-NAME
NORTHEAST CAPE
BOXER 3AY
BOXER 3AY
BOXER 3AY
BOXER 3AY
WANMAYEE CR
NOME
NOME
NOME
NOME
TELLER RD
TELLER RD
ANVIL MT
ANVIL iT
CAPE NOME
CAPE NOME
KOKRINES
KOKRINES
TWELVE MI
TWELVEMI
TWELVEMI
TWELVEMI
TWELVEMI
TWELVEMI
TWELVEMI SUMIT
TWELVEMI SUMIT
EAGLE SUMMIT
TELLER
PORT CLARENCE
PORT CLARENCE
WISEMAN
WISEMAN
OGOTORUK
OGOTORUK
OGOTORJK
OGOTORJK
OGOTORUK
OGOTORUK CR
OGOTORUK CR
KUKPUK RIVER
NUKA RIVER
KURUPA LAKE
ULO LAKE
ULO LAKE
CANNING RIVER
SADLEROCHIT R
14M
14M
SUMIT
SUMIT
SuUMIT
SUMIT
SUMIT
SUMIT
CR
CR
CR
CR
CR
BRULE
MT EDITH CAVE
FAIRY LAKE
CYPRESS CR
ANDERSON RIVER
FT«FRANKLIN
BOLSTEAD CR.
MACMILLAN PASS
OGRADY Le
SEKWI 2.
GRIZZLY
COLONEL
COLONEL
COLONEL
COLONEL
COLONEL MT
COLONEL MT
Se» NAHANNI Reo
ITSI RANGE* NW
LTL«HYLAND Re
SHEEP MTe
PTARMIGAN HRT
PTARMIGAN HRT
BEAR L
MT
MT
MT
MT
958
95e
954
956
966
963
934
962
934
933
970
949
949
949
B99
933
933
933
933
96n
93A
Q4R
923
954
96RA
963
926
929
900
926
925
967
932
932
957
957
957
957
964
964
938
88a
889
949
949
950
959
961
959
959
960
960
963
95n
952
963
963
947
948
919
968
960
935
965
92R
944
944
967
944
967
939
939
939
939
939
930
970
944
963
967
948
94A
COLLECTOR NAVE
CQO i. i.
ow CMOOOCOOTNEZEOOYNPY FC P=
TCACNYFITIITIPPrPADPPrrnNnerrnnwnnszezacr z=enoyp
ZICVPSPOTITITILTIT SE PtyH EC Pref Oc
rE Ww1> WC
SS ON Sa ae oe
i SS See Gr toe
VIERECK
VIER ECK
VIERECK
ARGUS
FOOTE
copy
PORSILD
CALOER
PORSILD
PORSILD
WELSH
LLANO
LLANO
BENNINGHOF
COVILLE
GEIST
GEIST
GEIST
GEIST
SAUER
ANDERSON
NUTILLY
ENAND=R
HELLER
PEGAU
PEGAU
PORSILD
THORNTON
BLAISDELL
SETGHEEE
MILLER
PALMER
HARMS
SETCHELL
SETCHELL
SHETLER
SHETLER
SHETLER
SHETLER
VIERECK
VIERECK
ANDERSON
COVILLE
COVILLE
JORDAL
JORDAL
JOHNSON
JOHNSON
JOHNSON
JOHNSON
JOHNSON
MELCHIOR
MELCHIOR
MELCHIOR
SPETZMAN
HODGSON
CHAUVIN
FLOCK
MCGREGOR
SPETZMAN
MACOUN
ARGUS
CALDER
HENRY
SCOTTER
PORSILD
WYNNE =E9WARDS
PORSIL)
CODY
PORSILD
Copy
RAUP
RAUP
RAUP
RAUP
RAUP
RAuUP
SCOTTER
PORSILD
YOUNGMAN
NEILSON
RAUP
RAUP
cOL NO
3155
3120
3?A6
595
ROA
6926
32412
8273
11223
16°94
11941
17993
a371
9422
9370
9374
9422
9370
12597
11386
408
1347
13781
13781
H=2%RB + NO
FSLC
FSEC
FSLC
SWA
FSLC
AO
SWA
YAO
CAN 4Y7RO4
CAN 4T7ROD
32Yy 91070
JS 2095301
‘NA %29072
VICH
IS %73479
ALA 29409
ALA 29510
ALA 29508
ALA 29509
CAN
TSC 256383
TSC 256384
A
ALA 27074
CAN
1S 1438560
IS 423726
NA 328916
ALA
US 1325631
ALA 34A32
54
3H
PLA 3R0u
ALA 3731
ALA 3702
GWA
=SLC 303
[SLC 301
TSC 256382
NS 73477
A
MITGH
“TCH
ALA 9166
ALA 9180
SWA
SwA
eWA
ALA 17848
ALA 17759
SwWA
NA %28908
GH
ALA ?3A76
ALA 21744
MICH
US 2032247
TSC 256165
SWA
ALA 17433
ISC 256167
5BWA
CAN 48278
CAN 4YA3B1
CAN 48380
DAO
CAN 48382
OAO
ALA 14761
ALA 14768
ALA 14762
ALA 16594
SWA
5WA
SAWA
TAN 48430
CAN 279643
CON 313087
ALA 19858
CAN 275688
Specimens Cited
TAXON SPC S V HY¥b PROV wUAD
1873
1873
1673
1673
16735
1673
1873
1873
1573
1873
1873
1673
1673
1073
1673
1873
1675
1673
1473
18673
1873
1673
1873
16735
1873
1673
1873
1673
1675
1873
1873
1873
1873
18673
1873
1875
1673
1673
1673
1473
1873
1873
1873
1873
1873
1473
1873
1873
92
1673
1873
1873
1673
1473
1873
1673
1673
1873
1673
1873
1873
1873
1873
1873
1873
1873
1673
1873
13873
1673
1873
13873
1873
1873
1873
1873
1873
1673
1873
1673
1873
1673
1873
1673
1873
1873
1873
1673
90
90
90
90
90
90
90
90
90
90
90
90
90
90
90
90
90
90
90
90
90
90
90
90
90
9u
90
90
90
90
90
390
90
90
90
90
90
90
90
90
90
90
90
90
90
90
90
90
12
12
2
12
12
12
12
12
12
12
12
12
12
12
12
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
PHLEBOPHYLLA
LAT
6130
b120
6130
6140
6145
0145
6130
6055
6055
6049
0057
6057
6022
603u
6034
6100
6112
6153
6155
6349
6306
6350
6239
6239
6239
6343
6325
6325
0330
6328
6326
6325
6325
6325
6325
6325
6330
6325
6330
6725
6725
0725
6836
6923
6926
6920
6928
6925
6920
6930
6813
6836
6839
6836
6745
6911
6933
6826
6740
6404
6002
5839
5841
5835
5941
6146
6146
0146
6146
6137
6137
6137
6137
6137
6137
6137
6137
6137
6259
6259
6257
6259
6259
6240
6340
0338
6351
LONG
13302
13300
13302
13302
13258
13258
13302
13843
13845
13844
13825
13825
13551
13300
13362
14139
14539
15421
15425
14109
14639
14925
14932
14932
14932
14915
15020
15020
15050
15011
15012
15020
15015
15020
15020
15029
15050
150350
15050
15007
15007
15007
14345
14339
14347
14500
14617
14910
13300
13347
13547
13403
13406
13403
13601
13907
13854
13848
14000
13925
12835
16054
15639
15551
15551
15808
15808
15808
15808
15930
15930
15930
15930
15930
15930
15930
15930
15930
15604
15604
15630
15622
15622
15230
14213
14450
14858
LOCALITY
CANOL RD MI 98
ROSE Re
UPPER ROSE Re
CANOL RD MI105
ROSE LAPIE 8
ROSE LAPIE &
UPPER ROSE R
SLIMS RIVER
SLIMS RIVER
OBSERVATION MT
KLUANE LAKE
KLUANE LAKE
LITTLE ATLIN L
CANOL 2D MI 10
CANOL RD M10
CHITINA Re
GLENN HWY M128
HEAD OF BIG Re
HEAD OF BIG Re
DIVIDE MT.
MACCLAREN R
SAVAGE R
TEKLANIKA R
TEKLANIKA R
TEKLANIKA 8
MT MCKINLEY PK
MT EIELSON
MT EIELSON
TOKLAT RIVER
STONY CREEK
MCK PK RD M63
MT EIELSON
MCK PK RD M64
MT EIELSON
MT EIELSON
THOROFARE R
TOKLAT RIVER
THOROFARE R
TOKLAT RIVER
WISEMAN
WISEMAN
WISEMAN
SHEENJEK R
JAGO RIVER
JAGO LAKE
LAKE PETERS
SHUBLICK:S SPR
SADLEROCHIT Re
KITTIGAZUIT
RICHAROS IS
CANOE LAKE
REINDEER STAe
REINDEER STAc
REINDEER STAe
HORNE Le
BUCKLAND HILLS
HERSCHL IS
SAM Le 6MI NW
OLD CROW R
DAWSON
WATSON L
HAGEMEISTER IS
KING SALMON
DUMPLING MT
WONDER LAKE
CANOE MT-
CANOE MT-e
CANOE MT-e
CANOE MTe
ANIAK
ANIAK
ANIAK
ANIAK REGION
ANIAK
ANIAK
ANIAK
ANIAK
ANIAK
TAKOTNA
TAKOTNA
GANES ROAD
YANKEE CREEK
YANKEE CREEK
KUSKOKWIM R
MT FAIRPLAY
HORN RIDGE
HEALY
DATE
12
3
11
10
10
10
il
21
21
06
12
03
03
30
25
oo (=) ooooo0°co
AMDOANNNDWDAO~TIN
oo oooo
NNDOODANOARDADNNNNNDAND
o
oooo°oo
NOOONN
94"
944
944
944
944
944
944
966
966
966
944
944
943
944
944
925
944
950
950
963
967
928
939
939
939
937
956
956
956
967
962
947
959
956
956
956
956
964
965
939
939
940
956
957
957
948
947
948
927
966
964
964
965
965
962
970
959
970
926
914
953
952
966
951
949
949
949
949
949
949
949
949
949
949
949
949
949
941
941
949
940
940
961
954
957
939
COLLECTOR NAME
TrrececomecuwnrrrrrremMmacooomrrrxrr=z=ztcirrizrivovc0c0 rrrrvrvrvp
CrUMOTrMMECCHEEESZEZEFEZAEFETFTFEEMOFMIESENAUNCAAQCCA YS
AANMNANMQANNAN
=
vVureoervrvevx7e>
>rreieniwo
[fiat 2 cet ek ey |
ee DW RS Be See Me eket i oy (cds oe ike seks, jc oo
>
PORSILD
PORSILD
PORSILD
PORSILD
PORSILD
PORSILD
PORSILD
MURRAY
MURRAY
MURRAY
RAUP
RAuUP
RAU?
PORSILD
PORSILD
LAING
ANDERSON
DRURY
DRURY
SPETZMAN
VIERECK
MEXTA
NELSON
NELSON
NELSON
SCAMMAN
ARGUS
ARGUS
ARGUS
FOOTE
RICHEY
SCAMMAN
SCHENE
VIERECK
VIERECK
VIERECK
VIERECK
VIERECK
WELSH
ANDERSON
ANDERSON
SCAMMAN
SCHALLER
CANTLON
CANTLON
SPETZMAN
SPETZMAN
SPETZMAN
PORSILD
SCOTTER
LARSEN
LARSEN
SCOTTER
SCOTTER
CALDER
WELSH
wood
WELSH
MURTE
MILVAIN
RAuP
MULLAR
SCHOFIELD
ESTEBROOK
MULLAR
DRURY
DRURY
DRURY
DRURY
DRURY
DRURY
DRURY
DRURY
DRURY
DRURY
DRURY
DRURY
DRURY
ANDERSON
ANDERSON
DRURY
SCAMMAN
SCAMMAN
VIERECK
SMITH
SPETZMAN
ANDERSON
11°99
10272
19762
9293
9293
9294
10362
46
ung
561
12585
12525
11396
11937
11037
217
Q2URZ
4?51
4?15
4O7%
R390
2019
Z5P4
4256
3594
5°96
646
644
690
Lvsheay
4990
L361A
1214
1N96A
1793
7417
GRO
5878
5878
2233
164
57911
571558
674
443
236
132
5773
COL "O HER3 + NO
CON 48427
CAN 48433
CAN 48429
CAN 48426
TSC 256160
5H
6H
SWA
GWA
CWA
ALA 19911
GWA
ALA 19912
CAN 4B431
Sd
CAN 48439
TSC 256162
CAN
CAN
CAN 299614
GWA
ALA 10708
ALA 503
ALA 504
TSC 256114
GWA
SWA
GWA
SWA
SWA
ALA 26099
GWA
ALA 25318
ALA 11625
ALA 3509
ALA 11621
ALA 11596
FSLC 293
TSC 247203
ALA 27501
TSC 256163
464
WTS
6H
6H
CAN 212062
TSC 256161
CAN 212063
GWA
5WA
CWA
SWA
GWA
GWA
AO
OTE 1192
CAN 258920
B32yY 91021
ALA 540
A
CAN
US 2176420
6H
ALA 32683
"IS 2176392
CAN
AN
CAN
CAN
CAN
CAN
CAN
CAN
CAN
CAN
CAN
CAN
CAN
ALA 541
TSC 256255
CAN
ALA 8373
GWA
FSLC
ALA 10730
US 2349537
TSC 256256
243
Appendix B
TaxON SPC S V HYB PROV QUAD LAT LONG LOCALITY DATE
1873 992 14 89 6315 15442 NIXON MINES 4 9
1673 0-92 14 = 89 6315 15442 NIXON MINES 4 9
lb/5 = 92 14% 91 0352 16047 UNALAKLEET 06
1873-92 14 93 ©6322 17117 POWOOTLIAK
1673-92 144 93 6322 17117 POWOOILIAK 17 08
1673 92 14 93 ©336 17026 MT ATUK 7) 07
1673-92 14 93 ©6330 17100 ST LAWRENCE IS
1473 92 14 93 ©0342 17029 ST LAWRENCE IS
1673 92 14 93 6320 17136 BOXER 3AY 27 06
ls73 92 14 95 6320 17136 BOXER 3AY 27 06
1873 92 14 93 6320 17136 BOXER BAY 12 08
1673-92 14 93 6334 17053 KANGEE
1673-92 14 95 6319 17127 SW CAPE 08
1873-92 14 95 632U 17136 BOXER 3AY 27 06
inva ‘U2 14 94 6430 16525 NOME 11 06
lu73 92 14 94 6435 16540 NOME 16 06
lays °" 92 14 94 0435 16540 NOME 21 08
1673 92 14 94 6435 16540 NOME 16 06
1373s 92 14 94 6435 16520 DEXTER CREEK 07 08
1873092 14 94 6435 16540 NOME 29 07
1673 92 14 94 6435 16540 NOME 21 08
1373-92 14 94 6448 16515 NOME RIVER 14 07
ls73 92 14 95 6435 16425 DICKSON 10 08
ls73 92 14 95 6433 16302 GOLOVIN 17 06
tu73° 92 14 99 6500 15038 MANLEY HOT SPR 26 06
1873-92 14 100 6453 14803 ESTER JOME 20 07
1673 92 14 102 6438 14120 EAGLE 09 08
1873 - 92 14 104 6530 14525 EAGLE SUMMIT 18 07
1673 92 14 104 6530 14600 TWELVE MI SUMM 15 06
ls73 92 14 104 6530 14600 TWELVE WI SUMM 14 07
1873 = 92 14 104 653u 14525 EAGLE SUMMIT 20 07
1673-92 14 104 6530 14525 EAGLE SUMMIT
1873092 14 104 6530 14525 EAGLE SUMMIT 20 07
1873-92 14 104 6531 14531 PORCUPINE DOME 12 07
1673-92 14 104 6527 14526 EAGLE SUMMIT 26 06
1873s 92 14 104 6525 14630 SOURDOJGH CR 08 08
1873-92 14 104 6527 14523 EAGLE CREEK 14 06
1873s 92 14 104 6526 14524 CRIPPLE CREEK 28 06
1673-92 14 104 6530 14525 EAGLE SUMMIT 25 07
ls73. (92 14 104 6530 14525 EAGLE SUMMIT 25 07
1573-92 14 105 6505 14730 PEDRO JOME 14 07
1873 92 14 105 6505 14730 PEDRO DOME 14 07
1673 «92 14 105 6505 14730 PEDRO DOME 14 07
1473 92 14 105 ©6505 14730 PEDRO JOME 14 08
1673 92 14 105 6512 14805 wICKERSHAM JOM 17 06
1873-92 14 105 6512 14805 WICKERSHAM 30M 17 06
lv73 0s 922 14 105 6512 14805 WICKERSHAM DOM 04 06
1673-92 14 105 6512 14805 WICKERSHAM DOM 04 07
1873-92 14% 105 6503 14726 CLEARY SUMMIT 15 06
1573 = 92 14 105 6503 14726 CLEARY SUMMIT 15 06
1473s 92 14 105 6519 14818 TATALINA 28 05
1473-92 14 105 6503 14726 CLEARY SUMMIT 15 06
lb73— 92 14 105 6519 14818 TATALINA 28 05
1873 92 14 105 0519 14818 TATALINA 28 05
1873s 92 14 105 6503 14726 CLEARY SUMMIT 24 06
1873 92 14% 106 6518 15100 TEXAS CREEK 13 08
1673 = 92 14 108 6425 14435 GOODPASTER R- 07 07
1873 92 14 111 6537 16805 WALES 19 08
1873-92 14 111 6516 16622 TELLER 23 06
1873 92 14 111 6546 16855 LTL DIOMEDE IS 14 08
1673 92 14 111 6512 16645 PORT CLARENCE 12 07
is73'* 92 14 113 6654 16235 KOTZEBUE 13 08
1873 92 14 114 ©6654 16038 KOBUK RIVER 15 08
1873-92 14 114 6617 16153 CHORIS PEN 15 09
1873 92 14 114 6617 16153 CHORIS PEN 15 09
1473-92 14% 124 6725 15007 WISEMAN 01 08
1873s 92 14 124 6725 15006 WISEMAN 01 o8
1873 92 14 126 ©6706 15815 ONION PORTAGE 24 07
1s73°" “92 14 126 6758 16432 KIVALINA 10 08
1873-92 14 128 6744 16432 KIVALINA 20 06
1473-92 14 129 6816 16535 KUPUK RIVER 06 07
1873 (92 14 130 6830 16351 IPEWIK RIVER 28 07
1873-92 14 130 6848 16420 PITYEGEA RIVER 10 07
1873 92 14 133 6822 15438 UPPER KURUPA R 03 07
1673 92 14 134 6805 15030 ULO LAKE 01 07
1873 92 14 134 6812 15241 CHANDLER LAKE 25 07
1873 92 14 136 6803 14500 OLD JOAN LAKE 16 08
1873 92 14 136 ©6804 14503 OLD JOHN LAKE 16 08
1673 92 14 137 6836 14345 SHEENJEK RIVER 28 06
1873 92 14 138 6950 14220 NUVAGAPAK PT 09 08
1473-92 14 138 6950 14220 NUVAGAPAK PT 09 08
1873 92 14 138 6950 14220 NUVAGAPAK PT 09 08
1673 92 14 138 6926 14347 JAGO RIVER 12 07
1675 92 14 139 6920 14500 LAKE SCHRADER 07
1873 92 14 140 6914 14700 SHAVIOVIK R 0S 06
1673 92 14 140 6925 14910 SADLEROCHIT R 26 07
1873s 992 14 141 6923 15210 UMIAT 28 07
1873 92 14% 141 ©6923 15210 UMIAT 28 07
1673 «92 14 142 6952 15350 IKPIKPUK R 04 09
1673 92 14 145 6902 16350 CAPE BEAUFORT 24 07
1673 92 14 145 6902 16350 CAPE BEAUFORT 27 07
244
949
949
921
9353
955
933
933
933
933
933
933
933
933
960
93a
966
966
966
965
966
966
966
948
938
965
967
965
935
963
951
947
945
947
947
957
953
965
966
964
964
965
956
956
966
965
965
965
965
954
954
953
954
953
953
961
961
956
938
93RA
926
899
938
950
931
931
939
939
967
938
960
963
961
958
952
963
956
967
957
956
966
966
966
957
948
947
948
966
966
947
966
966
COLLECTOR NAVE
H DRURY
4 DRURY
J PALMER
w GEIST
w GEIST
w GEIST
w GEIST
Ww GEIST
w GEIST
W GEIST
w GEIST
Ww GEIST
Ww GEIST
& SAUER
P ANDERSON
PEGAU
PEGAU
PEGAU
PEGAU
PEGAU
PEGAU
WELSH
ANDERSON
ANDERSON
WELSH
VIERECK
HARMS
ANDERSON
“ CHAUVIN
KESSEL
SCAMMAN
SCAYMAN
SCAMMAN
SCAMMAN
© SHELTER
% SMITH
TRENT
TRENT
A VIERECK
A VIERECK
w ARGUS
Ww ARGUS
W
W
L
i ee 2 or
ARGUS
ARGUS
HARMS
L HARMS
L HARMS
L HARMS
6 SMITH
@ SMITH
G SMITH
& SMITH
® SMITH
G SMITH
SPOONER
T NEAN
w JOHNSON
™ ANDERSON
P ANDERSON
F PORSILD
TRELEASE
& ANDERSON
H JORDAL
H ROUSE
H ROUSE
P ANDERSON
™ ANDERSON
SCHWEGER
P ANDERSON
BUCKNELL
® MELCHIOR
HULTEN
wv THOMPSON
HODGDON
“™ CHAUVIN
WIGGINS
SHETLER
SHETLER
SCHALLER
ARGUS
ARGUS
ARGUS
CANTLON
SCHOLANDER
SPETZMAN
SPETZMAN
SUDA
SUDA
F BLACK
ARGUS
ARGUS
OOAD<<FFruvcCcQOdQdONUHTMPEPCMIPCNOCCNONOFTFCABZHPCCHEAADUMNUNUNNNNK CK KOOOOFFTFCCHNOMMMMANcerunecunwmwABDAAMoCmMmoovoooc0cCC ooorz=ze
=t=ezv
z=
COL NO HFRAR + NO
3346
3345
ant
SeNe
SeNe
SeNe
278
onT
O31
17418
CAN
CAN
ALA 2026
ALA 29722
ALA 29781
ALA 29711
ALA 29738
ALA 29423
ALA 29421
ALA 29420
ALA 29418
ALA 29783
ALA 29782
WIS
TSC 258262
ALA 32707
ALA 32703
GWA
SWA
SWA
SWA
GWA
TSC 256252
TSC 256260
TSC 246969
GWA
ALA 32636
TSC 256246
ALA 23890
ALA 22908
GWA
GWA
SWA
GWA
ALA 3824
ALA 10652
ALA 30716
ALA 32689
FSLCN00299
ESUC 297
ALA 22492
ALA 4519
SwWA
5WA
ALA 32654
4LA 32581
ALA 32626
ALA 32600
ALA 10654
ALA 10731
ALA 10732
SWA
GWA
GWA
ALA 19064
ALA 190863
ALA 7094
TSC 256259
TSC 256253
US 1789737
4
TSC 253258
IS 2030195
ALA 2557
ALA 5979
ALA 27552
TSC 256257
ALA 34717
TSC 258254
SLA 26577
5WA
US 2384989
WIS
6H
ALA 23891
US 2264037
BLA 4381
GWA
WIS
GWA
GWA
GWA
64H
US 2031803
TSC 256249
US 2032121
GWA
GWA
TSC 256247
GWA
GWA
Specimens Cited
TaxON SPC S V HYB PROV QUAD LAT LONG LOCALITY DATE
1573 92 14 #4145 0902 16350 CAPE BEAUFORT 23 07
16735 92 14 145 6902 16350 CAPE BEAUFORT 23 07
1673 92 14 145 7038 16002 WAINWRIGHT 25 06
1873 92 14 147 7030 15730 MEADE RIVER PO 16 07
1873 92 14 147 7031 15722 USUKTUK RIVER 07 08
1673 92 14 147 7031 15722 USUKTUK RIVER 07 08
1873 92 14 #147 7028 15725 MEADE RIVER 07 OR
15675 92 14 147 7031 15722 USUKTUK R 07 07
1673 92 44 147 7031 15722 USUKTUK R 07 07
1673 92 14 #148 7049 15442 TESHEKPUK 30 08
13873 92 14 151 7910 14650 BULLEN 03 08
1673 92 14 151 7010 14650 BULLEN 03 08
13873 9e 14 151 7010 14650 BULLEN 03 08
1673 92 14 151 7010 14650 BULLEN 03 08
1873 92 14 151 7010 14650 BULLEN 03 08
1473 92 14% #$§J51 7010 14650 BULLEN 4 8B
1873 92 14% $151 #70910 14650 BULLEN 04 08
1873 92 144 151 7001 14510 KONGANEVIK PT 12 08
1873 92 14 152 7005 14335 BARTER IS 07 08
1lb75 92 14° «152 7005 14355 BARTER IS 07 08
1373 92 14 152 7005 14335 BARTER IS 07 08
1573 92 144 152 7005 14335 BARTER IS 07 08
is73 92 14 152 7005 14335 BARTER IS 26 06
1873 92 14 155 712u 15640 BARROW 1207
18673 92 14 153 712U 15640 BARROW 30 07
lo73 92 14 455 712uU 15640 BARROW 30 07
1873 92 14 4153 7120 15640 BARROW 12 07
1873 92 14 153 7120 15640 BARROW i BE rg
13873 92 14 1535 7120 15640 BARROW 11 07
1673 92 14 153 71235 15629 PT 3ARROW 11 06
1873 92 14 #155 7123 15629 PT BARROW 07 08
1473 92 14 1153 7117 15648 BARROW 20 06
18735 92 14% 155 7117 15648 BAR20W 20 06
1873 92 14 153 7115 15651 BARROW 13 07
1873 92 14 i535 7116 15640 BARROW 04 07
1873 92 14 155 7115 15651 BARROW 13:07
ln73 92 14% ##%41J53 7120 15640 BARROW 04 O08
1673 92 14 153 71148 15636 BARROW 01 07
Io POLARIS
1673 96 02 2 5937 13628 HAINES RD M75 O05 07
14873 96 02 6 5915 12945 CASSIAR 17 06
1873 96 02 16 5831 12434 SUMMIT PASS 20 07
1673 96 02 lo 5831 12434 SUMMIT PASS 18 07
1673 96 v2 lo 5831 12434 SUMYIT PASS 20 07
1673 96 02 lo 5831 12434 SUMMIT PASS 12 06
1873 96 02 lo 5831 12434 SUMMIT PASS 12 06
1873 96 02 lo 5831 12434 SUMMIT PASS 12 06
1673 96 02 21 5720 12802 COLD FISH L 23 07
1873 96 02 21 5720 12802 COLD FISH L er or
1673 96 02 35 5530 12245 AZOUZETTA L 04 0B
1873 96 U7 133 ©6935 13355 RICHARDS IS 21 07
1673 96 07 152 6813 13554 CANOE LAKE 18 07
1873 96 07 152 6813 13554 CANOE LAKE 18 07
1875 96 07 152 6813 13554 CANOE LAKE 18 07
1873 96 U7 152 6813 13554 CANOE LAKE 18 07
1673 96 12 15 6411 14021 BIG SWEDE DOME 14 08
1873 96 12 22 6358 13542 KENO HILL 30 07
1873 96 12 30 0247 13104 MT SHELDON 17 08
18735 96 12 36 6247 13104 MT SHELDON 07 08
1473 96 12 32 6149 13835 PTARMIGAN HART 16 07
1673 96 12 39 6058 13829 KLUANE LAKE 24 07
1873 96 12 39 ©045 13852 KASKAWJLSH GL 25 07
1873 96 12 39 6049 13844 OBSERVATION MT 06 07
1873 96 12 39 0057 13825 SLIMS RIVER 18 07
1673 96 12 39 ©0057 13825 SLIMS RIVER £6 O7
14673 96 12 39 6057 13825 SLIMS RIVER 18 07
1873 96 12 42 6022 13351 MT WHITE 15 08
1473 90 Le 42 of022 13351 MT WHITE 13 08
1673 96 12 151 6853 13906 MT SEDGWICK 19 07
1873 96 14 10 5858 13606 MUIR INLETe GL 29 6
1673 96 14 57 6033 160525 CAPE VANCOUVER 08 08
1573 96 14 57 6000 16600 NUNIVAK IS 22 05
1673 96 14 67 0137 14158 SKOLAI CR 21 07
1873 96 14 67 6137 14156 SKOLAI CR 21 07
1673 96 14 67 6136 14152 RUSSEL: GL 27 07
1673 96 14 61 6240 15230 KUSKOKWIM R 02 09
1673 96 14 61 0240 15230 KUSKOKWIM R 05 08
1673 90 14 Bl 6240 15230 KUSKOKWIM R 29 07
1873 96 14 81 6240 15230 KUSKOKAIM R 16 06
1873 96 14 81 6240 15230 KUSKOKWIM R 15 06
1673 9o 14 84 02368 14340 UPPER COPPER R 06 07
1673 96 14 86 6333 14548 RICH HwY M233 12 07
18673 96 14 86 6318 14520 GULKANA GL 19 07
1873 96 14 86 6318 14520 GULKANA GL 19 07
1873 96 14 86 6305 14536 DENALI HWY 5 26 07
1873 96 14 86 6305 14536 DENALI HWY M5 26 07
1673 96 14 86 6305 14545 DENALI HWY M12 04 08
1873 96 14 86 ©6305 14545 DENALI HWY M12 04 08
COLLECTOR NAME
AAMDNAANODNVAANVDEDADADNADNDANDNDDOVVVVDVDOAD
HH OOOQCC
4~9rrrrocnmrrrrraoVeadc eect lrtltztTLTOoucivwrracae£ dd SZSAZ ACP PaAataATIta =
>ro>rrrsxenntinvne ere
CC Sec Ge oe
ARGUS
A2GUS
LLANO
ARGUS
GEIST
GEIST
GEIST
GEIST
GEIST
BLACK
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
HENRY
ARGUS
ARGUS
ARGUS
ARGUS
BARKALOW JRe
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
MJIRDOCK
MURDOCK
THOMAS
THOMAS
WARD
WAR)
WARD
WIGGINS
WIGGINS
TAYLOR
TAYLOR
RAUe
RAUP
RAuUP
TAYLOR
TAYLOR
TAYLOR
SZCZAWINSKT
SZCZAWINSKI
CALDER
Copy
CODY
CODY
CODY
CODY
CALDER
PORSILD
PORSIL)D
PORSILD
RAP
ANDERSON
MURRAY
MURRAY
RAU?
RAUP
RAUP
RAUP
RAUP
CALDER
ARGUS
MACOUN
MILLER
MURRAY
MURRAY
SCOTT
VIERECK
VIERECK
VIERECK
VIERECK
VIERECK
HULTEN
ANDERSON
VIERECK
VIERECK
VIERECK
VIERECK
WEBSTER
WEBSTER
COL NO H=29 + NO
5474
5411
76A
52R2
SeNe
SeNe
GS eNe
5771
5767
5769
5768
5770
5B°7A
5833
50
58°09
5853
5ayu9
S5RAL
Pil
5172
5796
S707
5171
S157
5156
Q7
97
2011
anti
1193
1995
116
1290°9
19248
A19
372
1N697
1N6u7
in7n2
127
1°9
193
7
191
1unr6
13018
129?1
129295
12976
12975
44R7
1058
11767
1166
14718
94104
1099
5460
12528
12597
195A8
113°7
11239
34440
6468
253C
1043
1042
1919
5348
5262
5181
5034
5012
2219
2175
AitS
B3Al
8361
212
212
SWA
5WA
US 2995283
SwWaA
ALA 29139
ALA 29139
ALA 27870
ALA 27998
BWA
TSC 256248
BWA
GWA
eWA
aWA
BWA
5WA
nwa
ALA 30405
aWA
AWA
GWA
nWA
TSC 258250
GWA
BWA
CWA
AWA
GWA
BWA
5H
a4
ALA 6550
TSC 224R90
TSC 224818
TSC 224014
Qv
ISC 224913
TSC 224873
"ADO 32736
NAD 32740
ALA 20074
ALA 20073
ALA 20075
“AO 32739
N40 «632741
AO 3273R
NAD 32734
.A0 632735
7a0)=— 5570
740 625922
"40 32707
NAD 32705
NAO 32706
NAO 32708
40 32730
CAN 312754
64
ALA 20054
ISC 256474
AWA
mwa
ALA 20072
ALA 20071
5WA
ALA 20055
ALA 20070
NAO 32727
AWA
6H
ALA 2931C
GWA
GWA
GWA
FSLC 2009
FSLC 2005
FSLC 2008
SLC 2006
FSLC 2007
US 2384891
TSC 256245
ALA 8407
GWA
GWA
GWA
DAO 32732
US 2331312
245
Appendix B
TAXON SPC S V HYB PROV QUAD LAT LONG LOCALITY DATE COLLECTOR NAME COL NO HFRS + NO
1873 96 14 87 6334 14939 MCK PK RD M36 08 07 939 A NELSON 3719 ALA 538
1873 96 14 87 ©6334 14939 MCK PK RD M36 08 07 939 A NELSON 3719 ISC 256475
1873 96 14 87 6324 14857 CANTWELL 10 07 927 L J PALMER 1963 ALA 5197
1873 96 14 87 6335 14843 CARLO MT 30 07 967 L A VIERECK A413 GWA
1873 96 14 87 6335 14843 CARLO MT 30 07 967 L A VIERECK 8413 GWA
1873 96 14 87 6324 14735 W FK GLACIER 24 08 956 L A VIERECK 1819 GWA
1873 96 14 88 6328 15010 HIGHWAY PASS 21 07 956 G Ww ARGUS 623 GWA
1873 96 14 88 6328 15010 HIGHWAY PASS 03 07 954 G “¥ FROHNE 54311 ALA 21765
1573 96 14 88 ©6325 15020 CAMP EIELSON 14 08 947 E SCAMMAN 4991 SWA
1873 96 14 88 6325 15015 MCK PK RD “64 03 07 959 L SCHENE SeNe ALA 25319
1873 96 14 88 6330 15015 STONY CR 23 07 959 L SCHENE ALA 25316
1873 96 14 88 6325 15020 CAMP EIELSON 08 07 956 L f VIERECK 1127A ALA 3499
1873 96 14 88 6318 15018 MULDROW GL 17 07 956 L A VIERECK 1389 ALA 11616
1473 96 14 88 6318 15018 MULDROW GL 17 07 956 L A VIERECK 1389 SWA
1873 96 14 88 6331 15004 TOKLAT 22 07 965 S L WELSH 4854 SC 247208
1873 96 14 93 6325 17130 ST LAWRENCE IS 933 0 W GEIST SeNe ALA 29734
1873 96 14 93 6325 17130 ST LAWRENCE IS 933 0 W GEIST ALA 29716
1873 96 14 93 6320 17136 BOXER BAY 20 07 960 E & SAUER WIS
1873 96 14 104 6530 14530 EAGLE SUMMIT 21 07 951 B KESSEL 6 ALA 22927
1573 96 14 104 6530 14530 EAGLE SUMMIT 02 07 937 E SCAMMAN 736A ALA 8391
1873 96 14 104 6532 14525 BONANZA CR 16 07 SUD E SCAMMAN 5275 GH
1673 96 14 104 6530 14530 EAGLE SUMMIT 25 07 936 E SCAMMAN 120 GH
1873 96 14 104 6524 14559 TWELVE VI SUM 14 06 957 S G SHETLER ?6AF ALA 3706
1873 96 14 111 6516 16622 TELLER 06 08 S949 E SCAMMAN 5455 GH
1873 96 14 111 6©520 16629 TELLER 09 09 901 F A WALPOLE 2056 US 379175
1873 96 14 123 6736 14940 WIEHL MT 29 06 962 R BROCKMAN ALA 28442
1873 96 14 137 6836 14345 SHEENJEK R 10 07 956 B KESSEL S150 ALA 5046
1873 96 14 137 6836 14345 SHEENJEK R 10 07 956 B KESSEL S150 GWA
1873 96 14 138 6926 14347 JAGO LAKE 23 07 957 J © CANTLON 571305 SY
98 NOVAE~ANGLIAE
1873 +98 1 9 5857 11355 CARIBOU MTS 29 06 930 H “ RAUP 2154 NY
1873 96 1 34 5343 11313 DAVIS L 22 05 947 G H TURNER 5473 ws
1873 98 1 37 5245 11748 ATHABASCA RIVE 11 08 968 G W ARGUS 6935 GWA
1473 98 1 44 5007 11439 OYSTER CR 962 R T OGILVIE P6214 GWA
1873 98 2 16 5851 12506 RACING R 16 06 960 J A CALDER 25460 ALA 17429
1873 98 2 16 5831 12434 SUMMIT PASS 19 07 943 H ™ RAUP 10661 ALA 19864
1873 98 2 16 5847 12500 MACDONALD CR 25 06 968 S L WELSH 73°27 «GWA
1873 98 2 24 5714 12243 SIKANNI R 01 07 943 H RAUP 10358 ALA 19846
1673 98 2 24 5714 12243 SIKANNI R 01 07 943 H “ RAUP 10357 ALA 19845
1873 98 2 24 5714 12243 SIKANNI R 02 07 943 H “ RAUP 10411 ALA 19848
1473 98 2 24 5705 12235 BEATTON R 23 06 943 H ¥ RAUP 10263 ALA 19870
1873 98 2 24 5705 12235 BEATTON R 23 06 943 H ™ RAUP 1N261 ALA 19869
1873 98 2 24 5705 12235 BEATTON R 23 06 943 H “ RAUP 1N2A1 GWA
1673 98 2 56 5210 12220 WILLIAMS LAKE 06 06 960 J A CALDER 17090 GWA
1873 98 2 64 5130 12045 BRIDGE LAKE PO 25 05 956 J A CALDER 16693 GWA
1673 98 2 72 5050 12140 PAVILION LAKE 22 05 956 J f CALDER 16592 GWA
1473 98 2 74 5049 11616 BRISCO 22 05 958 RL TAYLOR 714 GWA
1873 98 2 79 4930 12030 PRINCETON 17 06 960 J A CALDER 17519 SWA
1673 98 7 307 6303 12855 OGRADY LAKE 28 7 967 W J CODY 16867 DAO 58180
1873 98 7 338 6222 12842 SELWYN MTSe 9 8 967 W J CODY 17868 940 58181
1873 98 7 338 6202 12810 FLAT RIVER 8 8 967 Ww J CODY 17714 AO 58177
1873 98 7 S545 6227 11422 YELLOWKNIFE 16 06 949 Ww J CODY 2144 WIS
1873 98 7 367 6125 12636 Se NAHANNI Re 3 7970 G w SCOTTER 12773 GWA
1873 98 7 369 6158 12325 CLI LAKE 11 8 961 Ww J CODY 12304 DADO 58184
1873 98 7 391 6046 11635 MACK HwY M24 10 07 959 J THIERT 5135 SY 1502781
1673 98 12 11 ©6502 13815 DEMPSTERHWYM75 15 07 964 E HULTEN GWA
1673 96 12 15 6404 13925 DAWSON 12 07 902 J ™ MACOUN 5445 6H
1873 98 12 32 ©6120 13910 ALA HWY M1101 21 08 966 G W ARGUS 6013 GWA
1473 98 12 32 6128 13917 ALA HWY M1110 28 07 966 S L WELSH 6053 SWA
1873 98 12 40 6047 13735 PINE CR 17 06 944 H ™ RAUP 117R84 ALA 19847
1673 98 12 40 6047 13735 PINE CR 02 06 948 H “ RAUP 13022 ALA 19909
1473 96 12 4y 6047 13735 PINE CR 02 06 94AR H “ RAUP 13035 ALA 19907
1873 98 12 40 6047 13735 PINE C8 02 06 948 H “ RAUP 13036 ALA 19905
1673 98 12 40 6047 13735 PINE CR 02 06 948 H “ RAUP 13078 ALA 19908
1873 98 12 40 6047 13735 PINE CR 17 06 944 H “™ RAUP 19891 ALA 19844
1873 98 12 40 ©6047 13735 PINE CR 10 06 948 H ™ RAUP 13059 ALA 19849
1873 96 12 40 6047 13735 PINE CR 02 06 94A H “ RAUP 13099 ALA 19904
1873 98 12 40 6047 13735 PINE CR 10 06 948 H “ RAUP 13058 ALA 19850
1673 98 12 40 6047 13735 PINE CR 02 06 948 H “ RAUP 13037 ALA 199190
1873 98 12 40 6047 13735 PINE CR 02 06 948 H ™ RAUP 13030 ALA 19903
1673 98 12 4Q 6047 13735 PINE CR 17 06 944 H “ RAUP 119891 SWA
1873 98 12 40 6047 13735 PINE CR 10 06 948 H “ RAUP 13059 GWA
1473 98 12 40 6041 13736 MACKINTOSH 17 08 957 WR SCHOFIELN A314 CAN 269641
1673 YH 12 41 6043 135U3 WHITEHORSE 31 07 944 J P ANDERSON 9693 ISC 256740
1873 96 12 41 ©6052 13537 TAKHINI R 11 06 949 J © GILLETT 32AR5 ISC 256140
1873 98 12 41 6051 13534 TAKHINI R 11 06 949 J “ GILLETT 3279 =6TSC 256143
1873 Jo 12 41 ©6032 13454 COWLEY 14 06 932 WwW SETCHELL 332 «GH
1673 98 12 44 6003 12840 WATSON LAKE 25 06 966 G W ARGUS 5044 GWA
1873 98 12 44 ©6003 12840 WATSON LAKE 28 06 966 G Ww ARGUS 5066 SWA
1873 «98 12 44 6003 12840 WATSON LAKE 28 06 966 G W ARGUS 50A7 SWA
1673 oes 12 44 6003 12840 WATSON LAKE 28 06 966 G W ARGUS 5065 WA
1873 98 14 66 6148 14512 wILLOW CREEK 01 08 967 L A VIERECK B4US6 SWA
1873 oT) 14 69 6156 14710 GLENN AWY 4128 13 06 944 J P ANDERSON AULGB TSC 256742
1673 Te) 14 69 6148 14730 GLENN 4wY M113 15 07 948 J © ANDERSON 1N674 SC 256137
1673 98 14 69 6156 14710 GLENN HwY M128 13 06 944 J P ANDERSON A4YUG SWA
1673 98 14 80 6230 15340 FAREWELL MT. 8 8 949 W 4 DRURY 3935 CAN
1873 98 14 80 6231 15354 FAREWELL 2 8 969 K “ REED A5% BYU ABALY
1873 98 14 83 6224 14504 TULSONA CREEK 08 08 960 L f* SPETZMAN 44AO CAN 299627
246
Specimens Cited
TAXON SPC S V HYB PROV QUApn LAT LONG LOCALITY DATE
1873 «98 14% 83 6216 14522 GULKANA BRIDGE 10 06
1873 98 14 85 6323 14321 TANACROSS 27 06
1873 98 14 85 6323 14321 TANACROSS 14 08
1873 98 14 87 6350 14925 SAVAGE RIVER 18 06
1873 98 14 87 6350 14925 SAVAGE RIVER 18 06
1873-98 14 87 ©6350 14925 SAVAGE RIVER 18 06
1873 98 14 100 6453 14750 COLLEGE 09 06
1873 98 14 100 6452 14752 UNIV EXP FARM 22 06
1673 98 14 100 6453 14750 COLLEGE 07 07
1873 98 14 100 6453 14750 COLLEGE 09 06
1873 98 14 100 6453 14750 COLLEGE 07 07
1873 98 14 100 6448 14755 CHENA PUMP RD 22 06
1873-98 14 100 6453 14751 SMITH LAKE 10 08
1873 «98 14 100 6453 14751 SMITH LAKE 14 06
1873 =«98 14 100 6430 14905 NENANA 19 06
1873 98 14 100 6453 14750 COLLEGE 14 06
1673-98 14 100 6448 14744 FAIRBANKS 20 06
1873 98 14 #100 6448 14744 FAIRBANKS 06
1873 98 14 100 6430 14905 NENANA 03 07
1873 98 14 100 6439 14825 WHISKEY IS 16 06
1873 98 14 100 6448 14744 FAIRBANKS 19 05
1673 98 14 100 6452 14752 UNIV EXP FARM 17 06
1873 98 14 100 6444 14805 ROSIE CR 08 09
1873 98 14 100 6444 14805 ROSIE CR 10 07
1873.98 14 100 6444 14805 ROSIE CR 12107
1873 «(98 14 100 6443 14809 SAM CHARLEY IS 08 07
187398 14 100 6443 14809 SAM CHARLEY IS 12 06
1673 98 14 100 6443 14713 MOOSE CREEK 16 06
1873 98 14 100 6450 14915 MINTO 24 06
1473 «98 14 100 6438 14909 MINTO 27 06
1873 «98 14 101 6412 14506 SHAW CR FLATS 22 06
1673 98 14 101 6401 14545 JARVIS CREEK 04 07
1473-98 14 102 6410 14147 FRANKLIN 13 07
1673 «98 14 104 6550 14404 CIRCLE 18 06
1573 498 14 104 6550 14404 CIRCLE 10 08
1873 §=98 14 104 6557 14408 YUKON R 09 07
1673-98 14 104 6557 14408 YUKON R 09 07
1873 98 14 105 6532 14833 LIVENGOOD 08 07
101 1 BRACHYCARPA SSPex BRACHYCARPA
1673 101 1 1 41 5150 11637 WATERFOWL LAKE 05 08
1673 1011 2 6 5927 12912 CASSIAR ROAD 19 06
1a73" LO" 2 2 lo 5845 12540 PETERSEN CREEK 23 08
PETS?) LOL’ 1 2 16 5845 12540 PETERSEN CREEK 23 08
$673 -LoL 2 2 lo 5845 12540 PETERSEN CREEK 23 08
1673 Lod) 1 2 16 5845 12540 PETERSEN CREEK 23 08
1873 101 1 2 16 5845 12540 PETERSEN CREEK 23 08
ba7zs 101" 2 2 16 5845 12540 PETERSEN CREEK 23 08
1873 101 1 2 16 5845 12540 PETERSEN CREEK 23 08
1673 1011 2 16 5845 12540 PETERSEN CREEK 23 08
1673 1011 2 16 5845 12540 PETERSEN CREEK 23 08
1873 LOL 2 16 5858 12545 MUNCHO LAKE 22 06
fs7s 20171 2 lo 5858 12545 MUNCHO LAKE 22 06
1673 101° 2 2 16 5858 12545 MUNCHO LAKE 22 06
1873 1011 2 16 5858 12545 MUNCHO LAKE 22 06
is73 Ou 1 2 lo 5851 12506 RACING RIVER 26 07
1875 LoL 1 2 lo 5839 12446 MACDONALD CR 24 07
1873 vor 2 2 lo 5900 12544 MUNCHO LAKE 29 07
18673 101 1 2 16 5900 12544 MUNCHO LAKE 29 07
14673 101 1 2 lo 5900 12544 MUNCHO LAKE 29 07
1873 101 1 2 24 5030 11603 ATHALMER 02 08
1873 101 1 2 34 5541 12430 MANSON CREEK 29 07
1873 101 1 2 35 5527 12240 PINE PASS 04 08
1673 1021 2 51 5301 11916 MT ROBSON 15 08
fa7s 210401 2 55 5243 12454 ITCHA MTS 16 08
1673 10M 1 2 62 5158 12430 TATLA L PeDO. 27 08
favs 101 1 2 79 4955 12053 ASPEN GROVE 17 06
1873 101 1 7 241 ©6517 12651 NORMAN WELLS O20 ta7
fe7s 104" 4 7 369 0158 12325 CLI LAKE iis ve
18673 101 1 7 4569 6152 12320 LTL DOCTOR L 08 08
tazs 102° 1 7 369 6152 12320 LTL DOCTOR L 08 o8
1873 101 1 12 32 6103 13831 KLUANE LAKE 23 07
1473 101 1 12 35 6155 13238 LOWER LAPIE R 20 06
havar LO%° 1 12 4Q 6041 13736 MACKINTOSH 18 08
1873) 101° 1 12 44 6002 12835 WATSON L 03 08
1873 101 1 12 44 6002 12835 WATSON LAKE 03 08
Lea7s wots ie 44 6002 12835 WATSON L 03 08
loi 4 BRACHYCARPA SSPe NIPHOCLADA
1873 101 4 2 16 5850 12532 TOAD RIVER 26 07
1873 101 4 2 lo 5848 12527 TOAD RIVER 13 06
1873 101 4 2 16 5853 12526 SUMMIT PASS 28 07
1873 101 4 2 16 5831 12434 SUMMIT PASS 15 07
1873 101 4 2 16 5831 12434 SUMMIT PASS 15 07
1673'- 101° 4 2 16 5853 12526 SUMMIT PASS 28 06
1673 101 4 2 16 5831 12434 SUMMIT PASS 15 07
1873 101 4 2 16 5847 12500 MACDONALD CR 25 06
967
944
960
928
928
928
957
956
966
957
966
956
956
956
965
965
927
933
932
965
965
964
966
966
966
966
966
966
965
965
966
968
941
932
963
965
965
944
968
960
966
966
966
966
966
966
966
966
966
966
966
966
966
960
960
943
943
943
953
954
954
956
956
960
956
953
961
961
961
944
944
957
943
94%
943
960
96n
943
943
943
943
943
96R
COLLECTOR NAME
COrrECWFFFFFFrrerr rr err Kec OQOD9D9O9O00 <<<FCF
TITEFEPrczeseEECCCrececLL IIc cagagagagagnqaqaqaqaqaqoaocean
A VIERECK
P ANDERSON
A SPETZMAN
MEXIA
MEXIA
MEXIA
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
HARMS
HARMS
PALMER
PALMER
SETCHELL
VIERECK
VIERECK
VIERECK
VIERECK
VIERECK
VIERECK
VIERECK
VIERECK
VIERECK
VIERECK
VIERECK
VIERECK
WELSH
ANDERSON
SETCHELL
SPETZMAN
VIERECK
VIERECK
ANDERSON
Grreztztzttzcecsz
UrlPpPrPrwrprprrr rp cS
Jo vv D>
w ARGUS
A CALDER
Ww ARGUS
w ARGUS
w ARGUS
vw ARGUS
w ARGUS
w ARGUS
w ARGUS
vw ARGUS
vw ARGUS
w ARGUS
w ARGUS
Ww ARGUS
Ww ARGUS
A CALDER
A CALDER
“™ RAUP
mM RAUP
v RAUP
A CALDER
A CALDER
& CALDER
JENKINS
& CALDER
& CALDER
P CALDER
ue Copy
! CODY
1 CODY
J CODY
P ANDERSON
F PORSILD
SCHOFIELD
v RAUP
“ RAUP
“ RAUP
Ps
A CALDER
A CALDER
mM RAUP
” RAUP
¥ RAUP
“ RAUP
“4 RAUP
t WELSH
COL NO
R2N7
8794
4545
2018
2018
2017
1010
425
5112
1010
5111
41
778
312
3721
36?1
1768
175
515
7643
7537
7189
8126
ROU?
B049
BOX
7934
79u3
76u8
7672
79u8
7821
7221
3A5
4967
7704
7705
A969
6877 ¢
25674
6031
6038
6030
4029
6027
6078
6032
6033
6029
79R2
49R3
49RD
YORI
27501
27362
1ngss
10855
19855
11363
14084
7in4
2n172
2nueo
175094
7442
12291
12120
1214
9379
9716
R320
110190
11911
11011
2742
25344
inguy
19592
1n501
inguy
19591
7325
HERB
+ NO
256744
299621
6711
45
256724
299622
256726
19721
13670
5705
46736
46764
269645
19723
19722
19389
19375
19390
247
Appendix B
TAXON SPC S V HYB PROV QUAD LAT LONG LOCALITY DATE
1873 101 4 7 134 6©949 12859 MICHOLSON IS 18 07
1873 101 4 7 134 6944 13120 ATKINSON POINT 21 07
1873 101 4 7 152 6812 13328 CAMPBELL LAKE 03 08
1873 101 4 7 153 6833 12828 ANDERSON RIVER 05 07
1873 101 4 7 153 6833 12828 ANDERSON RIVER 05 07
1873 101 4 7 214 6628 11728 MCTAVISH ARM 30 07
1873 101 4 7 276 6452 12541 NORMAN 13 07
1873 101 4 7 368 6137 12544 VIRGINIA FALLS 29 6
1873 101 4 12 1 6912 13830 KING PT 23 07
1873 101 4 12 2 6922 13932 FIRTH Re P07
1873 101 4 12 4 6823 13908 SAM LAKE 9 7
1873 101 4 12 5 ©6725 14059 RAMPART HOUSE 27 06
1873 101 4 12 7 6602 13844 CATHEDRAL ROCK 29 06
1873 101 4 12 32 6103 13831 KLUANE LAKE 23 07
1873 101 4 12 32 6103 13831 KLUANE LAKE 23 07
1873 10% 4 12 32 6122 13859 BURWASH 03 08
1873 101 4 12 32 6122 13859 BURWASH 27 06
1873 101 4 12 32 ©6122 13859 BURWASH 03 08
1873 101 4 12 32 6122 13859 BURWASH 03 08
1873 101 4 12 32 6103 13831 KLUANE LAKE 1107
1673 101 4 12 32 6103 13831 KLUANE LAKE 02 07
1873 101 4 12 32 6122 13843 KLUANE LAKE 08 07
1873 101 4 12 32 6103 13831 KLUANE LAKE 02 07
1673 101 4 12 32 6103 13831 KLUANE LAKE 11307
1873 101 4 12 32 6108 13833 KLUANE LAKE 05 07
1873 101 4 12 35 6150 13300 ROSE LAPIE R 01 07
1873 101 4 12 35 ©6155 13238 LOWER LAPIE R 20 06
1873 101 4 12 35 6155 13238 LOWER LAPIE R20 06
1873 101 4 12 35 6155 13238 LOWER LAPIE R 20 06
1873 101 4 12 35 6155 13238 LOWER LAPIE R_ 20 06
1873 101 4 12 39 6049 13844 OBSERVATION MT 06 07
1873 101 4 12 39 6055 13843 SLIMS RIVER 29 06
1873 101 4 12 39 6059 13833 SLIMS RIVER 22 07
1873 101 4 12 40 6047 13651 ALA HWY M9868 29 07
1873 101 4 12 40 6047 13738 ALSEK RIVER 24 06
1873 101 4 12 40 6047 13738 ALSEK RIVER 24 06
1873 101 4 12 40 6047 13738 ALSEK RIVER 24 06
1873 101 4 12 40 6047 13738 ALSEK RIVER 24 06
1873 101 4 12 40 6047 13738 ALSEK RIVER 24 06
1873 101 4 12 40 6047 13738 ALSEK RIVER 24 06
1873 101 4 12 40 6047 13738 ALSEK RIVER 24 06
1873 101 4 14 67 6135 14200 WRANGELL MT 31 07
is73' 101° 4& 14 68 6131 14426 CHITINA 07 07
1873 101 4 14 69 ©6132 14914 MATANUSKA 06 07
1873 101 4 14 69 6132 14914 MATANUSKA 06 07
1873 101 4 14 69 6117 14858 MATANUSKA 18 06
1673 101 4 14 71 6155 15425 HEAD OF BIG Re 10 7
1673 101 4 14 71 6155 15425 HEAD OF BIG Re 10 7
1873 101 4 14 71 6155 15425 HEAD OF BIG Re 9 7
1873 101 4 14 71 6155 15425 HEAD OF BIG Re 9 7
18735 101 4 14 71 6155 15430 HEAD OF BIG Re 9 7
1873 101 4 14 71 6155 15430 HEAD OF BIG Re 9 7
1873 101 4 14 71 6155 15425 HEAD OF BIG Re 7 7
1873 101 4 14 71 6155 15425 HEAD OF BIG Re 6 7
1873 101 4 14 80 6233 15336 FAREWELL MT. 8 8
is73 .1017%4% 14 80 ©6233 15336 FAREWELL MT. 8 8
1873 101 4 14 80 6232 15337 FAREWELL Le 6 8
1873 101 4 14 80 6232 15337 FAREWELL Le 6 8
1673 101 4 14 80 6233 15337 FAREWELL Le 20.8
1675: .10146 14 80 6235 15338 FAREWELL Le 2 8
1873 101 4 14 80 6231 15353 FAREWELL 14 «8
1873 101 4 14 61 6240 15230 KUSKOKwWIM R 14 06
1673 101 4 14 86 6355 14656 W FK LTL DELTA 06
1673 101 4 14 86 6355 14656 W FK LTL DELTA 10 07
1673 101 4 14 86 6355 14656 Ww FK LTL DELTA 03 06
1673 101 4 14 86 6325 14545 BLACK RP GLe 16 07
1873 101 4 14 86 6324 14543 CASTNER GL 25 07
1873 101 4 14 86 6324 14543 CASTNER GL 25 07
1873 101 4 14 87 ©6340 14930 TEKLANIKA R 20 07
14673 101 4 14 87 6344 14855 MCKINLEY RIVER 26 07
1873 101 4 14 87 6340 14930 TEKLANIKA R 20 07
1873 101 4 14 87 ©6344 14855 MCKINLEY RIVER 26 07
1673 101 4 14 87 ©6340 14930 TEKLANIKA R 20 07
1873 101 4 14 87 6351 14858 HEALY 27 08
1873 101 4 14 87 ©351 14858 HEALY 27 08
1873 101 4 14 87 6350 14925 SAVAGE R 06 07
1673 101 4 14 87 6350 14925 SAVAGE R 06 07
1673 101 4 14 87 6350 14925 SAVAGE R 06 07
1673 101 4 14 87 ©6350 14925 SAVAGE R 06 07
1873 101 4 14 87 6350 14925 SAVAGE R 06 07
1673 101 % 14 87 6350 14925 SAVAGE R 06 07
18735 101 4 14 87 6350 14925 SAVAGE R 06 07
1673 101 4 14 87 6350 14925 SAVAGE R 06 07
1873 101 4 14 87 6350 14925 SAVAGE R 06 07
1673 101 4 14 87 6350 14925 SAVAGE R 06 07
1673 101 4 14 87 6350 14925 SAVAGE R 06 07
1573 101 4 14 87 6350 14925 SAVAGE R 06 07
1873 101 4 14 67 6350 14925 SAVAGE R 06 07
1673 101 4 14 87 6350 14925 SAVAGE R 06 07
248
965
966
966
965
965
948
947
970
934
970
970
951
960
944
944
948
948
948
94a
944
944
944
944
944
968
944
944
944
944
944
966
967
944
946
944
944
944
944
944
944
944
967
935
931
931
965
950
950
950
950
950
950
950
950
949
949
949
940
949
949
949
961
941
941
943
957
967
967
956
956
956
956
956
950
950
928
926
928
926
928
928
928
928
928
926
926
926
926
926
COLLECTOR NAMF
<a K KK KKK KK KOOUDMMOOOFVOVTFTrrrrrezetSFSsteeEsFTsFEFTTEFEZAFAFAENCCCALTAIAITIIILALIcCIVOPrPrrrvnzrTIITIITIIIIeccececynynroarioaoaoaoa®m
WIE Ps SCS er ECC EL Tere Leth oe tr VN Vee S35 S25 2S 0S Ae SS See eS SSS Se ne Sees SS
SCOTTER
SCOTTER
SCOTTER
SCOTTER
SCOTTER
SHACKLETTE
PORSILD
SCOTTER
PORSILD
WELSH
WELSH
MARTIN
CALDER
ANDERSON
ANDERSON
RAUP
RAUP
RAUP
RAUP
RAUP
RAuUP
RAUP
RAUP
RAUP
WELSH
PORSILD
PORSILD
PORSILD
PORSILD
PORSILD
MURRAY
MURRAY
RAUP
ANDERSON
RAUP
RAUP
RAUP
RAUP
RAUP
RAUP
RAUP
SCOTT
ANDERSON
ANDERSON
ANDERSON
WELSH
DRURY
DRURY
DRURY
DRURY
DRURY
DRURY
DRURY
DRURY
DRURY
DRURY
DRURY
DRURY
DRURY
DRURY
DRURY
VIERECK
PALMER
PALMER
PALMER
VIERECK
VIERECK
VIERECK
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
HANSON
HANSON
MEXIA
MEXIA
MEXIA
MEXIA
MEXIA
MEXTA
MEXIA
MEXIA
MEXIA
MEXIA
MEXIA
MEXIA
MEXIA
MEXIA
cOL NO
6939
1nin7
19125
6910
3245
16705
HERB
GWA
GWA
GWA
GWA
GWA
GWA
6WA
GWA
GWA
8RyY
OTF
GWA
GWA
sc
NIA
ALA
ALA
ALA
ALA
ALA
ALA
ALA
GWA
GWA
GWA
Isc
Isc
GH
NY
US
GWA
GWA
ALA
1sc
ALA
ALA
ALA
ALA
GWA
GWA
GWA
GWA
TSsc
ALA
sc
TSC
CAN
CAN
CAN
CAN
CAN
CAN
CAN
CAN
CAN
CAN
CAN
CAN
CAN
CAN
CAN
FSLC
ALA
ALA
ALA
ALA
SWA
GWA
ALA
GWA
GWA
RM
RM
sc
TSC
+ NO
91102
1204
256220
19940
19941
19935
19942
19939
19466
19388
256681
256215
19565
256218
19936
19937
193684
19385
256217
27348
256221
246203
5199
5194
5156
8385
4751
256358
256358
10197
10200
Specimens Cited
TAXON SPC S V HYB PROV QUAD LAT LONG LOCALITY DATE
1873 101 4 14 87 6334 14936 CATHEDRAL MT
1873 101 4 14 87 6344 14855 MCKINLEY RIVER 23 08
1673 101 4 14 87 6344 14855 MCKINLEY RIVER 23 08
18673 101 4 14 87 6320 14735 WEST FORK GL 24 08
1673 101 4 14 87 6356 14728 ORY CREEK 27 06
1873 101 4 14 87 6340 14935 TEKLANIKA R 03 08
1873 101 4 14 87 6340 14935 TEKLANIKA R 03 08
1673 101 4 14 688 6328 15010 HIGHWAY PASS 21 07
1873 101 4 14 88 6328 15010 HIGHWAY PASS- 21 07
1673 101 4 14 88 6328 15010 HIGHWAY PASS~ 21 07
1873 101 4 14 88 6325 15032 MULDROW GL 23 07
1873 101 4 14 88 6325 15032 MULDROW GL 23 07
1873 101 4 14 88 6325 15032 MULDROW GL 23 07
1873 101 4 14 88 6325 15032 MULDROW GL 23 07
1873 101 4 14 88 6325 15032 MULDROW GL 23 07
1873 101 4 14 88 6325 15032 MULDROW GL 23 07
1873 101 % 14 88 6331 15003 TOKLAT 02 07
1673 101 4 14 88 6331 15003 TOKLAT 15 08
1873 101 4 14 88 6325 15020 THOROFARE R 04 07
1673 101 4 14 88 6325 15020 THOROFARE R 04 07
1673 101 4 14 88 6325 15035 MCKINLEY R 22 06
1673 101 4 14 88 6325 15035 MCKINLEY R 20 07
1873 101 4 14 88 6324 15025 GLACIER CREEK 13 07
1873 101 4 14 88 6330 15050 MCKINLEY RIVER 28 06
1873 101 4 14 868 6324 15025 MULDROW GL 10 07
1873 101 4 14 88 6324 15025 MULOROW GL 10 07
1873 101 4 14 88 6325 15033 MULDROW GL 13 07
1873 101 4 14 88 6324 15032 MULDROW GL 12 08
1873 101 4 14 88 ©6324 15025 MULDROW GL 10 07
1673 101 4 14 88 6325 15035 MCKINLEY R 22 06
1873 101 4 14 88 6324 15025 MULDROW GL 10 07
1873 101 4 14 88 6324 15025 GLACIER CREEK 13 07
1673 101 & 14 95 6458 16310 ETCHEPUK RIVER 05 09
1873 101 4 14 95 6458 16310 ETCHEPUK RIVER 05 09
1873 101 4 14 100 6444 14805 ROSIE CREEK 08 09
1873 101 4 14 100 6444 14805 ROSIE CREEK 28 06
1873 101 4 14 100 6443 14809 SAM CHARLEY IS 11 06
1873 101 4 14 100 6443 14809 SAM CHARLEY IS 08 07
14673 101 4 14 100 6443 14809 SAM CHARLEY IS 02 06
1873 101 4 14 100 6443 14809 SAM CHARLEY IS 11 06
18673 101 4 14 100 6443 14809 SAM CHARLEY IS 08 07
1673 101 4 14 100 6443 14809 SAM CHARLEY IS 02 06
1873 101 4 14 100 6452 14749 COLLEGE 10 06
1873 101 4 14 101 6403 14542 DELTA JUNCTION 26 07
18673 101 & 14 101 6402 14545 JARVIS CREEK 04 06
1873 101 4% 14 101 6406 14546 BIG DELTA 03 07
18673 101 4 14 102 ©410 14136 FRANKLIN ee
1673 101 4 14 102 6410 14136 FRANKLIN ae
1673 101 4 14 102 6410 14136 FRANKLIN 11 07
1673 101 4 14 102 6410 14136 FRANKLIN ee OF
1673 101 4 14 102 6410 14136 FRANKLIN 10°67
1673 101 4 14 102 6410 14136 FRANKLIN 22 oF
1873 101 4 14 102 6410 14136 FRANKLIN 11 07
1673 101 4 14 102 6410 14136 FRANKLIN hie
1873 101 4 14 102 6410 14136 FRANKLIN e"9
1673 101 4 14 102 6410 14136 FRANKLIN 12)>4
1673 101 4 14 102 6447 14112 EAGLE 27 06
1873 101 4 14 102 6447 14112 EAGLE 27 06
1873 101 4 14 102 6447 14112 EAGLE 27 06
1873 101 4 14 102 6447 14112 EAGLE 27 06
1673 101 4 14 102 6443 14117 EAGLE 27 06
1673 101 4 14 102 6443 14117 EAGLE 27 06
1673 101 14 113 6655 16231 KOTZEBJE 12 08
1873 101 4 14 113 6655 16231 KOTZEBUE 12 0A
1673 101 4 14 113 6655 16231 KOTZE8UE 11 08
1673 101 4 14 113 6655 16231 KOTZEBUE 11 08
18673 101 4 14 113 6655 16231 KOTZEBUE 11 08
1873 1ul 4 14 114 6658 16026 KIANA 15 06
1673 101 4 14 119 6635 14510 FORT YUKON 23 06
1673 101 4 14 120 6644 14334 BLACK RIVER 10 07
1673 101 4 14 121 6715 14140 PORCUPINE R 19 07
1873 101 4 14 121 6715 14140 PORCUPINE R 19 07
1673 101 4 14 121 6715 14140 PORCUPINE 8 19 07
1673 101 4 14 121 6715 14140 PORCUPINE R 19 07
1873 101 4 14 121 6715 14140 PORCUPINE R 19 07
1873 101 4 14 121 ©6715 14140 PORCUPINE R 19 07
1873 101 4 14 121 6715 14140 PORCUPINE R 19 07
1873 101 4 14 121 6723 14350 SMALL LAKE 25 07
1873 101 4 14 121 6723 14350 SMALL LAKE 29 07,
1673 101 4 14 121 6723 14350 SMALL LAKE 23 07
1873 101 4 14 124 6725 15007 WISEMAN 31 07
1673 101 4 14 124 6725 15007 WISEMAN 0B 07
1873 101 4 14 124 6725 15007 WISEMAN 20 06
1873 101 4 14 124 ©6725 15007 WISEMAN 20 06
1673 101 4 14 124 6726 15007 WISEMAN 28 06
1873 101 4 14 124 6725 15007 WISEMAN 01 08
1873 101 4 14 124 6725 15007 WISEMAN 01 08
1673 101 4 14 124 6725 15007 WISEMAN 01 08
1673 101 4 14 124 6725 15007 WISEMAN 01 08
1873 101 4 14 125 6757 15310 EASTER CREEK 05 08
1873 101 4 14 126 6706 15815 ONION PORTAGE 05 07
940
940
956
962
964
964
956
956
956
928
928
928
928
928
928
953
950
956
956
958
958
956
958
958
958
956
956
958
958
958
956
966
966
966
967
966
966
966
966
966
966
959
948
951
951
941
941
941
941
94)
941
94)
941
941
941
954
954
954
954
966
966
938
938
966
966
966
937
965
957
957
957
957
957
957
957
957
B57
957
957
939
962
962
962
949
940
940
940
940
965
967
COLLECTOR NAME
CKMMMMFADAACHUNNHOECCCCCCHEODDOCCHUNNHNNNHNNHNCCOCCCOCOCCCCEECFFFFFFFOrewearererreereerrrnr)rerr ye ex~<x<x<<<@Q@Orrrnrrrrer>
Ztetrrvrrerrcao
PrrRrvvUFv FVVRPT Ly
V9 TVDVGVIvIVIVIVVULCVUFYPKORPFF?F?F?
urraanD
o
MURIE
PALMER
PALMER
VIERECK
VIERECK
VIERECK
VIERECK
ARGUS
ARGUS
ARGUS
MEXIA
MEXIA
MEXIA
MEXIA
MEXIA
MEXIA
MURIE
MURIE
VIERECK
VIERECK
VIERECK
VIERECK
VIERECK
VIERECK
VIERECK
VIERECK
VIERECK
VIERECK
VIERECK
VIERECK
VIERECK
VIERECK
PEGAU
PEGAU
VIERECK
VIERECK
VIERECK
VIERECK
VIERECK
VIERECK
VIERECK
VIERECK
VIERECK
ANDERSON
CODY
CODY
ANDERSON
ANDERSON
ANDERSON
ANDERSON
ANDERSON
ANDERSON
ANDERSON
ANDERSON
ANDERSON
ANDERSON
SMITH
SMITH
SMITH
SMITH
WELSH
WELSH
ANDERSON
ANDERSON
ARGUS
AXGUS
ARGUS
CLARK
HARMS
SHETLER
BUCKLEY
BUCKLEY
BUCKLEY
BUCKLEY
BUCKLEY
BUCKLEY
BUCKLEY
SHETLER
SHETLER
SHETLER
ANDERSON
BROCKMAN
BROCKMAN
BROCKMAN
JORDAL
SCAMMAN
SCAMMAN
SCAMMAN
SCAMMAN
STAENIJER
SCHWEGER
COL NO HF&B + NO
GWA
ALA
NA
ALA
GWA
FSLC
FSLC
ALA
GWA
RM
ALA
“IN
A
NY
64
US
BWA
BWA
ALA
BWA
GWA
GWA
GWA
GWA
BWA
GWA
GWA
SWA
FSLC
FSLC
FSLC
FSLC
ALA
5WA
GWA
GWA
GWA
SWA
5WA
GWA
GWA
BWA
6WA
TSC
GWA
GWA
ALA
ALA
ALA
CAN
TSC
1SC
1SC
NA
NA
6H
ALA
ALA
GWA
GWA
GWA
SWA
TSC
NA
GWA
SWA
GWA
ALA
ALA
ALA
ALA
ALA
ALA
ALA
GWA
6WA
GWA
ALA
ALA
ALA
ALA
ALA
ALA
ALA
msc
5WA
SWA
VIN
_
;
WA
ALA
5622
11610
288
292
4752
10207
11609
32699
256182
514
512
537
256361
256362
256549
10642
10643
256213
10189
32599
3968
5100
5120
26765
5997
4697
4131
4664
27557
28501
28432
28508
256212
34707
249
Appendix B
TaxON SPC S V HYB PROV QUAD LAT LONG
1873 101 4% 14 129 6808 16540
1673 101 4 14 129 6808 16540
1873 101 4 14 129 6808 16540
1873 101 4 14 129 6808 16540
1873 101 % 14 129 6817 16532
1873 101 4% 14 129 6817 16532
1873 101 4 14 129 6808 16540
1873 101 4% 14 129 6817 16532
1873 101 4 14 129 6808 16540
1875 101 4 144 129 6845 16610
1873 101 4 14 129 6815 16528
1873 101 4 14 129 6814 16528
1873 101 4 144 #129 6814 16528
1873 102 4 14 131 6855 16110
1875 101 4 14 131 ©6855 16110
1875 101 4 14 137 6839 14100
1873 101 4 14 139 6905 14506
1873 101 4 14 14, 6920 15210
1673 101 4 14 #141 6922 15208
1873 101 4 14° 141 6922 15208
1873 101 4 144% 141 ©922 15208
1875 101 4 14 141 ©6923 15210
1s73 101 4 14° #141 6923 15210
1873 101 4& 14 #141 6923 15210
1873 101 4 14 142 6938 15452
18735 101 4 14 #142 6938 15452
1873 101 4 14% #145 6©902 16350
1673 101 4 14 #145 6902 16350
1875 101 4% 14 146 7010 16000
1873 101 4 14 #146 7008 15937
1873 101 4 14 147 7030 15730
1673 101 4 14° 147 7030 15730
1873 101 4 14 147 7030 15730
1873 101 4 14 147 7030 15730
1875 101 4 14° #147 7030 15730
1875 101 4 14 #147 7030 15730
1873 101 4 144 #147 7030 15730
1873 101 4 14 147 7030 15730
1873 101 4 14 247°) 7030 15750
1673 101 4 14 #147 7030 15730
1873 #101 4 14 #147 «7030 15730
16735 101 4 14° #147 7030 15730
1673 101 4 1G: 147) 7030) 45750
1673 101 4 14 #147 7030 15730
1873 101 4 14. 147 7030 15730
1675 101 4 10 247 7OS0 15750
1375 101 4 14 #147 7030 15730
1873 101 4 14 #147 7030 15730
1873 101 4 14° #147 7030 15730
102 1 RETICULATA SSPe RETICULATA
1873 102 1 2 2 5935 13629
l1s753 102 1 2 2 5935 13630
1873 102 1 2 2 5935 13629
1673 102 1 2 2 5938 13628
1873 102 1 2 16 5831 12434
1873 1021 2 24 5720 12356
1875 102 1 7 133 6944 13230
1873 102 1 7 135 ©6944 13230
1675 102 1 7 134 6942 12900
1873 lve 1 7 134 6942 12903
1873 102 1 7 #134 6956 12858
1873 102 1 7 134 6956 12858
1673 102 1 7 #135 6921 13341
1673 102 1 7 140 6905, 26505
1673 102 1 7 152 6813 13554
1673 102 1 7 #152 6824 13408
1873 102 1 7 152 6813 13554
1675. £02 1 7 241 6500 12715
18735 102 1 7 274 6447 12937
1873 102 1 7 274 60447 12937
1673 102 1 7 276 6431 12634
18673 102 1 7 276 6431 12634
1673 102 1 7 290 6418 09603
1673 102 1 7 297 6409 08318
1873 102 1 7 307 6305 12835
1673 102 1 7 307 6331 12840
1873 102 1 7 307 6331 12840
1675 102 1 7 307 6305 12835
1873 102 1 7 309 6308 12630
1673 102 1 7 309 6308 12630
1873 «102 1 7 335 6345 06832
16735 102 1 7 336 6224 12714
1873 102 1 7 338 6224 12714
1873 102 1 7 339 6205 12735
1873 102 1 7 339 ©6205 12735
2a75° 610241 7 368 6120 12428
16735 102 1 7 402 6050 09425
1873 102 1 7 402 6050 09425
250
LOCALITY
OGOTORUK CR
OGOTORUK CR
OGOTORUK CR
OGOTORUK CR
KUKPUK RIVER
KUKPUK RIVER
OGOTORUK CR
KUKPUK RIVER
OGOTORJK CR
UKINYIK CREEK
KUKPUK RIVER
KUKPUK RIVER
KUKPUK RIVER
DRIFTWOOD CR
DRIFTWOOD CR
FIRTH RIVER
CANNING RIVER
UMIAT
UMIAT
UMIAT
UMIAT
UMIAT
UMIAT
UMIAT
IKPIKPUK RIVER
IKPIKPUK RIVER
CAPE BEAUFORT
CAPE BEAUFORT
KETIK RIVER
KETIK RIVER
MEADE RIVER
MEADE RIVER
MEADE RIVER
MEADE RIVER
MEADE RIVER
MEADE RIVER
MEADE RIVER
MEADE RIVER
MEADE RIVER
MEADE RIVER
MEADE RIVER
MEADE RIVER
MEADE RIVER
MEADE RIVER
MEADE RIVER
MEADE RIVER
MEADE RIVER
MEADE RIVER
MEADE RIVER
PO
PO
PO
PO
PO
PO
PO
PO
PO
PO
PO
PO
PO
PO
PO
PO
PO
PO
PO
MT GLAVE
INSPECTOR CREE
THREE SUARDSME
HAINES RD MB2
SUMMIT PASS
FAIRY LAKE
WARREN PT.
WARREN POINT
ANDERSON Re
ANDERSON R
NICHOLSON PEN
NICHOLSON PEN
KITTIGAZIUT
VICTORIA IS
CANOE Le
REINDEER STAe
CANOE wv
DODO CANYON
KEELE Re
KEELE 2
STERILE Le
STERILE L
BAKER LAKE
CORAL HARBOUR
LTLe DIVIDE L
JUNE Le
JUNE L
LTL DIVIDE L
DAL Le
DAL LAKE
FROBISHER
MACKENZIE
MACKENZIE
BRINTNELL
BRINTNELL LAKE
SeNAHANNI Reo
MH MCCONNELL R
MH MCCONNELL R
BAY
MTS
MTS
LAKE
DATE
28
28
28
28
15
06
06
06
06
o
arINN
oo
NO
oo
oo
NNNOONDONWONNANONNNDWBDDNDWONNA
°o
oo oo
ONNDO
959
959
959
959
959
959
959
959
959
960
960
960
960
969
969
961
947
951
948
953
953
966
966
966
959
959
966
966
959
959
966
966
966
966
966
966
966
966
966
966
966
966
966
966
966
966
966
966
966
967
967
967
948
943
960
O57
957
959
959
963
963
940
959
963
957
963
953
963
963
967
967
947
948
967
967
967
967
963
963
94An
963
963
939
939
970
964
963
COLLECTOR NAME
JOHNSON
JOHNSON
JOHNSON
JOHNSON
JOHNSON
JOHNSON
JOHNSON
JOHNSON
JOHNSON
VIERECK
VIERECK
VIERECK
VIERECK
PEGAU
PEGAU
LITTLE?s
SPETZMAN
BUCKLEY
DUTILLY
SMITH
SMITH
SUDA
SUDA
SUDA
GEIST
GEIST
ARGUS
ARGUS
GEIST
GEIST
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
SUDA
SUDA
C711 =>oerrerprzetzttteczte
JR.
“ay GD | Te
CES SETETCTALECZEECLEZETEZESE
<<XDADAD ANAND AGVADANANANGVDANMDNGMNIOIOODGMIO<<<YNnM crmamaarrrryrrrrrrrrrp
=
w ARGUS
w ARGUS
w ARGUS
“ RAUP
“ RAUP
A CALDER
1 CODY
J CODY
W BARRY
W BARRY
A PARMELEE
A PARMELEE
ROBERTSON
vw THOMSON
J CODY
J Copy
J CODY
J CODY
KVALE
KVALE
“ SIMMONS
w SIMMONS
NN FREEMAN
J COpY
J Copy
J COOY
J CODY
' CODY
KVALE
KVALE
SENN
ROWLANDS
ROWLANDS
RAUP
RAUP
SCOTTER
MACINNES
MACINNES
AAQOQLTIMMIEMMETFETESEAS]S]YMMEETSFTSACCCCAAZEcCLTLTIAVDAAAD
rreeteecavse
COL NO HFRA + NO
215A ALA 26714
215A ALA 9187
215 ALA 26693
215 ALA 9193
673 ALA 26713
673 ALA 9191
215A GWA
673 SWA
215 GWA
4451 ALA 13233
4651 ALA 13234
4651 ALA 13235
4651 ALA 13235
16269 GWA
16169 GWA
184R3 GWA
405 ISC 256216
ALA 5032
6288 SC 256214
1847A ALA 10651
1847B ALA 10733
24766 GWA
24466 SWA
24666 GWA
SeNe ALA 28203
SeNe ALA 29309
5665 GWA
5663 GWA
SeNe ALA 28965
SeNe ALA 27597
5316 GWA
5238 GWA
5296 GWA
5315 GWA
5208 GWA
5270 GWA
5274 SwA
52%7 GWA
5275 GWA
5318 SWA
5312 GWA
5317 GWA
52n2 GWA
5252 swWwA
5277 SWA
5253 GWA
5269 SwWA
19666 GWA
21166 GWA
6780 SWA
6724 SWA
6767 GWA
13194 ALA 19378
10465 ALA 19382
27157 ALA 17426
1N?58 AO
10258 "AO
&4R DAD
448 NAO
2550 AO
2550 A090
28 SC 256495
wIS
12987 NAO
9R26 NAO
12987 AO
7698 YAO
241G AO
2416 A0
47 AO
47 AO
2 16¢ 256528
1819 TSC 256529
16559 AO
17°76 DAO
17°76 AO
16859 40
2 AO
2 940
3753 ISC 256531
51 A0
51 40
9253 ALA 16407
9371 ALA 16412
12575 GWA
47 wo
2°?6 UWO
Specimens Cited
TAxON SPC S V HYB PROV QUAD LAT
1873
1873
1873
1873
1873
1873
1873
1673
1873
1873
1873
1873
1873
1873
1873
1873
1873
1873
1873
1873
1873
1873
1873
1873
1873
1873
1873
1873
1873
1873
1873
1873
1873
1873
1873
1873
1873
1873
1873
1873
1873
1873
1873
1873
1673
13873
1873
1873
1873
1873
1873
18673
1873
1873
1873
1873
1873
1873
1873
1673
1873
1873
1873
18673
1873
1873
1673
1873
1873
1873
1873
1873
1873
1873
1873
1873
1873
1873
1873
1873
1873
1873
1873
1873
1873
1873
1873
1873
1873
1873
1873
102
102
102
102
102
102
102
102
102
102
102
102
102
102
102
102
102
102
102
102
102
102
102
102
102
102
102
102
102
102
102
102
102
102
102
102
102
102
102
102
102
102
102
102
102
102
102
102
102
102
102
102
102
102
102
102
102
102
102
102
102
102
102
102
102
102
102
102
102
102
102
102
102
102
102
102
102
102
102
102
102
102
102
102
102
102
102
102
102
102
102
Ll el el el el ce cl el el el ce ce lo lc a el ce el ed lo cl eh ce ee
12
12
12
12
12
12
12
6850
6850
6430
6430
6445
6445
6435
6430
6430
6445
6420
6345
6334
6331
6159
6103
6103
6103
6149
6103
6149
6145
6145
6129
6129
6140
6140
6058
6058
6049
6055
6055
6055
6055
6049
6057
6057
6042
6042
60047
6047
6047
6047
6047
6047
6048
6048
6044
6044
6043
6043
6043
6043
6033
6033
5555
5857
5858
5858
5858
5858
5859
5858
5858
5858
5858
5859
5858
5856
5858
5857
5855
5856
5818
5840
5830
5840
5825
5826
5818
5818
5818
5818
5818
5818
5818
5818
5825
5826
5255
5250
LONG
14033
14033
136828
13828
13830
13830
13820
13828
13828
13830
13200
13858
13728
13028
14037
13821
13821
13631
13835
13831
13835
13258
13258
12941
12941
12825
12825
13829
13829
13844
13843
13843
13843
13843
13844
13825
13825
13748
13748
13735
13735
13735
13735
13735
13735
13737
13737
13632
13632
13503
13503
13503
13503
13306
13306
13415
13602
13606
13606
13606
13606
13606
13606
13606
13606
13606
13606
13606
13606
13606
13602
13603
13603
13424
13410
13415
13410
13540
13435
13423
13423
13423
13423
13423
13423
13423
13423
13540
13435
LOCALITY
FIRTH RIVER
FIRTH RIVER
OGILVIE MTS
OGILVIE MTS
OGILVIE MTS
OGILVIE MTS
CHAPMAN LAKE
OGILVIE MTS
OGILVIE MTS
OGILVIE MTS
BONNET PLUME L
KLONDIK
MCQUESTEN
KEELE LAKE
WHITE RIVER
JENNY LAKE
JENNY L
KLUANE LAKE
PTARMIGAN HEAR
KLUANE L
PTARMIGAN HRT
CANOL RD M118
ROSE*LAPIE R
FINLAYSON R
FINLAYSON R
LTL HYLAND R
LTL HYLAND R
KLUANE L
KLUANE L
OBSERVATION MT
SLIMS RIVER
SLIMS RIVER
SLIMS R
SLIMS R
OBSERVATION MT
SLIMS RIVER
SLIMS R
DEZADEASH R
DEZADEASH R
PINE CR
PINE CR
PINE CR
PINE CR
PINE CR
PINE CR
MACKINTOSH
MACKINTOSH
MT BRATNOBER
MT BRATNOBER
WHITEHORSE
WHITEHORSE
WHITEHORSE
WHITEHORSE
JOHNSON CROSS
JOHNSONS CROSS
CORONATION ISL
MUIR INLET? GL
MUIR INLET? GL
MUIR INLETe GL
MUIR INLET? GL
MUIR INLET? GL
MUIR INLET GL
MUIR INLET? GL
MUIR INLET? GL
MUIR INLET? GL
MUIR INLETe GL
MUIR INLETe GL
MUIR INLETe GL
MUIR INLETe GL
MUIR INLETe GL
MUIR INLET? GL
MUIR INLET
MUIR INLET
JUNEAU
TAKU B NUNATAK
JUNEAU ICE FD
TAKU B NUNATAK
GUSTAVUS.
MENDENHALL GLA
MT ROSERTS TR
MT ROBERTS TR
MT ROBERTS TR
MT ROBERTS TR
MT ROBERTS
MT ROBERTS
MT ROBERTS
MT ROBERTS
GUSTAVUS.
MENDENHALL GLA
1725SEATTU ISLAND
16905
ANANULIAK IS
c—]
NNNNNNNANN NGS
oo
apo
953
962
966
966
966
966
964
963
963
963
966
902
948
966
944
966
966
944
948
944
948
944
944
887
887
903
963
944
944
966
967
967
967
967
966
944
944
965
965
944
944
944
944
944
944
957
957
962
962
944
944
944
944
959
959
959
967
967
967
967
967
967
967
967
967
967
967
967
967
967
967
967
9LM
917
952
952
952
967
967
967
967
967
967
967
967
967
967
967
967
945
962
COLLECTOR NAME
PRPITIICcczecCCzZEVUVVUVUwAAAADVINM
FOMNMNODONWDOVDOONDONODAONGVACMVNONDODONVVDOVOVOVVOOVOOD UNH SPHECCrr#AELITIIAIIP,PrPILTrVOVO0N0UDUCCTC VY
4444
eee eeseteetesennnnnininwod maces
Vn1VUVSP>
ZCECCTVATEC CETTE CTE ZZTEEALEZEEE
ZEEECECTCECEELE
MCEWEN
YOUNGMAN
PORSILD
PORSILD
PORSILD
PORSILD
YOUNGMAN
YOUNGMAN
YOUNGMAN
YOUNGMAN
BUTLER
MACOUN
CAMPBELL
BUTLER
ANDERSON
NEILSON
NEILSON
RAuP
RAuUP
RAuUP
RAUP
PORSILD
PORSILD
DAWSON
DAWSON
YOUNGMAN
YOUNGMAN
ANDERSON
ANDERSON
MURRAY
MURRAY
MURRAY
MURRAY
MURRAY
MURRAY
RAUP
RAuUP
PEARSON
PEARSON
RAUP
RAuUP
RAuUP
RAUP
RAUP
RAuP
SCHOFIELD
SCHOFIELD
SPETZMAN
SPETZMAN
ANDERSON
ANDERSON
PORSILD
PORSILD
SHETLER
SHETLER
KLEIN
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ANDERSON
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
SOULE
JOHNSON
COL NO HERB
CAN
CAN
CAN
CAN
CAN
CAN
CAN
CAN
CAN
CAN
CAN
CAN
CAN
CAN
Isc
CAN
CAN
ALA
ALA
ALA
ALA
Tsc
rs¢e
CAN
CAN
CAN
CAN
tsc
1sc
GWA
GWA
SWA
SWA
GWA
SWA
ALA
ALA
CAN
CAN
ALA
ALA
GWA
ALA
ALA
GWA
CAN
CAN
CAN
CAN
mse
TSsc
5
6H
CAN
CAN
ALA
SWA
GWA
SWA
SWA
GWA
GWA
GWA
SWA
GWA
GWA
GWA
SwWA
SWA
SWA
SWA
SWA
SWA
Isc
ALA
SWA
+ NO
226056
273835
303406
303405
283818
279587
279587
303134
54407
126891
303135
256522
313097
313097
19381
19383
19361
19383
256543
256543
18848
18848
283818
256519
256519
19377
LOATT
293846
19387
19380
19387
19380
269665
269665
279631
274631
256516
256516
297301
23056
85594
4732
256514
251
Appendix B
TAXON SPC
1873
1873
1873
1873
1873
1873
1873
1873
1873
1873
1873
1873
13873
1873
1873
1873
1873
1873
1873
1873
1873
1873
1873
1873
1873
1873
1873
1873
1873
1873
1873
1873
1873
1873
1673
1873
1873
1873
1873
1873
1873
1873
1673
1873
1673
1873
18673
1673
1673
1873
1873
1673
1873
1873
1873
1873
1873
1873
1873
1873
1873
1673
1673
1873
1873
1873
1873
1873
1673
1873
1873
18673
1673
1673
1673
1673
1673
1673
1473
1673
1873
1873
1873
1473
1673
1873
1673
1673
1873
1673
1873
252
102
102
102
102
102
102
102
102
102
102
102
102
102
102
102
102
102
102
102
102
102
102
102
102
102
102
102
102
102
102
102
102
102
102
102
102
102
102
102
102
102
102
102
102
102
102
102
102
102
102
102
102
102
102
102
1u2
102
102
102
102
102
102
102
102
102
102
102
102
1v2
102
102
102
102
102
102
102
102
102
102
102
102
102
102
102
102
102
102
102
102
102
102
S V HYB PROV QUAD LAT
See eee ed ed ee
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
21
22
22
22
22
22
22
23
23
24
25
25
35
35
38
38
38
38
39
43
56
56
56
57
57
57
57
57
59
67
67
68
69
69
69
69
69
69
69
69
69
71
71
75
77
79
79
80
80
80
80
8u
81
84
85
85
85
86
86
86
86
86
86
86
86
Bo
86
86
86
86
87
87
87
87
87
87
87
87
87
87
87
87
87
87
88
88
88
88
88
88
88
5300
5245
5245
5245
5245
5245
$245
5335
5324
5408
5459
5459
5705
5730
5710
5710
5710
5710
5834
5804
6024
6024
6030
6012
6012
60235
6018
6025
6029
6100
6100
6107
6148
6152
6145
6145
6145
6140
6140
6146
6146
6158
6152
6151
6205
6259
6259
6231
6233
6233
6233
6300
6240
6230
6308
6327
6310
6313
6357
6330
6345
6345
6340
6305
6357
6320
6320
6348
6310
6315
6352
6352
6325
6338
6350
6352
6352
6335
6350
6336
6344
6336
6344
6355
6328
6325
6325
6325
6330
6325
6326
LONG
16854
16858
16858
16858
16858
16858
16858
16650
16733
16546
16228
16228
15425
15413
17015
17015
17015
17015
16146
15304
17242
17235
17300
16656
16656
16611
16611
16611
16049
14139
14139
14544
14730
14710
14915
14915
14920
14900
14900
14918
14918
15438
15433
16535
16343
15604
15604
15353
15336
15336
15336
15400
15230
14330
14315
14340
14330
14536
14658
14535
14554
14554
14555
14545
14658
14535
14535
14432
14541
14537
14859
14859
14900
14930
14925
14859
14859
14915
14925
14932
14858
14932
14858
14828
15050
15020
15020
15020
15052
15015
15024
LOCALITY
ANANUDIAK IS
UMNAK ISLAND
UMNAK ISLAND
UMNAK ISLAND
UMNAK ISLAND
UMNAK ISLAND
UMNAK ISLAND
UNALASKA
CHERNOFSKI
AKUTAN ISLAND
COLD BAY
COLD BAY
OLGA BAY
KODIAK REFUGUM
ST PAUL ISLAND
ST PAUL ISLAND
ST PAUL ISLAND
ST PAUL ISLAND
HAGEMEISTER IS
RASPBERRY IS
ST MATTHEW IS
ST MATTHEW IS
ST MATTHEW IS
NUNIVAK ISLAND
NINIVAK ISLAND
NUNIVAK ISLAND
NUNIVAK ISLAND
NUNIVAK ISLAND
CORRAL CR
CHITINA R
CHITINA R
THOMPSON PASS
SHEEP MT
GLENN HWY M128
WILLOW CR RD
WILLOW CR RD
WILLOW CR RD
MATANUSKA VY
MATANUSKA VY
HATCHER PASS
HATCHER PASS
BIG RIVER
HEAD OF BIG Re
SCAMMON BAY
MOUNTAIN VIL
TAKOTNA
TAKOTNA
FAREWELL
FAREWELL Le
FAREWELL MT.
FAREWELL MTe
FAIRWELL
TONZONA R
NABESNA RD M29
TOKSLA HWY M33
MOON LAKE
SLIPPERY RK CR
RICH HwY M202
FK LTL DELTA R
RAINBOw MT
DONNELLY DOME
DONNELLY DOME
DONNELLY DOME
PAXON
FK LTL DELTA R
RAINBOW MT
RAINBOw MT
UPPER ORY CR
FIELDING LAKE
PAXON
HEALY
HEALY
CANTWELL
TEKLANIKA R
SAVAGE &
HEALY
HEALY
SAVAGE R
SAVAGE R
IGLOO CREEK
MCK NAT PK
IGLOO CREEK
MCK NAT PK
DRY CR
WONDER LAKE
CAMP EIELSON
CAMP EIELSON
CAMP EIELSON
WONDER LAKE
CAMP EIELSON
MCK PK RD
DATE
29
03
13
04
07
08
08
07
937
962
962
962
962
962
962
938
941
934
971
971
945
962
938
941
897
914
970
946
938
966
957
938
965
929
929
965
926
925
925
935
948
944
931
931
951
940
940
965
965
950
950
961
941
941
949
949
949
949
953
961
962
944
96
957,
935
941
965
965
964
964
965
941
966
966
957
961
965
939
939
956
957
965
950
950
928
928
939
939
939
939
962
956
956
956
939
962
962
COLLECTOR NAME
J
M
M
M
M
M
M
J
L
I
M
M
=
M
J
L
J
J
W
J
Vv
D
J
6
W
W
D
w
H
H
J
J
J
J
J
G
e
L
i)
i)
W
W
E
H
J
J
W
wW
W
W
S
L
M
J
V
L
J
J
V
v
V
V
V
L
Cc
Cc
L
R
=
J
J
G
F
B
D
D
Y
Y
A
A
A
A
L
p
G
G
G
A
R
R
H STEENIS
JOHNSON
JOHNSON
JOHNSON
JOHNSON
JOHNSON
JOHNSON
COLE
NORBERG
i Saw
WILLIAMS
WILLIAMS
BEALS
JOHNSON
COLE
MACOUN
MACOUN
DICK
EYERDAM
rw sew
HARMS
KLEIN
U
BOS
MILLER
MILLER
SEIM
MILLER
LAING
LAING
oO1NDD
FROHNE
PALMER
PALMER
WELSH
WELSH
DRURY
DRURY
HULTEN
Be sg? it cual mal <ul yr 0 Jo Yn oT © f° J
KYLLINGSTAD
ANDERSON
ANDERSON
DRURY
DRURY
DRURY
DRURY
SMITH
VIERECK
POZE LELIV Vv
SHARROCK
ANDERSON
HARMS
HARMS
HARMS
HARMS
HARMS
HARMS
PALMER
PARKER
PARKER
Sererer ts Pray
f SPETZMAN
SPOONER
WELSH
ARGUS
DEAN
FABER
HANSON
HANSON
MEXIA
MEXIA
NELSON
NELSON
NELSON
NELSON
A VIERECK
Ow Ore
ANDERSON
w ARGUS
w ARGUS
w ARGUS
NELSON
RICHEY
RICHEY
ANDERSON
ANDERSON
ANDERSON
ANDERSON
ANDERSON
ANDERSON
ANDERSON
ANDERSON
ANDERSON
SPETZMAN
ANDERSON
ANDERSON
ANDERSON
ANDERSON
cOL NO
4532
1047
1114
2nk
1067
1010
1031
4213
2855
2971
102
in2
4059
90
5200
3948
5354
al
3880
159C
159C
1573
uy
47
1889
19746
B445
998
998
Bice
1n4
396
4690
4707
4303
3989
GS eNe
7434
7H34
2915
2410
2776
2745.2
1726
50468A
A744
3594
735
2149
512
3576
3535
Pane
2800
4197
534
RMU
RU72
990
P1n0
4315
5760
5760
595
48
50
50
2010
2074
3737
4533
S750
3533
56R0
638
638
628
3976
HERS
WIS
WIS
WIS
WIS
wIs
wIS
wIs
SC
TSC
SC
GWA
GWA
CAN
wis
ISC
1SC
CAN
CAN
GWA
Isc
TSC
ALA
ALA
1Sc
ALA
ALA
ALA
ALA
ALA
CAN
CAN
1SC
tse
TSC
TSC
ALA
ALA
ALA
ALA
TSC
TSC
CAN
nAN
+ NO
256488
256515
256496
256511
256513
18875
93812
256546
256510
32663
6669
256523
32286
2027
23508
31950
2033
256498
256544
256518
256509
27300
21826
5164
5963
247241
247098
IS 2384897
Tsc
Isc
256480
256492
549
246111
256506
256506
4514
24941
31907
256485
256485
10198
10205
256490
256489
546
547
256497
4728
22466
548
26100
26097
Specimens Cited
TAXON SPC
1873
1873
1873
1873
1873
1873
1873
1873
1873
1873
1873
1873
1873
1873
1873
1873
1873
1873
1873
1873
1873
1873
1873
1873
1873
1873
1873
1873
1873
1873
1873
1873
1873
1873
1673
1673
1873
18673
1673
1873
1873
1873
1873
1873
1873
1873
1873
1873
1873
1873
1873
1873
1873
1873
1873
1873
1873
13873
1873
1873
1873
1673
1873
1873
1873
1673
1673
1873
1873
1873
1873
1673
1873
1873
1873
1873
1873
1873
1873
1873
1873
1873
1873
1673
1873
1673
1873
1873
1873
1873
1873
102
102
102
102
102
102
102
102
102
102
102
102
102
102
102
102
102
102
102
102
102
102
102
102
102
102
102
102
102
102
102
102
102
102
102
102
102
102
102
102
102
102
102
102
102
102
102
102
102
102
102
102
102
102
102
102
102
102
102
102
102
102
102
102
102
102
102
102
102
102
102
102
102
102
102
102
102
102
102
102
102
102
102
102
102
102
102
102
102
102
102
S V HYB PROV QUAD LAT
6325
6325
6330
6324
6324
6324
6324
6347
6330
6321
6435
6430
6430
6430
6430
6435
6446
6436
6426
6437
6433
6433
6410
6530
65235
6525
6425
6522
6531
6529
6520
6534
6533
6654
6658
6655
6803
6803
6803
68035
6730
6725
6725
6730
6730
6757
6757
6706
0815
6815
6815
6815
6745
6853
6806
6806
6806
6810
6810
6810
6817
6800
6805
6848
6848
6850
6805
6805
6814
6810
682e
6822
6822
6839
6950
6950
6959
6923
6952
6946
6902
6902
6902
6902
6902
6902
6912
7030
7030
7030
7030
124
125
125
127
1238
129
134
134
136
137
137
137
137
138
138
139
141
142
145
145
145
145
145
145
145
145
147
147
147
147
Te lee ee ee ee eh lh lh oe el el lol el el ee ee
~
+
LONG
15015
15015
15015
15025
15025
15025
15025
17145
17030
17134
16532
16525
16530
16530
16530
16532
16630
14530
16250
16215
14455
14455
14147
14600
14555
14520
14520
14556
14513
14526
14550
16309
16751
16235
16026
14516
14500
14500
14500
14500
15007
15007
15007
15007
15000
15315
15315
15815
16600
16600
16600
16600
16430
16613
16545
16545
16545
16538
16538
16538
16532
16530
16535
16420
16420
15425
15030
15030
15230
14530
14355
14355
14355
14100
14220
14220
14454
15210
15350
16303
16350
16350
16350
16350
16350
16350
16200
15730
15730
15730
15730
LOCALITY
CAMP EIELSON
CAMP EIELSON
STONY CR
MULDROW GL
MULDROW GL
MULDROW GL
MULDROW GL
ST LAWRENCE IS
ST LAWRENCE IS
ST LAWRENCE IS
NOME
NOME
NOME
NOME
NOME
NOME
CAPE
NOME
GOLOVIN
ELIM
BIG DELTA
BIG DELTA
FRANKLIN
TWELVE MI SUM
TWELVE MI SUM
EAGLE SUMMIT
EAGLE SUMMIT
TWELVE MI SUM
MILLER HOUSE
EAGLE SUMMIT
STEES HWY M88
IMURUK L
TIN CITY
KOTZEBUE
KIANA
FT YUKON
OLD JOHN
OLD JOHN
OLD JOHN
OLD JOHN
WISEMAN
WISEMAN
WISEMAN
WISEMAN
WISEMAN
LONELY LAKE
LONELY LAKE
ONION PORTAGE
CAPE THOMPSON
CAPE THOMPSON
CAPE THOMPSON
CAPE THOMPSON
KIVALINA
CAPE LISBURNE
OGOTORUK CREEK
OGOTORUK CREEK
OGOTORUK CREEK
OGOTORUK CREEK
OGOTORUK CREEK
OGOTORUK CREEK
KUKPUK R
KUKPUK R
KUKPUK R
CAPE SABINE
CAPE SABINE
OOLAMNAGAVIK R
ULO L
ULO L
ULO L
ARCTIC VILLAGE
OLD WOMAN CR
OLD WOMAN CR
OLD WOMAN CR
FIRTH RIVER
NUVAGAPAK PT
NUVAGAPAK PT
COLLINSON PT
UMIAT
IKPIKPUK R
POINT LAY
CAPE BEAUFORT
CAPE BEAUFORT
CAPE BEAUFORT
CAPE BEAUFORT
CAPE BEAUFORT
CAPE BEAUFORT
KOKOLIK R
MEADE RIVER PO
MEADE RIVER PO
MEADE RIVER PO
MEADE RIVER PO
WOOLLEY
Cree
962
959
959
958
958
958
956
938
933
960
938
950
954
954
929
966
969
966
938
938
956
956
941
963
967
961
959
957
955
965
964
947
938
938
937
949
957
957
957
957
939
939
962
946
937
965
965
967
960
960
960
960
960
938
966
966
966
959
961
959
960
962
964
958
958
946
963
963
956
965
956
956
956
961
966
966
914
966
947
938
966
966
966
966
966
966
947
966
966
966
966
COLLECTOR NAWE
RICHEY
SCHENE
SCHENE
VIERECK
VIERECK
VIERECK
VIERECK
ANDERSON
GEIST
5 SAUER
ANDERSON
HANSON
HELLER
HELLER
MILLER
PEGAU
PEGAU
WELSH
ANDERSON
ANDERSON
JOHNSON
JOHNSON
ANDERSON
CHAUVIN
HARMS
HERRIEK
P OSFARRELL
SHETLER
6 SMITH
TRENT
A VIERECK
G SIEH
P ANDERSON
p
9
t
DEVUVPrvrv YS
9
‘D
= Vice Vrs
ANDERSON
CLARK
COOK
SHETLER
SHETLER
SHETLER
SHETLER
P ANDERSON
P ANDERSON
BROCKMAN
JOHNSON
SCAMMAN
STAENDER
STAENDER
SCHWEGER
BELSON
BELSON
BELSON
BELSON
BUCKNELL
ANDERSON
ARGUS
ARGUS
ARGUS
JOHNSON
JOHNSON
JOHNSON
MELCHIOR
MELCHIOR
STREVELER
THOMSON
THOMSON
CHAPMAN
CHAUVIN
FLOCK
WIGGINS
HARMS
KESSEL
KESSEL
KESSEL
LITTLE>
ARGUS
ARGUS
JOHANSEN
SUDA
F BLACK
P ANDERSON
w ARGUS
w ARGUS
w ARGUS
WwW
Ww
W
|
DD=z £=t=z=zeu i ste De sks
=
=
wee
c
a
ARGUS
ARGUS
ARGUS
BLACK
w ARGUS
w ARGUS
w ARGUS
Ww ARGUS
MDODMDADONONGVNGVONGVDGDCAXNGVOAMAWWIcHCUNACCOLTLIEPAPANDOCYPFFTrFFTrFrOaAceMEDCCHUNHNMUYHVEOCCCFTFCHUNADWCUNCHEKYPCCUNADABWEAOOTCMocrrnrrrnrrnrrD
JRe
COL NO
SeNe
SoNe
31A0
31A0
310
1079
3698
SeNeo
HERB + NO
ALA 26098
ALA 25319
ALA 25315
LAV
FSLC 346
GWA
ALA 8283
TSC 256526
ALA 29396
WIS
ISC 256525
ISC 256486
ALA 27012
ALA 268624
ALA 2024
GWA
GWA
GWA
TSC 256527
1SC 256507
ALA 7072
ALA
TSC 256493
ALA 23845
ALA 34631
ALA 18480
ALA 9618
ALA 3718
ALA 10734
ALA 30641
FSLC 308
ISC 256545
Isc 256524
Isc 256508
ALA 10191
TSC 256487
ALA 4282
ALA 4268
ALA 4337
ALA 4326
Isc 256505
ALA 27553
ALA 28485
ALA 10192
GWA
SeNe
ALA
ALA 34706
ALA 28752
ALA 28862
ALA 28834
ALA 28806
ALA 26601
TSC 256542
BWA
BWA
GWA
ALA 9178
ALA 17680
SWA
ALA 17835
SWA
wis
WIS
WIS
TSC 256547
ALA 23889
ALA 21752
US 2264121
ALA 32618
ALA 5052
ALA 22872
ALA 22870
SWA
SWA
GWA
CAN
GWA
TSC 256534
ISC 256541
GWA
6WA
SWA
5WA
GWA
GWA
TSC 256537
GWA
GWA
GWA
SWA
253
Appendix B
TAXON SPC S V HYB PROV QUAD LAT LONG LOCALITY DATE COLLECTOR NAME COL NO HERB + NO
1675 102 1 14 147 7030 15730 MEADE RIVER PO 15 07 966 G W ARGUS 5251 GWA
1875 102 1 14 147 7031 15724 MEADE RIVER PO 19 08 960 O W GEIST ALA 29189
1873 102 1 14 147 7000 15700 MEADE RIVER PO 06 08 960 O W GEIST ALA 28183
1873 102 1 14 147 7031 15724 MEADE RIVER PO 06 08 960 O W GEIST ALA 27678
1873 102 1 14 148 7049 15442 HALF MOON 3 30 08 947 R F BLACK ISC 256536
1873 102 1 14 i468 7047 15500 HALF MOON 3 RA 07 962 D L CHESEMORE 3 ALA 29358
1873 102 1 14 149 7033 15142 ATIGARU PT 29 08 947 R G BLACK TSC 256538
1873 102 1 14 151 7010 14650 BULLEN 03 08 966 G W ARGUS 5726 GWA
1873 102 1 14 151 7010 14650 BULLEN 03 08 966 G W ARGUS 5728 GWA
1873 102 1 14 151 7010 14650 BULLEN 03 08 966 G W ARGUS 5727 GWA
1873 102 1 14 151 7010 14650 BULLEN 03 08 966 G W ARGUS 5725 SWA
1873 102 1 14% 152 7005 14335 BARTER ISLAND 07 08 966 G Ww ARGUS 5868 GWA
1873 102 1 14 152 7005 14335 BARTER ISLAND 07 08 966 G W ARGUS 5869 GWA
1873 102 1 14 152 7005 14335 BARTER ISLAND 26 06 951 F S BARKALOWe JR. P10 ISC 256539
1873 102 1 14 153 7123 15628 POINT BARROW 08 947 P F SCHOLANDER TSC 256540
1873 102 1 14 153 7114 15656 BARROW 13 07 952 G H WARD 1117 YSC 224851
102 2 RETICULATA SSPe GLABELLICARPA
1873 102 2 2 44 5305 13300 MOSQUITO LAKE 1 8 964 J A CALDER 36479 NAO 22233
18735 102 2 2 44 5340 13230 MCCINTON BAY 18 6 957 J & CALDER 21621 DAO 22232
1873 102 2 2 52 5245 13200 TAKAKTA LAKE 25 7 964 J A CALDER 36347 DAO 22234
1873 102 2 14 011 5818 13423 MT ROBERTS TR 09 7 967 G W ARGUS 6632 GWA
1875 102 2 14 O11 5818 13423 MT ROBERTS TR 09 7 967 G W ARGUS 6620 GWA
1873 102 2 14 O11 5818 13423 MT ROBERTS TR 09 7 967 G W ARGUS 6633 GWA
1673 102 2 14 O11 5818 13423 MT ROBERTS TR 09 7 967 G W ARGUS 6645 GWA
1875 102 2 14 O11 5818 13423 MT ROBERTS TR 09 7 967 G W ARGUS 6644 GWA
103 3 LANATA SSPe RICHARDSONII
1873 103 3 2 2 5935 13630 INSPECTOR CREE 12 7 967 G W ARGUS 6736 GWA
1873 103 3 2 2 5935 13630 INSPECTOR CREE 12 7 967 G W ARGUS 6736 GWA
1873 103 3 2 2 9935 13629 THREE GUARDSME 14% 7 967 G W ARGUS 6770 GWA
1873 103 3 2 2 5935 13629 THREE SUARDSME 14 7 967 G W ARGUS 6770 GWA
1873 103 3 2 2 5935 13627 HAINES RD M58 O04 08 967 L A VIERECK 8537 GWA
1873 103 3 2 2 5935 13628 HAINES RD M56 O04 08 967 L A VIERECK 8523 GWA
1873 103 3 2 2 5935 13628 HAINES RD M56 L A VIERECK R525 GWA
1873 103 3 2 2 5950 13639 HAINES RD M83 O04 08 967 L 4 VIERECK 8547 GWA
1873 103 3 2 4% 5940 13327 BOULDER CR 09 06 960 J A CALDER 25205 ALA 17424
18673 103 3 2 16 5831 12434 SUMMIT PASS 19 07 943 H ™ RAUP 10669 ALA 19857
1873 103 3 7 133 6938 13254 TOKER PT 28 07 957 W J CODY 10337 DAO 32636
1873 103 3 7 133 6930 13347 RICHARDS IS 24% 07 966 G W SCOTTER 10122 GWA
1673 103 3 7 134 6952 12900 MICHOLSON IS 18 07 965 G w SCOTTER 6942 GWA
1873 103 3 7 134 6938 12843 ANDERSON RIVER 21 07 965 G W SCOTTER 6950 GWA
1873 103 3 7 140 6903 10550 CAMBRIDGE BAY 20 07 938 C © GILLHAM 19 I1SC 256621
1873 103 3 7 152 6818 13340 INUVIK 5 7 957 Ww J CODY 9849 DAO 5740
18735 103 3 7 152 6813 13554 CANOE L. 16 7 963 Ww J CODY 12806 DAO
1875 103 3 7 152 6820 13330 CAMPBELL Le 15 6 927 A © PORSILD 1°03 CAN 48279
1873 103 3 7 208 6349 12728 BANANA Le 5 8 967 Ww J CODY 17506 DAO
1873 103 3 7 245 6543 11847 LEITH PENe NE 19 7 948 H T SHACKLETTE 3090 CAN 199879
1673 103 3 7 306 6310 13008 MACMILLAN PASS 31 8 944 A F PORSILD 11227 CAN 48274
1873 103 3 7 307 6331 12840 JUNE Le 2 68 967 w J! CODY 17288 DAO
1873 103 3 7 307 6305 12850 OGRADY Le NE 29 7 967 W J CODY 17005 5AO
18735 103 3 7 339 6205 12735 BRINTNELL L 05 07 939 H ™ RAUP 9373 ALA 14812
1873 103 3 7 339 6205 12735 BRINTNELL L 05 07 939 H “ RAUP 93°1 ALA 14760
1673 103 3 7 340 6242 12640 LITTLE DAL Le 6 6 967 Ww J CODY 174652 AO
1873 103 3 7 367 6142 12710 HOLE-IN=-WALL=L 10 68 967 W J CODY 17936 AO
1873 103 3 12 2 6930 13920 FIRTH Re 6 8 953 E H MCEWEN 168 CAN 226060
1873 103 3 12 2 6922 13932 FIRTH RIVER 1 7 970 S L WELSH 1N162A OTF 1211
1873 103 3 12 4 6807 13928 BLACK FOX CR 20 7 970 J ¥ RIGBY 97 OTF 1210
1873 103 3 12 5 6725 14100 RAMPART HOUSE 26 07 951 C fF LOAN 666 GWA
1673 103 3 12 15 6404 13925 DAWSON 25 6 914 A EASTWOOD 385 CAN 48312
1873 103 3 12 17 6406 13914 HUNKER CR 24 +7 902 J “ MACOUN SeNe CAN 48308
1873 103 3 12 29 6241 13238 MT-«SHELDON Se 7 8 944 A F PORSILD 11961 CAN 48294
1873 103 3 12 30 6238 13108 MT»SHELDON NE+ 13 8 944 A F PORSILD 11073 CAN 48293
1673 103 3 12 32 6159 14033 WHITE R 21 07 944 J © ANDERSON 9315 ISC 256616
1873 103 3 12 32 6122 13859 BURWASH 03 08 948 H “ RAUP 139%4 ALA 19749
1873 103 3 12 35 6125 13300 CANOL RD MI 85 6 7 944 A © PORSILD 19166 CAN 46300
1873 103 3 12 35 6120 13300 ROSE Re 3 7 944 A © PORSILD 10274 CAN 48301
18673 103 3 12 35 6155 13238 LOWER LAPIE Re 14% 6 944 A F PORSILD 9460 CAN 48296
1673 103 3 12 35 6140 13302 CANOL RD MI105 10 6 944 A © PORSILD 9292 CAN 48297
1873 103 3 12 35 6140 13302 ROSE LAPIE R 10 06 944 A © PORSILD 9300 1SC 256607
1673 103 3 12 35 6155 13238 LOWER LAPIE R 14% 06 944 A © PORSILD 9460 ISC 256615
1673 103 3 12 39 6056 13829 KLUANE L 24 07 944 J P ANDERSON 9411 ISC 256604
1673 103 3 14 23 5353 16632 DUTCH HARBOR 29 05 938 J P ANDERSON 3151 ISC 255843
1673 103 3 14 41 5842 15639 KING SALMON 06 07 952 Ww 8 SCHOFIELD 2010 SWA
1873 103 3 14 67 6129 14253 KENNICOTT 27 05 955 G Ww ARGUS a3 GWA
1673 103 3 14 67 6137 14158 SKOLAI PASS 21 07 967 D © MURRAY 977 GWA
18673 103 3 14 67 6137 14201 SKOLAI RIVER 20 06 967 R W SCOTT 1611 GWA
1873 103 3 14 67 6137 14201 SKOLAI RIVER 04 07 967 R W SCOTT 1679 GWA
1873 103 3 14 67 6137 14201 SKOLAI RIVER 22 07 967 R w SCOTT 1799 GWA
1873 103 3 14 67 6137 14201 SKOLAT RIVER 20 06 967 R w SCOTT 1609 GWA
1873 103 3 14 67 6137 14201 SKOLAI RIVER 23 06 967 R Ww SCOTT 1619 GWA
1873 103 3 14 69 ©6156 14710 GLENN HWY M128 13 06 944 J P ANDERSON 8448 GWA
1873 103 3 14 69 6156 14710 GLENN HWY M128 15 06 944 J © ANDERSON R4AKL SWA
1873 103 3 14 69 6156 14710 GLENN HWY M128 14 06 944 J P ANDERSON A46B TSC 256012
1875 103 3 14 69 6156 14710 GLENN HWY M128 13 06 944 J P ANDERSON B44B TSC 256601
1873 103 3 14 69 6156 14710 GLENN HWY M128 15 06 944 J P ANDERSON BR4B4 TSC 256600
1873 103 3 14 69 6156 14710 GLENN HWY M128 14 06 944 J © ANDERSON 8469 TSC 256602
254
Specimens Cited
TAXON SPC S V HY3 PROV QUAD LAT LONG LOCALITY DATE
1875 103 3 14 69 6245 14915 WILLOW CR RD 28 06
1873 103 3 14 69 6140 14900 MATANUSKA
1873 103 3 14 71 6152 15433 HEAD OF BIG Re 8 6
1873 103 3 14 71 6155 15425 HEAD OF BIG Re 10 7
1873 103 3 14 72 ©0132 15841 NAPAIMIUT 22 06
1873 103 3 14 72 6132 15841 NAPAIMIUT 22 06
1873 103 3 14 73 6137 15930 ANIAK 18 6
1873 103 3 14 75 6151 16535 SCAMMON BAY 16 06
1673 103 3 14 77 6254 16353 HAMILTON 07 07
1873 103 3 14 77 +6254 163535 HAMILTON 07 07
1673 103 3 14 78 6212 15946 HOLY CROSS 22 08
1873 103 3 14 79 6259 15604 TAKOTNA 27 07
1673 103 3 14 79 6259 15604 TAKOTNA 27 07
1673 103 3 14 80 6233 15336 FAREWELL MT. 8 8B
1873 105 3 14 80 6257 15536 MCGRATH 23 8B
1873 103 3 14 80 6257 15536 MCGRATH 23) 8
1873 103 3 14 81 6219 15006 TALKEETNA 10 08
16735 103 3 14 81 6240 15230 KUSKOKWIM R 18 06
1673 103 3 14 81 6240 15230 KUSKOKWIM R 18 06
1673 103 3 14 84 6323 14345 ALA HWY M1345 12 08
1873 103 3 14 85 6319 14236 TETLIN JCT 28 05
1873 103 3 14 86 6347 14545 DONNELLY DOWE 30 06
1873 103 3 14 86 6357 14658 W FK LTL DELTA 06
1873 103 3 14 86 6357 14658 W FK LTL DELTA 06
1873 103 3 14 86 ©6333 14548 RICH HWY M227 27 06
1673 103 3 14 86 6315 14540 RAINBOW MT 05 08
4873 103 3 14 86 6348 14545 DONNELLY DOME 25 07
1873 103 3 14 87 6335 14935 IGLOO CR 28 07
1873 103 3 14 87 6335 14935 IGLOO CR 28 07
1875 103 3 14 87 6340 14930 TEKLANIKA R 21 07
1875 103 3 14 87 6323 14836 CANTWELL 27 07
1873 103 3 14 87 ©6323 14836 CANTWELL 27 07
1873 103 3 14 87 ©6356 14728 DRY CR 23 06
1875 103 3 14 87 ©6356 14728 DRY CR 29 06
1873 103 3 14 87 6335 14845 CARLO 30 07
1873 103 3 14 88 6327 15052 MCKINLEY BAR 29 07
1873 103 3 14 88 6330 15003 TOKLAT 26 06
1673 103 3 14 68 6325 15047 WONDER LAKE 30 07
1873 103 3 14 88 ©6325 15047 WONDER LAKE 30 07
1873 103 3 14 92 6331 16217 STEBBINS 20 06
1873 103 3 14 92 6302 16318 PASTOLIAK 19 07
1873 103 3 14 92 6302 16318 PASTOLIAK 19 07
1873 103 3 14 94 6454 16503 SALMON LAKE 14 07
1873 103 3 14 94 6454 16503 SALMON LAKE 14 07
1873 103 3 14 95 6437 16215 ELIM 31 08
1873 103 3 14 106 6448 14755 CHENA PUMP RD 22 06
1873 103 3 14° #100 6451 14743 FAIRBANKS 15 08
1873 103 3 14 101 6400 14530 CLEARWATER R 21 08
1873 103 3 14 101 6400 14530 CLEARWATER R 21 08
18673 103 3 14 104 6522 14556 TWELVE MILE CR 18 06
1873 103 3 14 104 6522 14543 STEESE HWY “92 22 06
1873 103 3 14 104 6520 14550 STEESE HWY M88 25 07
1873 103 3 14 104 6559 14411 YUKON R 12 07
1873 103 3 14 105 6532 14833 LIVENGOOD 08 07
1873 103 3 14 113 6655 16231 KOTZEBUE 11 08
1873 103 3 14 113 6655 16240 KOTZEBUE 09 07
1875 103 3 14 113 6655 16240 KOTZEBUE 09 07
1875 103 3 14 114 6658 16026 KIANA 04 06
1873 103 3 14 115 6655 15652 KOBUK 18 07
1873 103 3 14 121 6710 14140 OLD RAMPART 21 07
1873 103 3 14 121 6710 14140 OLD RAMPART 21 07
1873 103 3 14 121 6710 14210 PORCUPINE R 27 07
1873 103 3 14 121 6715 14140 PORCUPINE R 27 07
1873 103 3 14 121 6710 14140 OLD RAMPART 21 07
1873 103 3 14 121 6710 14210 PORCUPINE R 27 07
1873 103 3 14 123 6700 15000 KOYUKUK R 01 08
1873 103 3 14 123 ©6700 15000 KOYUKUK R 01 08
1873 103 3 14 123 6700 15000 KOYUKUK R 01 08
1873 103 3 14 124 6925 15007 WISEMAN 01 08
1873 103 3 14 124 6925 15007 WISEMAN 01 08
1873 103 3 14 124 6925 15007 WISEMAN 01 08
18735 103 3 14 124 6728 15015 NOLAN CR 17 06
1873 103 3 14 124 6925 15007 WISEMAN 08 06
1873 103 3 14 124 6925 15007 WISEMAN 29 05
1873 103 3 14 124 6925 15007 WISEMAN 15 06
18735 103 3 14 124 6925 15007 WISEMAN 15 06
1873 103 3 14 124 6925 15007 WISEMAN 29 06
1873 103 3 14 129 6806 16545 OGOTORUK CREEK 11 08
1873 103 3 14 129 6815 16532 KUKPUK R 06 08
1873 103 3 14 129 6845 16602 UKINYAK CR 01 08
1873 103 3 14 129 6845 16602 UKINYAK CR 30 07
1873 103 3 14 134 6808 15007 ULO VALLEY
1873 103 3 14 136 ©6803 14500 OLD JOHN LAKE 08 08
1873 103 3 14 136 6803 14500 OLD JOHN LAKE 06 08
1873 103 3 14 136 6803 14500 OLD JOHN LAKE 08 08
1873 103 3 14 136 6803 14500 OLD JOHN LAKE 06 08
1873 103 3 14% 136 6803 14500 OLD JOHN LAKE 06 08
1873 103 3 14 137 6840 14101 FIRTH RIVER 10 08
18673 103 3 14 137 6822 14355 SHEENJEK R 05 06
1873 103 3 14 137 6822 14355 SHEENJEK R 05 06
1873 103 3 14 137 6822 14355 SHEENJEK R 05 06
941
940
950
950
930
930
949
961
931
931
925
941
941
949
949
949
941
961
961
944
961
957
941
941
947
965
967
956
956
956
967
967
962
962
964
967
959
956
956
938
931
931
966
966
938
956
963
956
956
957
957
964
966
944
966
966
966
937
924
957
957
957
957
957
957
940
940
940
939
939
939
962
962
962
946
946
949
966
964
960
960
957
957
957
954
954
961
956
956
956
COLLECTOR NAME
WWOMMOODYVNYVNEFFTFIDCZEEDAADCCCMMMCECCCCCFOMWMNOACFTOMNNOOCFOCHNOOCFFFOFCFrrrrragorrmrromirrczteteccroomszzt£BEr Cc
GGtreseseveuvuzrrtetVvVe ts anu Dvara
PPrYrPrYrrPrEetErp
rere Se ea 2S oe maa sa ae et 6 oe ee
uu
DADC PrrPeVveEL
ANDERSON
PALMER
DRURY
DRURY
MILLER
MILLER
DRURY
HULTEN
ROUSE
ROUSE
PALMER
ANDERSON
ANDERSON
DRURY
DRURY
DRURY
ANDERSON
VIERECK
VIERECK
RAUP
LITTLE»
ARGUS
PALMER
PALMER
SCAMMAN
VIERECK
VIERECK
ARGUS
ARGUS
ARGUS
VIERECK
VIERECK
VIERECK
VIERECK
VIERECK
FOOTE
SCHENE
VIERECK
VIERECK
ANDERSON
ROUSE
ROUSE
WELSH
WELSH
ANDERSON
ARGUS
VIERECK
ARGUS
ARGUS
SHETLER
SHETLER
VIERECK
VIERECK
ANDERSON
ARGUS
WELSH
WELSH
CLARK
RYNNING
BUCKLEY
BUCKLEY
BUCKLEY
BUCKLEY
BUCKLEY
BUCKLEY
SCAMMAN
SCAMMAN
SCAMMAN
ANDERSON
ANDERSON
ANDERSON
BROCKMAN
BROCKMAN
BROCKMAN
JOHNSON
JOHNSON
JORDAL
ARGUS
MELCHIOR
VIERECK
VIERECK
CHENEY
SHETLER
SHETLER
SHETLER
SMITH
SMITH
HULTEN
KESSEL
KESSEL
KESSEL
COL NO HERB + NO
7093
425
3960
4071
AN10AF
S65AF
1010AF
2554A
2554A
S4
Su
S4
TSC 256606
ALA 5222
CAN
CAN
ALA 2025
TSC 256605
CAN
US 23864902
ALA 2n41
ALA 25430
ALA 29319
ALA 551
™SC 256610
CAN
CAN
CAN
TSC 255972
GWA
FSLC
ALA 19856
GWA
GWA
ALA 5624
TSC 256613
GWA
GWA
SWA
ALA 4735
GWA
BWA
GWA
SWA
GWA
GWA
FSLC 294
SWA
ALA 25274
ALA 11614
GWA
TSC 256603
ALA 25435
ALA 2052
GWA
TSC 253615
Isc 256612
SWA
FSLC 49
ALA 4899
65WA
ALA 3725
ALA 3777
FSLC 307
FSLC
TSC 256609
SWA
GWA
TSC 253687
ALA 10193
ALA 2030
ALA 26779
ALA 5078
ALA 5638
ALA 5099
GWA
GWA
ALA 8396
GWA
SWA
ALA 256611
ALA 27500
ISC 256614
ALA 286513
ALA 28511
ALA 28492
ALA 10187
ALA 27149
TSC 256396
GWA
GWA
ALA 13225
ALA 13227
ALA 32025
ALA 4291
ALA 4261
GWA
ALA 10739
GWA
US 2384903
ALA 3534
ALA 3534
ALA 3534
255
Appendix B
TaxON SPC S V HYB PROV QUAD LAT LONG LOCALITY DATE
1873) 103 3 14 137 6822 14355 SHEENJEK R 05 06
1673 103 3 14 137 6839 14101 FIRTH RIVER 16 06
1873 103 3 14 138 6950 14220 NUVAGAPAK PT 08 08
1873 103 3 14 138 6950 14220 NUVAGAPAK PT 09 08
1873 103 3 14 139 6930 14500 SADLEROCHIT R 02 08
1873 103 3 14 141 6920 15210 UMIAT 22 06
1873 103 3 1% 141 6920 15210 UMIAT 22 06
1873 103 3 14 141 6©922 15210 UMIAT 22 06
1873 103 3 14 141 6922 15210 UMIAT 12 08
1873 103 3 14 142 6952 15442 IKPIKPUK R 21 07
1873 103 3 144 145 6902 16350 CAPE BEAUFORT 25 07
1873 103 3 14 145 6902 16350 CAPE BEAUFORT 24 07
1873 103 3 144 145 6931 16245 KUKPOWRUK R 04 07
1873 103 3 14 146 7008 15937 KETIK R 21 08
1873 103 3 14 146 7008 15942 KUK RIVER 15 07
1873 103 3 14 147 7030 15730 MEADE RIVER 15 07
1873 103 3 14 147 7030 15730 MEADE RIVER 14 07
1873 103 3 14 147 7030 15730 MEADE RIVER 18 07
1873 103 3 14 #147 7031 15724 MEADE R 19 08
1873 103 3 14 147 7028 15725 MEADE R 07 08
1873 103 3 14 147 7031 15722 USUKTUK R 07 07
1673 103 3 14 151 6905 14506 CANNING R 28 07
1873 103 3 14 153 7120 15630 BARROW 21 07
105 1 ROTUNDIFOLIA SSP. ROTUNDIFOLIA
18753 105 1 7 307 6305 12850 0 GRADY LAKE 29 07
1873 105 1 7 307 6305 12850 0 GRADY LAKE rate (thr
1873 105 1 V7 307 “ie5a7 V28LW ETE DIVIDE tL 26 07
1873 1051 12 32 6127 13805 GLADSTONE CR 26 07
1873 105 1 14 15 5132 17900ESEA OTTER PT 27 07
1873 #105 1 14 15 5132 179008SEA OTTER PT 27) OF
1873 105 1 14 15 5132 17900FSEA OTTER PT 27 07
1873 105 1 14 15 5132 17900ESEA OTTER PT 27 07
1875 105 1 14 17 5150 17608 LTL TANAGA IS 07
1873 1051 14 17 5155 17645 ADAK IS 25 06
1873 105 1 14 17 5152 17637 SWEEPER COVE 14 09
1873 105 1 14 18 5207 17430 ATKA 30 06
18735 105 1 14 18 5207 17430 ATKA 30 06
1873 105 1 14 18 5207 17430 ATKA 30 06
1873 105 1 14 18 5207 17430 ATKA 21 09
18673 1051 14 23 5352 16632 UNALASKA 23 07
1875 105 1 14 23 5352 16632 UNALASKA 02 08
1873 105 1 14 23 5352 16632 UNALASKA 12 07
18735 105 1 14 27 5538 15940 FOX BAY 31 07
1673 1051 14 34 5746 15312 TERROR BAY 16 08
1873 105 1 14 34 5730 15413 KODIAK REFUGIM 18 06
1873 105 1 14 34 5746 15224 POPOF ISLAND 08 07
1873 1051 14 34 5715 15330 SAINTS BAY 05 08
1673 105 1 14 34 5747 15247 KODIAK 20 08
1875 105 1 14 35 5705 15425 OLGA BAY 28 06
18735 105 1 14 38 5710 17015 ST PAUL IS 08 07
1873 105 1 14 38 5636 16932 ST GEORGE IS 24 06
1673 105 1 14 38 5636 16932 ST GEORGE IS 24 06
1873 1051 14 39 5834 16146 CAPE PEIRCE ae f
18735 105 1 14 39 5834 16146 CAPE PEIRCE 11 6
1873 105 i 14 50 5939 15128 PALMER CREEK 27 08
1873 105 1 14 56 6024 17238 ST MATTHEW IS 16 07
1873 105 1 14 56 6040 17306 HALL ISLAND 14 07
1873 105 1 14 56 6024 17238 ST MATTHEW IS O01 07
1873 105 1 14 56 6024 17238 ST MATTHEW IS 01 07
1873 105 1 14 56 6024 17238 ST MATTHEW IS 19 07
18673 105 1 14 62 6013 15008 ICEBERG LAKE 04 07
1873 105 1 14 63 6006 14927 MARATHON MT 04 07
1873 105 1 14 63 6023 14950 COOPER MT 20 08
1873 105 1 14 64 6110 14540 WORTHINGTON GL 08 08
1873 105 1 14 68 6059 14702 COLUMBIA GL 26 06
1673 105 1 14 68 6108 14547 THOMPSON PASS 01 08
16735 105 1 14 69 6245 14515 WILLOW CR RD 28 06
1873 105 1 14 69 6245 14515 WILLOW CR RD 12 07
1673 105 1 14 69 6245 14515 WILLOW CR RD 12 07
18673 105 1 14 69 6245 14515 WILLOW CR RD 07 08
1675 105 1 14 69 6245 14515 WILLOW CREEK 19 07
1673 105 1 14 69 6140 14900 MATANUSKA VY
1673 105 1 14 69 6140 14900 MATANUSKA VY
18673 105 1 14 69 6104 14905 GIRDWOOD MINE 07 07
1873 105 1 14 69 6146 14918 HATCHER PASS 16 07
16735 105 1 14 82 6317 14631 MACLAREN GL i1 08
1673 105 1 14 82 6317 14631 MACLAREN GL 11 06
1873 105 1 14 86 6315 14530 GULKANA GL veh laiey
1873 105 1 14 86 6305 14622 PAXON 18 07
18735 105 1 14 87 6351 14858 HEALY 23 07
1873 105 1 14 87 6346 14900 MT HEALY 03 08
1673 105 1 14 87 6344 14855 MCKINLEY PARK 22 07
1673 105 1 14 88 6327 15052 MCKINLEY BAR 17 07
1873 105 1 14 88 6325 15020 MCKINLEY PARK 11 08
1673 105 1 14 8B ©3525 15020 MCKINLEY PARK 11 08
1873 105 1 14 8B 6325 15020 MCKINLEY PARK 11 08
1673 105 1 14 8&8 6328 15014 STONY PASS
1673 105 1 14 93 6322 17117 POWOOILIAK 17 08
256
956
961
966
966
948
951
951
952
964
959
966
966
951
959
951
966
966
966
960
960
960
947
959
967
967
967
966
967
967
967
967
950
966
949
932
932
932
944
938
932
899
913
950
962
899
963
900
939
941
923
923
970
970
949
938
899
966
966
957
951
951
952
957
899
967
941
931
931
949
940
940
940
965
965
958
958
957
968
939
954
954
940
939
939
939
933
COLLECTOR NAME
OTOCODMDDHOAGDGDOSOODCCLTAOMD
Zvectetzt=z
ONPFrrrrqgaocvnoartTTIywrrrere@ececrnroucooccencec I rremMmMmMrmMnwWaASSEWDEMCeOFZZFETrKaAzseEZEECZEEZEE
cCreeeter
zzz
=
Ee i a a
MS occ es =
x
> arr
=zuvuvvu>
Ss ur e£x4t-sr rca
w
KESSEL
LITTLE?
ARGUS
ARGUS
SPETZMAN
BUCKLEY
BUCKLEY
LINDSAY
WEST
GEIST
ARGUS
ARGUS
CHAMBERS
GEIST
WIGGINS
ARGUS
ARGUS
ARGUS
GEIST
GEIST
GEIST
SPETZMAN
GEIST
CODY
CODY
CODY
NEILSON
KLEIN
KLEIN
KLEIN
KLEIN
BANK
HARMS
JORDAL
EYERDAM
EYERDAM
EYERDAM
YORK
ANDERSON
HULTEN
JEPSON
GRIGGS
CLARK
JOHNSON
KINCAID
NYBAKKEN
WALPOLE
LOOFF
COLE
JOHNSTON
JOHNSTON
DICK
DICK
LUTZ
ANDERSON
COVILLE
HARMS
HARMS
KLEIN
KLEIN
CALDER
KLEIN
VIERECK
COVILLE
VIERECK
ANDERSON
ANDERSON
ANDERSON
FROHNE
PALMER
PALMER
PALMER
VIERECK
WELSH
SHACKLETTE
SHACKLETTE
ARGUS
WELSH
ANDERSON
FROHNE
FROHNE
MURTE
NELSON
NELSON
NELSON
WARREN
GEIST
cOL NO
S4
1A492
5879
5913
1032
2256
V7509
SeNe
5557
5440
110
SeNe
12723
5265
5192
5366
SeNe
SeNe
UiaB |
SeNe
HERB + NO
ALA 3534
BWA
BWA
5WA
US 2032216
ALA 5042
GWA
US 2312671
GWA
ALA 27930
GWA
GWA
US 2264235
ALA 27591
US 2263928
GWA
GWA
BWA
ALA 27616
ALA 27869
ALA 29140
TSC 256618
ALA 28925
DAO 32702
DAO 32699
940 32701
rAN 313089
cs
cs
BWA
GWA
BLA 6720
ALA 32641
US 2009694
6H
US 1631382
TSC 256656
ISC 256653
TSC 256654
6H
US 887216
US 1072109
TSC 256655
WIS
US 378363
wis
US 376197
A
ISC 256657
6H
GH 1289508
6WA
GWA
NA 319224
TSC 256640
US 373502
GWA
ALA 32668
ALA 6584
1SC 256637
6H
ALA 23032
GWA
A
BWA
1SC 256629
TSC 256633
ALA 27353
RM
NA
ALA 5965
ALA 5985
6WA
1SC 247139
US 2387739
MICH
GWA
BWA
ISC 256630
RY
ALA 21779
RY
RM
TSC 256628
ALA 552
RM
ALA 29424
Specimens Cited
1873 105
1673 105
14% 153 7125 15630 POINT 3ARROW 11 08
14 153 7125 15650 POINT BARROW 11 08
TAXON SPC S V HYB PROV QUAD LAT LONG LOCALITY DATE
1675 105 1 14 95 6322 17117 POWOOILIAK 17 08
18735 105 1 14 93 6320 17136 BOXER BAY 27 06
1873 105 1 14 93 6320 17136 BOXER BAY I 27 06
1673 105 1 14 93 6342 17029 ST LAWRENCE IS
1673 105 1 14 93 ©6319 17127 SW CAPE 08
1673 105 1 14 93 6334 17053 KANGEE
1875 105 1 14 93 ©6336 17026 MT ATUK 17 07
1873 1051 14 93 6322 17117 POWOOILIAK 17 08
18735 105 1 14 95 6426 16500 CAPE NOME 07 08
16735 105 1 14 104 6529 14525 EAGLE SUMMIT 07 07
1873 105 1 14 104 6529 14525 EAGLE SUMMIT 20 07
1873 105 1 14 110 6536 16313 IMURUK LAKE 14 07
1873 105 1 144 111 65335 16751 TIN CITY 19 08
1873 105 1 14 111 6516 16622 TELLER 08 08
1873 105 1 14 123 6735 14935 WIEHL MT 29 07
1873 105 1 14 123 6735 14935 WIEHL MT 29 06
1873 105 1 14 128 6758 16432 KIVALINA 10 08
1673 105 1 14 129 6853 16613 CAPE LISBURNE 07 08
1873 1051 14 129 6806 16545 OGOTORUK CR 11 08
1873 105 1 14 129 6809 16559 CAPE THOMPSON 06
1673 105 1 14 129 6806 16545 OGOTORUK CR 17 06
1873 105 1 14° #129 6806 16545 OGOTORJUK CR 18 06
1873 105 1 14 129 6806 16545 OGOTORJK CR 20 06
1873 105 1 14 129 6806 16545 OGOTORUK CR 17 06
16735 105° 1 14 129 6806 16545 OGOTORJK CR 22 06
1873 105 1 14 129 6806 16545 OGOTORUK CR 20 06
18673 105 1 14 129 6806 16545 OGOTORUK CR 18 06
£6735" (205 1 14 129 6806 16545 OGOTORJK CR 25 06
14673 105 1 14 129 6806 16545 OGOTORUJK CR 06 07
1873 105 1 14 129 6839 16613 CAPE DYER 15 07
18735 105 1 14 129 6842 16615 CAPE DYER 20 07
1675 105 1 14 130 6855 16430 CAPE SABINE 13 07
1873 105 1 14 130 6855 16430 CAPE SABINE 13 07
1873 «#105 1 14 130 ©6855 16430 CAPE SASBINE 14 07
1875 105 1 14° 136 6808 14532 ARCTIC VILLAGE 20 07
16735 105 1 14 138 ©6950 14220 NUVAGAPAK PT 09 08
1873 105 1 14 138 6950 14220 NUVAGAPAK PT 09 08
18673 105 1 14 139 6923 14404 OKPILAK RIVER 08 08
1675 105 1 14 139 6935 14445 SADLEROCHIT R 17 08
1875 105 1 14 139 6940 14445 SUNSET PASS 13 08
1873 «10S 1 14 139 6920 14500 LAKE PETERS 11 07
te73. 105-1 14 139 6923 14604 EAGLE CR 05 08
1673 105 1 14° #140 6950 14815 FRANKLIN BLUFF 31 07
1873 #105 1 14 145 6946 16303 POINT LAY 16 08
1675 105 1 14 145 6902 16350 CAPE BEAUFORT 28 7
1673 105 1 14 145 6902 16350 CAPE BEAUFORT 25 07
1873 105 1 14 145 ©6902 16350 CAPE BEAUFORT 25 07
1873 105 1 14 145 6902 16350 CAPE BEAUFORT 25 07
1673 105 1 14° #145 6902 16350 CAPE BEAUFORT 23 07
16735 105 1 14 #145 6902 16350 CAPE BEAUFORT 27 07
1673 105 1 14 145 6902 16350 CAPE BEAUFORT 25 07
1873 i105 1 14 145 6902 16350 CAPE BEAUFORT 23 07
1673 105 1 14 145 6©902 16350 CAPE BEAUFORT 25 07
1873 105 1 14 145 6902 16350 CAPE BEAUFORT 25 07
1875 105 1 14° #145 6902 16350 CAPE BEAUFORT 28 7
1873 105 1 14 #145 6902 16350 CAPE BEAUFORT 24 07
1873 105 1 14 145 6902 16350 CAPE BEAUFORT 23 07
18673 «105 1 14 145 6902 16350 CAPE BEAUFORT 25 07
1673 105 1 14 145 6902 16350 CAPE BEAUFORT 25 07
1873 105 1 14 145 6902 16350 CAPE BEAUFORT 25 07
1673 105 1 14° 145 6902 16350 CAPE BEAUFORT 24 07
1873 105 1 14 145 6902 16350 CAPE BEAUFORT 27 07
18735 105 1 14 145 6902 16350 CAPE BEAUFORT 25 07
1873 105 1 14 145 6902 16350 CAPE BEAUFORT 25 07
1873 105 1 14 145 6902 16350 CAPE BEAUFORT 25 07
1673 105 1 14 145 ©6902 16350 CAPE BEAUFORT 25 07
1873 105 1 14 145 6902 16350 CAPE BEAUFORT 27 07
1875 105 1 14 #145 6902 16350 CAPE BEAUFORT 25 07
1873 105 1 14° #145 6902 16350 CAPE BEAUFORT 04 08
16735 105 1 14 145 6902 16350 CAPE BEAUFORT 05 08
1873 105 1 14 146 7038 16002 WAINWRIGHT 04 08
1873 #105 1 14 #147 7030 15730 MEADE RIVER PO 14 07
1673 105 1 14 147 7030 15730 MEADE RIVER PO 14 07
1673 105 1 14 147 7030 15730 MEADE RIVER PO 14 07
1873 105 1 14 147 7030 15730 MEADE RIVER PO 14 07
1673 105 1 14 #147 7030 15730 MEADE RIVER PO 14 07
1873 105 1 14 147 7051 15848 PEARD BAY 06 08
1673 105 1 14 149 7033 15142 ATIGARU PT 29 08
1673 105 1 14 151 7010 14650 BULLEN 03°08
1873 105 1 14 151 7010 14650 BULLEN 05 08
18673 105 1 14 151 7010 14650 BULLEN 03 08
1873 105 1 14 151 7010 14650 BULLEN 03 08
1873 105 1 14 151 7010 14650 BULLEN 03 08
1873 105 1 14 151 7010 14650 BULLEN 03 08
1873 105 1 14 152 7005 14335 BARTER ISLAND 07 08
1873 105 1 14 152 7005 14335 BARTER ISLAND 07 08
1873 105 1 14 4152 7005 14335 BARTER ISLAND 07 08
1873 105 1 14 153 7120 15640 BARROW 03 08
1873 «#105 1 14 153 7125 15630 POINT BARROW 10 07
1
1
933
933
933
933
933
933
933
933
948
937
947
947
938
901
962
962
938
938
966
960
959
959
959
959
961
959
959
959
960
960
960
959
959
959
950
966
966
957
948
948
948
947
958
938
966
966
966
966
966
966
966
966
966
966
966
966
966
966
966
966
966
966
966
966
966
966
966
966
961
961
938
966
966
966
966
966
947
947
966
966
966
966
966
966
966
966
966
938
966
966
966
COLLECTOR NAME
NODC ODANVDV OVD ADAD ADA GVNADVD AOD C AAD AVNVDVNVANDAVNDAVGVANGV GVANGANV AVA ANNAONANOVNDOAODCCFEFFKFOTOCMOODCTFMNUNTLTITL PEP YrPrPAYPPrPracCcAADUACCMMMODODODCIOCSD
ZEEztezeeeE
ZzZECIEVY 07
r7TD7DrrzvVset=e
==
ZEEEVAZATESCTFTSETTEMNNSESTEEEVCCTECSTTFFTETFCETESFEECTEEATATLETETAZEEELTELEZE IEP rvrerD
GEIST
GEIST
GEIST
GEIST
GEIST
GEIST
GEIST
GEIST
LEPAGE
SCAMMAN
SCAMMAN
SIEH
ANDERSON
WALPOLE
BROCKMAN
BROCKMAN
ANDERSON
ANDERSON
ARGUS
BELSON
JOHNSON
JOHNSON
JOHNSON
JOHNSON
JOHNSON
JOHNSON
JOHNSON
JOHNSON
MELCHIOR
VIERECK
VIERECK
SHETLER
SHETLER
SHETLER
JORDAL
ARGUS
ARGUS
CANTLON
SPETZMAN
SPETZMAN
SPETZMAN
SPETZMAN
THOMSON
ANDERSON
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
STONE
STONE
ANDERSON
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
BLACK
BLACK
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ANDERSON
ARGUS
ARGUS
ARGUS
cOL NO
180
o7
HERB + NO
ALA 29412
ALA 29427
ALA 29419
ALA 29394
ALA 29782
ALA 29723
ALA 29394
ALA 29718
ISC 256636
GWA
GWA
ISC 256476
TSC 256639
US 378891
ALA 28552
ALA 28550
TSC 256638
ISC 256646
SWA
ALA 28873
GWA
GWA
GWA
ALA 9161
ALA 17681
ALA 9163
ALA
ALA 9179
ALA 17761
ALA 13223
ALA 13220
MICH
MICH
MICH
MICH
GWA
SWA
GH
US 2032289
US 2032274
US 2032029
TSC 256648
wIS
ISC 256643
GWA
GWA
GWA
GWA
GWA
GWA
GWA
6WA
GWA
GWA
GWA
GWA
GWA
GWA
5WA
GWA
GWA
SWA
GWA
GWA
GWA
GWA
GWA
GWA
RV
Qu
TSC 256684
GWA
GWA
GWA
GWA
GWA
TSC 256244
TSC 256651
GWA
GWA
GWA
GWA
GWA
GWA
GWA
GWA
GWA
TSC 256645
GWA
GWA
GWA
257
Appendix B
TAXON
1873
1873
1873
1873
1873
1873
1873
1873
1873
1873
1673
1873
1873
1873
1873
1873
1873
1873
1873
1873
1673
1873
1873
1673
1873
1873
1873
1673
1873
1873
1873
105 2
1673
1873
1873
1873
1873
1673
1873
1873
1873
1673
1673
1673
1673
1873
1673
1673
1873
1873
1673
1873
1873
1873
1673
1873
1873
1873
1873
1673
1873
1673
1873
1673
1873
1873
1873
1873
1673
109
1873
1873
1873
18673
1673
1673
1673
1873
1873
1873
1873
1673
14673
1573
1673
258
SPC S V HYB PROV QUAD LAT LONG
105 1 14% 153 7125 15630
105 1 14 153 7125 15630
105 1 14 153 7125 15630
105 1 14 153 7125 15630
105 1 14 153 7125 15630
105 1 14 153 7125 15630
105 1 14 153 7125 15630
105 1 14 153 7120 15640
105 1 14 153 7120 15640
105 1 14 153 7120 15640
105 1 14 153 7120 15640
105 1 14 153 7120 15640
105 1 14 153 7120 15640
105 1 14 153 7120 15640
105 1 14 153 7120 15640
105 1 14 153 7120 15640
105 1 14 153 7110 15600
105 1 14 153 7115 15651
105 1 14 153 7049 15442
105 1 14 153 7120 15640
105 1 14 153 7118 15646
105 1 14 153 7030 15600
105 1 4) USSe 7125) 15630
105 1 14 153 7120 15640
105 1 14 153 7108 15701
105 1 14 153 7116 15640
105 1 14 153 7115 15651
105 1 14 153 7115 15651
105 1 14 153 7115 15651
105 1 14 153 7120 15640
105 1 14 153 7120 15640
ROTUNDIFOLIA SSP. DODGEANA
105 2 7 170 0757 13627
105 2 7 274 6427 12801
105 2 7 274 6427 12801
105 2 7 275 6430 12755
105 2 7 275 6430 12755
105 2 7 307 6308 12816
105 2 7 308 6349 12728
105 2 7 308 06328 12723
105 2 7 313 6328 12834
105 2 7 339 6255 12638
105 2 7 339 6255 12638
105 2 7 339 60224 12714
105 2 12 32 6103 13831
105 2 12 32 6115 14010
105 2 12 32 6115 14010
105 2 12 32 6103 13831
105 2 12 32 6103 13831
105 2 12 39 6055 13843
105 2 12 39 6055 13843
105 2 12 39 6049 13844
105 2 12 39 6059 13830
105 2 14 69 ©6245 14515
105 2 14 69 6146 14918
105 2 14 86 6332 14550
105 2 14 86 ©0348 14436
105 2 14 88 6318 15018
105 2 14 8B 6318 15018
105 2 14 8&8 6318 15018
105 2 14 94 6430 16520
105 2 14 105 6535 14725
105 2 14 124 6725 15006
105 2 14 124 6725 15006
105 2 14 124 6725 15006
105 2 14 131 6847 16000
105 2 144 139 6920 14500
105 2 14 140 6925 14910
105 2 14 140 6925 14910
SCOULERIANA
109 2 2 5935 13630
109 2 16 5831 12434
109 2 17 5830 12245
109 2 17 5850 12235
109 2 24 5705 12235
109 2 24 5705 12235
109 2 31 5625 12040
109 2 35 5530 12245
109 2 35 5530 12245
109 2 40 5445 12700
109 2 4O 5445 12700
109 2 40 5445 12700
109 2 44 5340 13245
109 2 44 5340 13245
109 2 44 5310 13201
LOCALITY
BARROW
B3ARROW
BARROW
BARROW
BARROW
BARROW
BARROW
POINT
POINT
POINT
POINT
POINT
POINT
POINT
BARROW
BARROW
BARROW
BARROW
BARROW
BARROW
BARROW
BARROW
BARROW
BARROW
BARROW
POINT 3ARROW
BARROW
BARROW
POINT BARROW
POINT 3ARROW
BARROW
BARROW
BARROW
BARROW
BARROW
BARROW
POINT BARROW
BARROW
RICHARDSON MTS
BOLSTEAD CR
BOLSTEAD CR
CANOL RD MB2E
CANOL RD MIB2E
LTL DIVIDE L
BACKBONE RANGE
SEKWI BROOK
SEKWI 3ROOK
REDSTONE R
REDSTONE R
MACKENZIE MTSe
KLUANE L
STEELE GL
STEELE GLACIER
KLUANE L
KLUANE L
SLIMS RIVER
SLIMS RIVER
OBSERVATION MT
SLIMS RIVER
WILLOW CR RD
HATCHER PASS
RAPIDS RH
UPPER ORY CR
MULDROW GL
MULDROW GL
MULDROW GL
NOME
WHITE MTS
WISEMAN
WISEMAN
WISEMAN
LAKE NOLUCK
LAKE PETERS
SADLEROCHIT R
SADLEROCHIT R
INSPECTOR CREE
SUMMIT PASS
PROPHET RIVER
FORT NELSON
BEATTON R
BEATTON R
TAYLOR
PINE RIVER.
PINE RIVER.
MACLURE LAKEe
MACLURE LAKEe
MACLURE LAKEe
GRAHAM IS
GRAHAM IS
SKIDEGATE
DATE
11
il
11
11
10
11
10
11
12
12
30
12
12
11
31
11
08
20
966
966
966
966
966
966
966
966
966
966
966
966
966
966
966
966
946
950
947
947
959
960
961
961
950
950
952
952
952
966
952
962
944
944
944
944
967
967
944
944
963
963
963
944
967
967
944
944
967
967
966
962
941
965
935
9577,
956
956
956
926
953
939
939
949
950
959
948
948
967
943
966
949
943
943
943
967
967
967
967
967
9577.
957
957
COLLECTOR NAME
HUNODAODICAROCODAADAVVDDNDDDANANGVDGVNVNANGDNDGDNADND
TrwmrreecorvrrrrencrogoolzrzroocmmnmnrrzztrPrrrc
CrEeEtrEta GoOtZORUGVNAZAZSZR ER ZEB EZ EZ CASAS ZS
ann a>
Ne>r-e>errvrwgrdV~=nrAntrnresennAvasA HMC Go
>rePernDrervde
PREZ ES £2 2 5274, =
A
A
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
BLACK
BLACK
BLACK
BLACK
GEIST
GEIST
STONE
STONE
THOMAS
THOMPSON
WARD
WARD
WARD
WELSH
WIGGINS
CALDER
PORSILD
PORSILD
PORSILD
PORSILD
CODY
CODY
PORSILD
PORSILD
KVALE
KVALE
ROWLANDS
ANDERSON
MURRAY
MURRAY
RAuP
RAUP
MURRAY
MURRAY
MURRAY
SPETZMAN
ANDERSON
WELSH
ANDERSON
SPETZMAN
VIERECK
VIERECK
VIERECK
PORSILD
GJAEREVOLL
ANDERSON
ANDERSON
JORDAL
SPETZMAN
SHETLER
SPETZMAN
SPETZMAN
ARGUS
RAUP
ARGUS
GILLETT
RAUP
RAuUP
RAUP
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
CALDER
CALDER
CALDER
COL NO HERB + NO
5996B
6000
5996
5996A
5147
5991
5124
5159
5174
5166
5705
5167
5169
5168
5718
5160
16
4 7ABK
835
625
2079
1150
1105
1196
1105
5695
129°1
34147
R274
8274
11800
11800
166°
17494
11861
11661
24
a1
44
9410B
1429
1429
12167
12168
798
918
514
114
6997
u7N>g
2201
977A
1370
1370
1370
1328
527
5874
5874
2223
4335
3416
968
968
6739
1N761
4968
3003
10038
10078
10010
6N71
6066
6NB7
6086
6NAB
22743
21298
21345
GWA
GWA
GWA
GWA
GWA
GWA
GWA
GWA
GWA
GWA
GWA
GWA
GWA
GWA
SWA
GWA
ISC 256647
1SC 256652
ISC 256649
ISC 256650
ALA 29182
ALA 28096
Qu
Rv
64
Qu
PM
TSC 224822
TSC 224A21
BWA
RM
NAO 32721
CAN 46N63
CAN 46063
CAN 46062
CAN 46062
NAO 32716
MAO 32717
CAN 46061
CAN 46061
NAO 32720
DAO 32719
NAO
ISC 256641
GWA
BWA
ALA 19719
ALA 19720
GWA
GWA
GWA
CAN 274620
TSC 256629
TSC 247096
1SC 256634
ALA 6871
ALA 3502
GWA
6H
CAN 46055
CAN 225053
ALA 27554
ISC 256631
US 2900395
NA 329290
MICH
CAN 212040
NA 321726
GWA
ALA 19836
RWA
ISC 256666
ALA 19935
ALA 19842
ALA 19854
RWA
GWA
GWA
GWA
SWA
SWA
AWA
5WA
Specimens Cited
TaxON SPC S V HYB PROV QUAD LAT LONG LOCALITY DATE
1673 109 2 44 5315 13205 @ CHARLOTTE 13 07
1873 109 2 46 5330 12830 SKEENA RIVER. 22 07
1873 109 2 46 5330 12830 SKEENA RIVER+ 22 07
1873 109 2 52 5230 13120 MORESBY ISLAND 06 07
1873 109 2 52 5302 13155 MORESBY ISLAND 28 06
1873 109 7 339 6205 12735 BRINTNELL L 20 06
1873 109 7 339 6205 12735 BRINTNELL L 22 06
1873 109 7 388 6030 12328 LIARD R VALLEY 8 8
1873 109 7 388 6010 12330 LIARD 2 VALLEY 20 7
1873 109 12 5 6725 14059 RAMPART HOUSE 01 06
1673 109 12) 15 6400 13945 BEAR CREEK b..7
1673 109 12 24 6310 13008 MACMILLAN PASS 31 8
1873 109 12 27 6244 13641 DAWSON RD M162 15 05
1873 109 12 27 6205 13618 CARMACKS 16 06
1873 109 12 34 «6137 13553 CONGLMERATE MT 14 05
1873 109 12 35 6145 13300 ROSE=LAPIE Re 29 6
1873 109 12 35 6155 13238 LOWER LAPIE Re 15 6
1873 109 12 35 6130 13302 UPPER ROSE Re 17 7
1673 109 12 35 6155 13238 CANOL 2D M132 15 06
1873 109 12 35 6155 13238 CANOL RD M132 15 06
1673 109 12 35 6155 13238 CANOL RD M132 30 06
1673 109 12 40 6045 13630 DEZADEASH Re 20 6
1873 109 12 40 6045 13730 HAINES JCT- 13 6
1873 109 12 41 6010 13442 CARCROSS 25 8B
1673 109 12-41) «6043 13503 WHITEHORSE 01 06
1873 109 12 41 6032 13423 ALA HWY M886 24 06
1873 109 12. 42 6054 13258 NISUTLIN Re 23 7
1873 109 12 42 6016 13255 TESLIN LAKE 02 07
1673 109 12 43 6006 13035 RANCHERIA 22 06
1873 109 12 43 6006 13035 RANCHERIA 22 08
1873 109 12 43 6006 13035 RANCHERIA 22 08
1873 109 12 44 6007 12848 WATSON LAKE 07 07
1873 109 14 3 5531 13139 KETCHIKAN 01 08
1873 109 14 3 5526 13140 KETCHIKAN. 9 6
1873 109 14 3 5527 13153 GUARD ISLANDS 07 08
1873 109 14 3 5531 13139 KETCHIKAN 22 06
1873 109 14 3 5531 13139 KETCHIKAN 09 06
1873 109 14 3 5527 13149 PT HIGGINS 10 08
1873 109 14 3 5526 13146 WHIPPLE CREEK 10 08
1873 109 14 4 5529 13239 HOLLIS 23 08
1873 109 14 4 5529 13239 HOLLIS 23 08
1873 109 14 6 5628 13223 WRANGELLe MI+ 21 7
1873 109 14 6 5648 13258 PETERSBURG. ee:
1873 109 14 11 5840 13410 TAKU B=NUNATAK 10 08
1873 109 144 45 5427 13519 SKAGWAY 25 07
1873 109 14 45 5926 13612 BIG BOULDER CR 12 7
1673 109 14 45 5926 13612 BIG BOULDER CR 12 7
1873 109 14 45 5926 13612 BIG BOULDER CR 12 7
1873 109 14 45 5928 13602 MOSQUITO LAKEe 11 7
1873 109 14 45 5926 13612 BIG BOULDER CR 12 7
1873 109 14 45 5907 13520 HAINESr MIe 6 13 7
1873 109 144 45 5907 13520 HAINES* MUD BA 13 7
1873 109 14 45 5907 13520 HAINESe MUD BA 13 7
1873 109 14 45 5927 13519 SKAGWAY 14 05
1873 109 14 45 5927 13519 SKAGWAY 15 05
1873 109 144 45 5913 13525 HAINES 03 08
1873 109 14 45 5928 13602 MOSQUITO L 08 06
1873 109 144 50 5936 15110 ISMAILOF IS 10 06
1873 109 14 50 5939 15132 HOMER 13 06
1873 109 14 62 6029 15103 SOLDOTNA 02 09
1873 109 14 62 6033 15116 KENAI 15 07
1873 109 14 63 6007 14925 RESURRECTION B 16 05
1873 109 144 63 6024 14922 KENAI LAKE 03 06
1873 109 144 63 6024 14922 KENAI LAKE 22 05
1873 109 14 63 6047 14926 HOPE JUNCTION 17 06
1873 109 14 63 6037 14931 SUMMIT LAKE 12 06
1673 109 14 63 6031 14936 STERL HWY M41 12 06
1873 109 14 67 6126 14256 MCCARTHY 08 06
1873 109 14 68 6157 14517 COPPER CENTER 08 06
1673 109 14 68 6131 14426 CHITINA 25 06
1873 109 14% 68 6101 14501 COPPER R FLATS
1873 109 14 68 6101 14501 COPPER R FLATS
1873 109 14 68 6111 14547 WORTHINGTON GL 01 08
1873 109 14 68 0133 14525 RICH HwY M70 01 08
1673 109 14 69 6148 14828 CHICKALOON 07 07
1873 109 14 69 6113 14953 ANCHORAGE 15 06
1873 109 144 69 6113 14953 ANCHORAGE 29 06
1873 109 14 69 6115 14941 FT RICHARDSON 15 06
1873 109 14 69 6148 14745 GLENN HWY M102 10 06
1873 109 14 69 6127 14922 EKLUTNA R 14 06
1873 109 14 70 6145 15003 wILLOw 28 06
1873 109 14 79 6259 15609 TAKOTNA 26 07
1873 109 14 79 6259 15609 TAKOTNA 26 07
1873 109 14 79 6257 15630 OPHIR REGION 10 9
1873 109 14 80 6254 15530 KUSKOKWIM R. 25 8
1873 109 14 80 6255 15509 KUSKOKWIM Re 9 8
1873 109 14 80 6255 15509 KUSKOKWIM Re 9 8
1873 109 14 80 6254 15530 KUSKOKWIM Re. 25 8
1873 109 14 81 6220 15006 TALKEETNA 10 08
1873 109 14 63 6220 14630 LAKE LOUISE 05 06
1873 109 14 84 6255 14342 MENTASTA PASS 28 05
957
967
967
957
957
939
939
959
959
951
948
944
960
932
960
944
944
944
944
944
944
967
965
951
944
966
944
968
948
966
966
949
927
967
901
931
932
967
967
967
957
967
967
966
934
967
967
967
967
967
967
967
967
961
961
967
965
961
967
950
949
951
951
951
967
967
967
961
961
931
960
960
967
967
948
947
948
948
967
967
941
941
941
949
949
950
950
950
941
961
961
COLLECTOR NAME
MmMmMCeezEEZEBaCCCrrrm CrrPrPremMmMmMrrreccecxocrmMmunrmMmmMaqgagMDaODOOCAGVOAADFFTFTAFECAOCCHOCUHV SPY YET STEP YPPYFYPPHEFCZECFTCMZEIICCOOC
U?>> rcreoepeprppr>>pyp PrrYrprrtteztzteczztztty zt cea rpevp ZzveEezteTvurwnmare-anrseesnraAnnwn> PNIYVOG EEC SSrve=zset>
re vErtTrITrEVvvIPr>s
CALDER
ARGUS
ARGUS
CALDER
CALDER
RAUP
RAuUP
JEFFREY
JEFFREY
LOAN
CAMPBELL
PORSILD
CALDER
SETCHELL
CALDER
PORSILD
PORSILD
PORSILD
PORSILD
PORSILD
PORSILD
PEARSON
PEARSON
PORSILD
PORSILD
WELSH
PORSILD
WELSH
ANDERSON
ARGUS
ARGUS
GILLETT
ANDERSON
ARGUS
FLETT
SETCHELL
SETCHELL
VIERECK
VIERECK
HURD
HURD
ARGUS
ARGUS
BESCHEL
ANDERSON
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
LITTLE»
LITTLE?
VIERECK
WELSH
LITTLE»
VIERECK
CHRIST
LUTZ
CALDER
CALDER
CALDER
VIERECK
VIERECK
VIERECK
LITTLE»
LITTLE?
SETCHELL
SHEETS
SHEETS
VIERECK
VIERECK
ANDERSON
DUTILLY
LEPAGE
SPETZMAN
VIERECK
VIERECK
ANDERSON
ANDERSON
ANDERSON
DRURY
DRURY
DRURY
DRURY
DRURY
ANDERSON
LITTLE?
LITTLE»
JR.
JR.
JR.
JRe
JRe
JR.
JR.
COL NO HFRB
22469
6246
6847
22186
21903
9146
9234
497
297
396
296
11221
244%6
367
24403
10126
9507
10451
9598
9507
10033
67-109
318
18459
9144
5515
10791
7615
10467
6014
6015
36273
2A506
6089
1896
3
273
8710
8703
591
591
6844
6240
154Aa4
1772
6693
6691
6690
6675
6696
6751
6741
6742
1R109
1A1?1
8506
4045
1R408
R268
20127
36
4902
5044
4906
R245
R256
R255
14372
1a3n2
, 48
SeNe
SeNe
R474
R462
10539
20076
23373
R216
B2ARL
7011
7415
7415
3472
3186
4536
4535
3187
7678
18365
1R257
GWA
GWA
GWA
GWA
GWA
ALA
ALA
CAN
CAN
CWA
CAN
CAN
GWA
NIA
GWA
CAN
CAN
CAN
TSC
+ NO
14833
14832
260975
260972
126893
49501
332716
49507
49506
49508
256675
256672
256674
316890
293825
208680
256673
49509
25639N
256660
25620n
232913
320470
US 1566845
GWA
GWA
GWA
GWA
GWA
GWA
GWA
Tsc
GWA
GWA
GWA
GWA
GWA
GWA
GWA
GWA
GWA
GWA
GWA
US
SWA
GWA
NA
NA
GWA
GWA
GWA
SWA
GWA
GWA
GWA
GWA
NA
ALA
ALA
GWA
SWA
msc
TSsc
rsc
MIN
SWA
GWA
msc
ALA
rsc
CAN
CAN
CAN
CAN
CAN
sc
GWA
GWA
256658
246236
322259
332911
320468
25133
25123
256676
256663
256662
256659
553
256661
256677
259
Appendix B
TAXON
1573
1873
1873
1873
1873
1873
1873
1873
1873
1873
1873
1873
1873
1473
1873
1673
18735
1873
14673
1873
1873
1873
1873
1873
1673
1873
1873
1873
1873
1873
1873
1873
1873
1873
1673
1873
1673
1873
1873
1873
1873
1873
1873
1673
1873
1873
1873
1873
1673
1873
1873
1873
1673
1873
1873
113
1673
1873
1673
1873
1873
1873
1873
1873
1673
1873
1873
1873
1673
1873
1873
1873
1873
1873
1873
1673
1873
1873
1873
1873
1873
1873
1873
1873
1873
1873
1873
1873
260
SPC S V HYB PROV QUAD
109
109
109
109
109
109
109
109
109
109
109
109
109
109
109
109
109
109
109
109
109
109
109
109
109
109
109
109
109
109
109
109
109
109
109
109
109
109
109
109
109
109
109
109
109
109
109
109
109
109
109
109
109
109
109
113
113
113
113
113
113
113
113
113
113
113
113
113
113
113
113
113
113
113
113
113
113
113
113
113
1135
113
113
113
113
113
113
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
14
SETCHELLIANA
85
85
86
86
86
86
86
86
88
88
89
89
89
89
89
98
100
100
100
100
100
100
100
100
100
100
100
100
100
100
100
100
100
100
100
100
100
100
100
100
100
100
100
100
100
100
100
100
101
101
101
105
105
105
105
32
32
32
32
32
32
32
32
32
32
32
40
40
46
46
67
69
69
69
69
69
69
71
71
80
80
81
82
Be
85
86
86
LAT
6332
6320
6347
6345
6335
6348
6358
6358
6337
6337
6315
6315
6315
6315
6315
6445
6451
6451
6447
6447
6453
6454
6454
6451
6451
64535
6447
6447
6453
6454
6454
6451
6451
6453
6451
6451
6451
6451
6451
6451
6451
6451
6451
6451
6452
6451
6451
6446
6453
6453
6453
6515
6515
6515
6515
6105
6137
6137
6103
6122
6122
6122
6122
6125
6120
6103
6040
6045
5910
5910
6105
6132
6136
6136
6132
6136
6136
6155
6155
6231
6230
6240
6228
6228
6345
6346
6342
LONG
14225
14258
14545
14443
14430
14545
14545
14545
15001
15001
15442
15442
15442
15442
15442
15530
14759
14759
14805
14805
14752
14751
14751
14759
14752
14800
14805
14805
14752
14751
14751
14759
14752
14800
14759
14759
14748
14743
14743
14743
14743
14743
14743
14743
14749
14752
14752
14816
14656
14656
14656
14810
14810
14810
14810
13825
13945
13945
13822
13859
13859
13859
13859
13910
13845
13823
13750
13735
13825
13823
14200
14914
14906
14906
14914
14906
14906
15425
15425
15353
15340
15230
15115
15115
14353
14445
14435
LOCALITY
TETLIN JCT
TOK JUNCT
DONNELLY DO
TANANA R V
UPPER BERRY
DONNELLY DO
FT GREELY
FT GREELY
CURRY
CURRY
NIXON MINE
NIXON MINE
NIXON MINE
NIXON MINE
NIXON MINE
RUBY
COLLEGE
COLLEGE
ESTER
ESTER
SMITH L
COLLEGE
COLLEGE
COLLEGE
UNIV EXP FA
SHEEP CR
ESTER
ESTER
SMITH L
COLLEGE
COLLEGE
COLLEGE
UNIV EXP FA
SHEEP CR
COLLEGE
COLLEGE
COLLEGE
FAIRBANKS
FAIRBANKS
FAIRBANKS
FAIRBANKS
FAIRBANKS
FAIRBANKS
FAIRBANKS
COLLEGE
COLLEGE
COLLEGE
FAIRBANKS
HOT SPS RD
HOT SPS RD
HOT SPS RD
WICKERSHAM
WICKERSHAM
WICKERSHAM
WICKERSHAM
KLUANE LAKE
DONJEK R
DONJEK R
KLUANE LAKE
BURWASH LDG
BURWASH LDG
BURWASH LDG
BURWASH LDG
DUKE R
JACQUOT IS
KLUANE LAKE
ALSEK R
ALSEK R
ALSEK R
ALSEK R
CHITINA R
MATANUSKA
PALMER
PALMER
MATANUSKA
PALMER
PALMER
HEAD OF BIG
HEAD OF BIG
FAREWELL
FAREWELL LA
TONZONA R
KAHILTNA GL
KAHILTNA GL
TANANA R
TANANA R
TANANA RIVE
ME
CR
ME
RM
RM
M23
M23
M23
DOM
DOM
DOM
DOM
Re
Re
KE
R
DATE
28
08
30
10
03
25
02
02
17
17
COLLECTOR NAME
961
944
O57
944
957
967
964
964
939
939
949
949
949
949
949
932
957
957
O57.
957
956
957
957
B57
956
956
957
O57
956
9517
957
O57
956
956
957
957
949
933
933
933
933
931
931
931
967
967
967
967
967
965
965
957
957
957
e)Sii/
944
948
946
966
948
944
- 948
944
962
962
968
965
957
965
965
925
931
944
944
931
944
949
950
950
949
949
961
956
956
957
961
957
DOOOCrCTOKFTOTTFTTrTrFreKT aga OOODDODDAVOVVOVVODGVOOVOAOVGOVAZALZASAPYPrrrerexzmdcm
Cer re £2 22 CC CC a2) Ni eee oC Cc
Erpprpp-_exz<rr
eS OS a ees oe
=
c
spa => Pp > SC Cit
=
PepXVyEerrpresEeEeesETMVV|I ==
vuoud
Poeprrrvrrtrrte
LITTLE» JRe
RAuUP
ARGUS
RAUP
SPETZMAN
VIERECK
VIERECK
VIERECK
NELSON
NELSON
DRURY
DRURY
DRURY
DRURY
DRURY
SeMoHe ls
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
GUILSHER
PALMER
PALMER
PALMER
PALMER
PALMER
PALMER
PALMER
VIERECK
VIERECK
VIERECK
VIERECK
VIERECK
VIERECK
VIERECK
ARGUS
ARGUS
ARGUS
ARGUS
ANDERSON
ANDERSON
ANDERSON
MURRAY
RAUP
RAUP
RAUP
RAUP
SPETZMAN
SPETZMAN
WELSH
PEARSON
SCHOFIELN
VIERECK
VIERECK
LAING
ANDERSON
ANDERSON
ANDERSON
ANDERSON
ANDERSON
LEPAGE
DRURY
DRURY
DRURY
DRURY
VIERECK
VIERECK
VIERECK
SPETZMAN
SPETZMAN
SPETZMAN
cOL NO
1R235
12719
1061
12773
19?°0
R326
7152
7408
4173
4173
335¢e
to Boi Ag A
3274
3371
3364
44s
1159
LUST
LAT 7
1178
726
1170
1152
1160
426
477
1177
1178
7°6
1170
1152
1160
46
477
1159
LS7
3
192
164
veal
2en2
shal
11
12
R191
B1k4
RIAY
R2R7B
R31
75a1
7530
1198
1199
1198
1199
9493
19515
in3eal
365
13306
12271
13306
12271
113
12
7852
55
7568
7698
7698
BAY
8431
A431
RBG
R431
25242
3973
4N32
2A92
3036
5000
1054
1054
SeNe
4633
516
HERR + NO
GWA
ALA 19853
GWA
ALA 19837
ALA 6A42
GWA
ESEG
PSEC
ALA 255986
Isc 556
CAN
CAN
CAN
CAN
CAN
‘A 315821
ALA 6802
ALA 6817
ALA 6795
ALA 6A13
ALA 4570
ALA 6796
ALA 6792
ALA 6825
ALA 4096
ALA 4B4UYy
GWA
GWA
GWA
GWA
GWA
SWA
GWA
GWA
SWA
GWA
ALA 3103
ALA 5280
ALA 5274
ALA 5275
ALA 5993
ALA 25554
ALA 5969
ALA 5970
GWA
GWA
GWA
SWA
GWA
FSLC
FSLC
ALA 6935
ALA 6938
SWA
SWA
TSC 255968
ISC 255966
TSC 255965
BWA
ALA 19725
ALA 19724
GWA
GWA
CAN
CAN
GWA
CAN
CAN
GWA
FSLC 336
CAN
ALA 18348
ALA = 554
GWA
ISC 255967
TSC 255969
CAN
CAN
CAN
CAN
CAN
LAV
ALA 3498
SWA
ALA 69847
CAN
CAN
Specimens Cited
TaxON SPC S V HYB PROV QUAD LAT LONG LOCALITY DATE COLLECTOR NAME COL NO H&83 + NO
1873 113 14 86 6342 14435 TANANA RIVER 10 07 957 L A SPETZMAN 596 US 2349813
1873 113 14 87 6340 14930 TEKLANIKA R 28 07 956 G W ARGUS 694 ALA 4521
LS735 L103 14 87 6340 14930 TEKLANIKA R 28 07 956 G W ARGUS 6°94 GWA
1673 113 14 88 6330 15004 TOKLAT R 27 07 956 G W ARGUS 695 ALA 22493
1673 113 14 88 6330 15004 TOKLAT R 27 07 956 G W ARGUS 685 ALA 4520
1875 113 14 88 6330 15004 TOKLAT R 27 07 956 G W ARGUS 686 ALA 4516
1873 113 14 88 6330 15004 TOKLAT R 27 07 956 G ® ARGUS 625 GWA
1873 «113 14 86 6330 15004 TOKLAT R 27 07 956 G W ARGUS 686 GWA
1873 113 14 88 6325 15032 MULDROw GL 31 08 932 D KAYE 1 UWS 2441267
1873 «113 14 88 6326 15015 THOROFARE PASS 12 07 928 Y MEXIA 2095 ALA 10195
1873 113 14 88 ©6325 15032 MULDROW GL 12 07 932 WA SETCHELL 587 JS 2441268
1873 113 14 88 6325 15032 MULDROW GL 12 07 932 W f SETCHELL 527 wis
1873 113 14 88 6324 15020 THOROFARE R 04 07 956 L “A VIERECK 1098 ALA 116290
1873 113 14 88 6325 15035 MULDROW GL 25 06 958 L A VIERECK 3066 GWA
ta7s m3 14 88 6325 15030 THOROFARE R 02 08 964 L A VIERECK 7419 SLC 284
1873 113 14 108 6416 14519 TANANA R 29 05 965 L A VIERECK 7560 SWA
116 SITCHENSIS
1673 116 2 1 5935 13630 INSPECTOR CREE 12 7 967 G W ARGUS 6731 GWA
1873 116 2 2 5941 13632 CHILKAT PASSe 13 7 967 G W ARGUS 6815 GWA
1873 116 2 2 5941 13632 CHILKAT PASSe 13 7 967 G W ARGUS 6A15 GWA
1673 116 2 2 5935 13629 THREE GUARDSME 14 7 967 G W ARGUS 6200 GWA
1873 116 2 35 5530 12245 PINE RIVER» 7 6 967 G W ARGUS 6070 SWA
1873 116 2 35 5530 12245 PINE RIVER. 7 6 967 G W ARGUS 6967 GWA
1673 116 2 35 5530 12245 PINE QIVER- 7 6 967 G W ARGUS 6068 SWA
1873 116 2 49 5320 12245 PRINCE GEORGE 7 6 967 G ™ ARGUS 6977 GWA
1873 116 2 49 5330 12830 SKEENA RIVER+e 22 07 967 G W ARGUS 6R4B GWA
1873 116 14 3 5555 13002 HYDER 28 06 939 J © ANDERSON 5444 SC 255993
1873 116 14 3 5526 13140 KETCHIKANe 9 6 967 G ™ ARGUS 6090 GWA
1873 116 14 3 5559 13114 BURROUGHS BAY 19 09 915 E WALKER 1015 F 466614
1873 116 14 3 5555 13002 HYDER 02 06 924 K WHITED 1172B NA 332064
1673 116 14 6 5637 13250 PETERSSURG. 19 7 967 G W ARGUS 6232 GWA
1873 116 14 6 5648 13258 PETERSBURG. 19 4 967 G W ARGUS 6R34 GWA
1673 116 14 6 5637 13234 PETERSBURGe MI 18 7 967 G W ARGUS 6228 GWA
1873 116 14 6 5628 13223 WRANGELLe PATS 21 7 967 G w ARGUS 6242 GWA
1873 116 14 6 5628 13223 WRANGELL. 21 7 967 G wW ARGUS 6P45 GWA
1873 116 14 6 5628 13223 WRANGELL 19 09 950 J CHRIST 2n212 NA 320586
1873 116 14 6 5628 13223 WRANGELL 27 07 S14 A EASTWOOD 993 CAN
1873 116 14 9 5751 13459 FRESHWATER BAY 10 06 937 J © ANDERSON 30R9 TSC 255998
1873 116 14 9 5704 13516 SITKA ROAD TO 17 7 967 G W ARGUS 6P27 GWA
1873 116 14 9 5704 13516 SITKA E+eBRANC 17 7 967 G W ARGUS 6P26 GWA
1873 116 14 10 5840 13620 GEIKE INLET 29 06 928 J P ANDERSON 752 TSC 255995
1873 116 14 10 5854 13603 MUIR INLETe GL 29 6 967 G W ARGUS 6297 GWA
1873 116 14 10 5854 13603 MUIR INLETe GL 27 06 967 G Ww ARGUS 6256 GWA
1873 116 14 10 5854 13603 MUIR INLETe GL 27 06 967 G W ARGUS 6355 GWA
1873 116 14 10 5859 13606 MUIR INLETe GL 01 7 967 G W ARGUS 6598 GWA
1873 116 14 10 5859 13606 MUIR INLETe GL 01 7 967 G W ARGUS 6485 GWA
1873 116 14 10 5859 13606 MUIR INLETe GL 01 7 967 G W ARGUS 6481 GWA
1873 116 14 10 5858 13606 MUIR INLETe GL 29 6 967 G W ARGUS 6453 GWA
1873 116 14 10 5857 13602 MUIR INLETe GL 29 6 967 G W ARGUS 6424 GWA
1873 116 14 10 5855 13603 MUIR INLET? GL 28 6 967 G W ARGUS A3R3A GWA
1873 116 14 10 5855 13603 MUIR INLETr GL 28 6 967 G W ARGUS 6374 GWA
1873 116 14 10 5855 13603 MUIR INLETe GL 28 6 967 G W ARGUS 6364 GWA
1873 116 14 10 5832 13724 CRILLON LAKE 13 07 956 D- BRINK 10 F 885198
1873 116 14 11 5825 13433 MENDENHALL GL 24 06 917 J © ANDERSON 411 TSC 255983
1873 116 14 11 5825 13433 MENDENHALL GL 31 05 925 J © ANDERSON 2a15 TSC 255997
1873 116 14 11 5825 13433 MENDENHALL GL 16 05 941 J © ANDERSON 6439 ISC 255990
14673 116 14 11 5818 13425 JUNEAU 23 04 946 J P ANDERSON 6227 ALA 555
1873 116 14 11 5820 13436 DOUGLAS ISLAND 07 7 967 G W ARGUS 6595 SWA
1873 116 14 11 5825 13540 GUSTAVUS. 04 07 967 G WwW ARGUS 6594 GWA
1873 116 14 11 5822 13436 MENDENHALL RIV 13 6 967 G W ARGUS 6105 GWA
1873 116 14 11 5820 13436 DOUGLAS ISLAND 07 7 967 G W ARGUS 6597 GWA
1473 116 14 11 5826 13435 MENDENHALL GLA 10 7 967 G W ARGUS 6665 SWA
1673 116 14 11 5831 13445 EAGLE RIVERe 110 7 967 G WwW ARGUS 6667 SWA
18673 116 14 11 5825 13540 GUSTAVUS. 04 07 967 G WwW ARGUS 6573 SWA
£875 216 14 11 5818 13423 MT ROBERTS TR 09 7 967 G W ARGUS 6642A GWA
1873 116 14 11 5818 13423 MT ROZERTS TR 09 7 967 G Ww ARGUS 6651 GWA
1873 116 14 11 5823 13440 JUNEAU 13. 6 967 G W ARGUS 6099 6WA
1873 116 14 11 5817 13424 MT ROBERTS 06 08 967 L * VIERECK A606 GWA
1873 116 14 34 5715 15332 THREE SAINTS B 07 07 963 B 4 NYBAKKEN 1008 4SwWA
1873 116 14 45 5926 13612 BIG BOULDER CR 12 7 967 G W ARGUS 6492 GWA
1873 116 14 45 5926 13612 BIG BOULDER CR 12 7 967 G W ARGUS 6695 GWA
1873 116 14 45 5925 13603 KLEHINI RIVER 12 7°967 G Ww ARGUS 6688 GWA
1873 116 14 45 5926 13612 BIG BOULDER CR 12 7 967 G Ww ARGUS 6694 GWA
1873 116 14 45 5907 13520 HAINESe MIe 6 13 7 967 G W ARGUS 6746 GWA
1873 116 14 45 5928 13605 MOSQUITO LAKE 45 7 967 G W ARGUS 6679 SWA
1873 116 14 45 5928 13602 MOSQUITO LAKEe 11 7 967 G Ww ARGUS 6671 GWA
1873 116 14 45 5928 13602 MOSQUITO LAKEe 11 7 967 G W ARGUS 6672 GWA
1873 116 14 45 5901 13533 DAVIDSON GL 06 07 929 W COOPER 111 F 74592A
1873 116 14 45 5928 13519 SKAGWAY 26 08 916 M MALTE CAN 122114
1873 116 14 45 5928 13519 SKAGWAY 25 08 916 M- MALTE CAN 122111
1873 116 14 46 5952 13945 YAKUTAT BAY. 16 6 967 G W ARGUS 6190 GWA
1673 116 14 46 5952 13945 YAKUTAT BAY. 16 6 967 G w ARGUS 6191 GWA
1873 116 14 46 5955 13945 YAKUTAT BAY. 15 6 967 G Ww ARGUS 6180 GWA
1873 116 14 46 5952 13945 YAKUTAT BAY. 16 6 967 G W ARGUS 6192 GWA
1673 116 14 46 5952 13945 YAKUTAT BAY. 14% 6 967 G W ARGUS 6117 GWA
1873 116 14 46 5952 13945 YAKUTAT BAY. 16 6 967 G W ARGUS 6186 GWA
1873 116 14 46 5952 13945 YAKUTAT BAY. 14 6 967 G W ARGUS 6136 GWA
ie73° 116 14 46 5952 13945 YAKUTAT BAY. 16 6 967 G W ARGUS 6186 GWA
261
Appendix B
TAXON SPC S V HYB PROV QUAD LAT LONG LOCALITY DATE
1873 116 14 46 5915 13830 TANIS LAKE LI
1873 116 14 46 5915 13830 TANIS LAKE 21.6
1675 116 14 46 5915 13830 TANIS LAKE LOG
1873 116 14 46 5932 13940 YAKUTAT. 23 6
1873 116 14 46 5915 13830 TANIS LAKE- 226
18673 116 14 46 5933 13930 SITUK RIVER. 23 6
1473 116 14 46 5935 13940 YAKUTAT 10 06
1873 116 14 46 5935 13940 YAKUTAT 12 06
1873 «116 14 46 5935 13940 YAKUTAT 10 06
18735 116 14 46 5935 13940 YAKUTAT 10 06
1873 +116 14 46 5930 13940 YAKUTAT 09 06
1873 «116 14 46 5906 13822 E ALSEK R 10 06
1875 116 14 46 5930 13940 YAKUTAT 09 06
1873 116 14 46 5930 13940 YAKUTAT 09 06
1873 «116 14 46 5908 13822 E ALSEK R 10 06
1873 116 14 46 5930 13940 YAKUTAT 09 06
1873 116 14 50 5940 15132 HOMER 13 06
1673 116 14 62 6003 15140 NINILCHIK CAMP 13 06
1873 116 14 62 6002 15141 DEEP CREEK 13 06
1673 116 14 63 6055 14939 HOPE 03 06
1873 116 14 63 6007 14927 SEWARD 30 05
1873 116 14 63 6007 14927 SEWARD 30 05
1873 116 14 63 6055 14939 HOPE 24 05
1873 116 14 63 ©6019 14921 SNOW R DELTA 28 05
1875 116 14 63 6003 14804 PORT SAN JUAN 12 07
1873 116 14 63 6007 14927 SEWARD 27 05
1875 116 14 63 6003 14804 PORT SAN JUAN 12 07
1875 116 14 63 6048 14852 PORTAGE 24 07
1873 116 14 63 6055 14900 PORTAGE GL 07 06
1873 116 14 63 60035 14904 DAY HARBOR 16 06
1873 116 14 63 6003 14904 DAY HARBOR 16 06
1873 116 14 63 6048 14855 PORTAGE GL RD 11 06
1873 116 14 63 6048 14855 PORTAGE GL RD 11 06
1873 116 14 63 6048 14850 PORTAGE GL 04% 07
1873 116 14 63 6027 14820 NELLIE JUAN GL 17 08
1873 116 14 63 6045 14923 SEWARD HWY M59 12 06
1873 «116 14 63 6048 14850 PORTAGE GL 04 07
1873 «116 14 64 6033 14545 CORDOVA 28 07
1873 116 14 64 6040 14543 ORCA 0s
1873 116 14 64 6100 14500 COPPER R FLATS
1873 «116 14 64% 6100 14500 COPPER R FLATS
1873 «116 14 68 6107 14616 VALDEZ 05 07
1873 116 14 68 6111 14619 VALDEZ GLACIER 02 08
1873 116 14 68 6112 14619 VALDEZ 02 08
16735 116 14 668 6112 14619 VALDEZ 02 08
116 STOLONIFERA
1673 118 02 002 5935 13630 INSPECTOR CREE 12 7
1873 118 02 002 5935 13630 INSPECTOR CREE 12 7
1873 «118 02 002 5935 13630 INSPECTOR CREE 12 7
1873 118 02 G02 5935 13630 INSPECTOR CREE 12 7
1673 118 02 002 5935 13629 THREE GUARDSME 14 7
1673 118 02 002 5935 13629 MT GLAVE ee
1873 1186 02 v02 5935 13629 MT GLAVE 14 #7
1873 «118 02 002 5935 13629 MT GLAVE 14° #7
1673 118 02 002 5935 13629 MT GLAVE LOO iT,
1673 118 02 002 5935 13629 MT GLAVE 14° #7
1673 118 02 002 5935 13629 MT GLAVE i4 7
1873 118 02 002 5935 13629 MT GLAVE 1G 7
1673 118 02 002 5935 13629 MT GLAVE 14° #7
1873 116 02 002 5935 13629 THREE GUARDSME 14 7
1673 «118 02 v02 5935 13629 THREE GUARDSME 14 7
18735 118 02 002 5935 13629 THREE GUARDSME 14 7
1873 1186 02 v02 5935 13630 INSPECTOR CREE 12 7
1673 118 02 002 5935 13630 INSPECTOR CREE 12 7
1873 118 02 002 5935 13630 INSPECTOR CREE 12 7
1673 118 02 002 5935 13630 INSPECTOR CREE 12 7
1873 118 02 002 5935 13630 INSPECTOR CREE 12 7
1873 118 02 G02 5935 13630 INSPECTOR CREE 12 7
18735 118 02 002 5935 13630 INSPECTOR CREE 12 7
1673 118 02 002 5935 13630 INSPECTOR CREE 12 7
1873 118 02 002 5935 13630 INSPECTOR CREE 12 7
1673 118 02 U02 5935 13630 INSPECTOR CREE 12 7
1873 116 02 002 5935 13630 INSPECTOR CREE 12 7
1673 118 02 02 5935 13629 THREE GUARDSME 14% 7
1675 118 02 002 5935 13629 MT GLAVE 14 7
1873 116 02 v02 5935 13629 THREE GUARDSME 14 7
1873 118 02 2 5938 13628 HAINES RD M81 13 07
1673 118 02 2 5935 13627 HAINES RD M58 O04 08
16735 116 02 2 5935 13627 HAINES RD M58 O04 08
1673 116 02 3 5938 13508 WHITE PASS 23 07
1873 118 02 3 9938 13508 WHITE PASS 23 07
1675 118 02 3 5938 13508 WHITE PASS 23 07
1673 118 02 3 5938 13508 WHITE PASS 23 07
1673 118 02 3 5938 13508 WHITE PASS 23 07
1673 118 02 3 5938 13508 WHITE PASS 23 07
1873 118 02 3 5938 13508 WHITE PASS 10 09
18673 1186 02 3 9936 13508 WHITE PASS 10 09
1873 118 02 13. 5830 13020 DEASE LAKE 03 08
262
967
967
967
967
967
967
966
966
966
965
965
965
965
965
965
967
967
967
941
941
941
951
951
948
945
948
967
966
952
952
967
967
957
957
967
957
940
940
960
960
935
967
967
967
967
967
967
967
967
967
967
967
967
967
967
967
967
967
967
967
967
967
967
967
967
967
967
967
967
967
967
967
967
967
956
967
967
914
914
914
914
914
914
900
900
962
COLLECTOR NAME
rCroeeprwerrrrrrvu0<<ZZEcccececrrrrrrrrr e<<<caanaaoo
YVNMNPSFP FPP EFTFTAVDNDAANVNVDNDNADNDAGNVAMNNAVNYVVNGVNAANNVANNNNGDNDNANGNDNDNODD
Pretutcepesvd~v~vP SPs > rere KePrrrreteteeeE
rCePrparprrvyr:>
Pv PTdU
PRP EZLTECEECLCEETTZTEETSEALETEEEEELCTTEEZEEEEET
ARGUS 6240
ARGUS 6274
ARGUS 6?10
ARGUS 6343
ARGUS 6306
ARGUS 6353
HARMS 5172
HARMS 5216
HARMS 5165
HARMS 5205
VIERECK 7597
VIERECK 761
VIERECK 7595
VIERECK 7597
VIERECK 76Nn1
VIERECK 7595
VIERECK 8269
VIERECK 8260
VIERECK 8263
ANDERSON 6539
ANDERSON 6462
ANDERSON 6462
CALDER 4926
CALDER 4965
EYERDAM 5845
EYERDAM 3672
EYERDAM 5845
HARMS 5896A
HARMS 5086
KLEIN 1
KLEIN 2
VIERECK R230
VIERECK 8229
VIERECK 2033
VIERECK 2240
VIERECK A2u3
VIERECK 2033
EYERDAM
NORBERG
SHEETS SeNe
SHEETS SeNe
ANDERSON 1845
VIERECK A749
VIERECK R748
VIERECK A743
ARGUS 6697
ARGUS 6726
ARGUS 6706
ARGUS 6707
ARGUS 6776
ARGUS 6797
ARGUS 6794
ARGUS 6796
ARGUS 6789
ARGUS 6788
ARGUS 6784
ARGUS 6782
ARGUS 67B1
ARGUS 6773
ARGUS 6772
ARGUS 6777
ARGUS 6710
ARGUS 6711
ARGUS 6716
ARGUS 6722
ARGUS 6725
ARGUS 6734
ARGUS 6733
ARGUS 6697
ARGUS 6726
ARGUS 6706
ARGUS 6707
ARGUS 6776
ARGUS 6799
ARGUS 6775
TAYLOR 1351
VIERECK A539
VIERECK B539
EASTWOOD 927
EASTWOOD 927
EASTWOOD 9°7
EASTWOOD 927
EASTWOOD 9°7
EASTWOOD 927
WALPOLE 1249
WALPOLE 1249
MACDONALD 4us
COL NO HE8S + NO
GWA
GWA
GWA
GWA
GWA
GWA
ALA
ALA
ALA
ALA
FSLC
FSLC
FSLC
GWA
GWA
GWA
GWA
GWA
GWA
sc
Woe
ALA
GWA
GWA
CAN
CAN
msc
ALA
ALA
ALA
ALA
GWA
GWA
GWA
GWA
SWA
ALA
NA
CAN
ALA
ALA
YSE
SWA
GWA
SWA
32574
32571
32576
32572
334
327
332
255991
255992
558
255985
34637
32643
22977
22975
8388
332115
25125
25121
256600
SWA
GWA
SWA
SWA
SWA
SWA
GWA
SWA
GWA
GWA
GWA
SWA
GWA
GWA
GWA
SWA
SWA
SWA
SWA
SWA
SWA
SWA
SWA
GWA
GWA
SWA
GWA
SWA
GWA
GWA
AO
GWA
GWA
CAN
CAN
32737
46435
46435
A
US 1011915
A
11S 1911915
US 378231
NS 378231
CAN 273999
Specimens Cited
TAXON SPC S V HYB PROV QUAD LAT LONG LOCALITY DATE
1873 118 02 13 5830 13020 DEASE LAKE 03 08
1673 118 02 20 5800 13110 TELEGRAPH CR 15 08
1873 118 02 20 5800 13110 TELEGRAPH CR 15 08
1873 118 02 33 5530 12748 SKEENA CROSS 30 08
1873 118 02 33 5530 12748 SKEENA CROSS 30 08
1873 118 14 3 5555 13001 CHICKAMIN GL 17 07
1873 118 14 4 5553 13414 PIN PEAK 09 06
1673 116 14 10 5854 13603 MUIR INLET? GL 29 6
1873 118 14 10 5859 13606 MUIR INLETe GL O01 7
1873 118 14 10 5858 13606 MUIR INLET? GL O01 7
1873 118 14 10 5858 13606 MUIR INLET*e GL 01 7
1873 118 14 10 5858 13606 MUIR INLETe GL 01 7
1873 118 14 10 5858 13606 MUIR INLETe GL 01 7
1673 118 14 10 5858 13606 MUIR INLETe. GL 29 6
1873 118 14 10 5858 13606 MUIR INLETe GL 29 6
1873 118 14 10 5858 13606 MUIR INLETe GL O1 7
1873 118 14 10 5858 13606 MUIR INLET? GL 29 6
1873 118 14 10 5858 13606 MUIR INLET? GL O01 7
1873 118 14 10 5858 13606 MUIR INLETe GL 01 7
1873 118 14 10 5858 13606 MUIR INLETe GL 01 7
1873 118 144 10 5858 13606 MUIR INLET? GL 01 7
1873 118 14 10 5858 13606 MUIR INLET? GL 29 6
1873 118 14 10 5858 13606 MUIR INLETe GL 29 6
1873 118 14 10 5858 13606 MUIR INLETe GL O01 7
1873 118 14 10 5859 13606 MUIR INLETe GL 01 7
1673 118 14 10 5859 13606 MUIR INLETe GL O01 7
1873 118 14 10 5859 13606 MUIR INLET? GL 01 7
1873 118 14 10 5859 13606 MUIR INLET? GL 01 7
1673 118 144 10 5859 13606 MUIR INLET? GL 01 7
1873 118 14 10 5859 13606 MUIR INLETe GL 01 7
1873 118 14 10 5859 13606 MUIR INLET? GL O01 7
1673 118 14 10 5859 13606 MUIR INLET* GL 01 7
1873 118 14 10 5859 13606 MUIR INLET? GL O01 7
1873 118 14 10 5859 13606 MUIR INLETe GL 01 7
1873 118 14 10 5859 13606 MUIR INLETe GL 01 7
1873 118 14 10 5855 13603 MUIR INLET 28 06
1873 118 144 10 5855 13603 MUIR INLETe GL 28 6
1873 118 14 10 5840 13710 MOUNT CRILLON 14 07
1873 118 144 10 5858 13611 MUIR INLET 08 06
1873 118 14 10 5858 13611 MUIR INLET 08 06
1873 118 144 10 5900 13610 MUIR GLACIER 29 06
1873 118 14 10 5900 13610 MUIR GLACIER 09 06
1873 118 144 11 5818 13425 JUNEAU 15 06
1873 118 14 11 5818 13425 JUNEAU 15 06
1873 118 14 11 5817 13424 MT ROBERTS 17 06
1673 118 144 11 5826 13434 MENDENHALL oe 7
1873 118 144 11 5830 13420 AMHERST PK 26 07
1873 118 14 11 5826 13435 MENDENHALL GLA 10 7
1873 118 14 11 5826 13435 MENDENHALL GLA 10 7
1873 118 14 11 5818 13423 MT ROBERTS TR 09 7
1673 118 14% 11 5818 13423 MT ROBERTS TR 09 7
1873 118 14 11 5840 13410 TAKU B 10 08
1873 118 1% 11 5840 13415 TAKU C 21 07
1873 118 14 11 5830 13415 DEATH VALLEY 27 07
1673 118 144 11 5818 13423 MT ROBERTS TR 09 7
1873 118 14 11 5818 13423 MT ROBERTS TR 09 7
1873 118 14 11 5818 13423 MT ROBERTS TR 09 7
1873 118 14 11 5818 13423 MT ROBERTS TR 09 7
1873 118 14 11 5818 13423 MT ROBERTS TR 09 7
1873 118 14 11 5818 13423 MT ROBERTS TR 09 7
1873 118 14 11 5818 13423 MT ROBERTS TR 09 7
1873 118 14 11 5818 13423 MT ROBERTS TR 09 7
1673 118 14 11 5818 13423 MT ROBERTS TR 09 7
1873 118 14 11 5818 13423 MT ROBERTS TR 09 7
1873 118 14 11 5818 13423 MT ROBERTS TR 09 7
1873 118 14 11 5818 13423 MT ROBERTS TR 09 7
1873 118 14 11 5818 13423 MT ROBERTS TR 09 7
1873 118 144 11 5818 13423 MT ROBERTS TR 09 7
1873 118 14 11 5826 13435 MENDENHALL GLA 10 7
1873 118 14 11 5817 13424 MT ROBERTS 23 07
1673 118 14 11 5817 13424 MT RORERTS 06 08
1873 118 14 11 5817 13424 MT ROBERTS 06 08
1873 118 14 29 5503 16219 KING COVE 26 07
1673 118 14% 29 5503 16219 KING COVE 26 07
1673 116 144 32 5651 15418 CAPE ALITAK 23 06
1673 118 14 34 5747 15247 KODIAK 08 06
1873 118 14 34 5747 15247 KODIAK 28 07
1673 118 14 34 5747 15247 KODIAK 20 08
1673 118 14 34 5747 15247 KODIAK 20 08
1873 116 14 34 5747 15247 KODIAK 20 08
1873 118 14 34 5747 15247 KODIAK 20 08
1873 118 14 34 5747 15247 KODIAK 22 08
1873 118 14 34 5747 15247 KODIAK 22 08
1673 118 14% 35 5710 15418 CANNERY MT 28 06
1673 118 144 35 5710 15418 CANNERY MT 10 06
1873 118 14% 35 5710 15418 CANNERY MT 10 06
1873 118 14 39 5834 16146 CAPE PEIRCE 7 6
1673 116 14 39 5834 16146 CAPE PEIRCE 26 5
1873 118 14 39 5834 16146 CAPE PEIRCE 28 5
1873 118 14 39 5834 16146 CAPE PEIRCE 30 5
1673 118 14 39 5834 16146 CAPE PEIRCE mF
COLLECTOR NAME
COOK KMMMAUAUNAAUNROANADEEFTPADDADAAVDAVUNANDANDAANMNVNANDAAANAVAVNADCCCCTETEAURWAVNAONAD VDDVDANADNDODANANDADADAVDVMAAAVNAOAOADADAOANDOIACCAAYW
> PDP
SEEFCTELEETETTEAESLTTEEECSECLTEZTEEEAETEEE
<<
PRPPr>> PCI NCCP "EEL TZTTELSEAZELETLEECTZEELETLZEACTzAZTVVII
MACDONALD
MCCABE
MCCABE
CALDER
CALDER
MCCABE
KLEIN
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
BATES
COVILLE
COVILLE
EVANS
TRELEASE
ANDERSON
ANDERSON
ANDERSON
ANDERSON
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
THRONE
VIERECK
VIERECK
EYERDAM
EYERDAM
GRIGGS
COVILLE
PIPER
WALPOLE
WALPOLE
WALPOLE
WALPOLE
WALPOLE
WALPOLE
LOOFF
LOOFF
LOOFF
DICK
DICK
DICK
DICK
DICK
COL NO
445
RBAT
8887
15219
15219
9006
460
6395
6477
6474
6473
6471
6470
6461
6460
6469
6448
6469
6469
6470
6471
6452
6448
6473
6511
6517
6511
6519
6524
6526
6527
6528
HERB
CAN
NT
NA
US
ys
NA
ALA
GWA
GWA
GWA
GWA
GWA
GWA
BWA
GWA
BWA
aWA
GWA
5WA
BWA
GWA
CWA
GWA
AWA
GWA
6WA
GWA
GWA
GWA
OWA
GWA
GWA
BWA
BWA
GWA
GWA
SWA
GH
us
US
US
A
ALA
ISC
Isc
Isc
ALA
GWA
BWA
GWA
GWA
GWA
GWA
5WA
BWA
GWA
BWA
GWA
BWA
BWA
6WA
BWA
GWA
GWA
AWA
GWA
GWA
BWA
SWA
ws
BWA
GWA
Us
WS
US
US
US
US
US
US
US
Us
US
A
A
NY
GWA
SWA
5WA
5WA
GWA
+ NO
273999
319578
319578
2349127
2349127
319577
17864
373565
376921
378396
559
256239
256224
85589
4736
1631461
1631461
1072095
373564
421028
378193
378194
378194
378193
378199
378200
263
Appendix B
264
SPC S V HYB PROV QUAD LAT
118 14
118 14
118 14
118 14
118 14
118 14
118 14
118 14
118 14
118 14
118 14
118 14
118 14
118 14
118 14
118 14
118 14
116 14
118 14
118 14
118 14
118 14
118 14
DRUMMOND IANA
119 1
119 1
119 i
119 1
119 1
119 1
119 1
119 2
119 2
119 2
119 2
119 2
119 2
119 2
119 2
119 2
119 2
119 2
119 11
119 11
119 11
119 12
119 12
119 12
119 12
1 ALAXENSIS VAR.
122 1 2
122 il 2
122 1 2
122 1 2
i2e 02 2
122 01 2
122 01 2
122 01 7
i220 1 7
122°'0 1 7
122 01 7
t2e 0 4 7
i220 i 7
122 0 1 7
i220 1 7
122 01 Us
tee 0 2 7
uzZe 0 *5 7
122 01 7
122 1 7
122 1 12
122 1 12
122 1 12
122 1 12
122 1 12
ize 0 4 12
i122 01 12
t2e, 0 i 12
ice 0 1 12
122 01 12
i122 01 14
122 is 14
122 1 14
122 1 14
122 1 14
ALAXENSIS
5802
5927
5915
5915
5915
5915
5915
5915
5915
§915
5933
5933
5933
5933
5952
6025
6030
6030
6001
6146
6146
6104
6104
5644
5644
5644
5644
5443
5241
4901
5822
5835
5748
5847
5714
5705
5705
5705
5705
5714
5550
5901
5901
5901
6005
6003
6003
6003
5941
5941
5941
5935
5839
5831
5821
6842
6813
6750
6559
60456
6416
6409
6336
6525
6237
6242
6242
6127
6922
6807
6850
6840
6734
6103
6122
6145
6016
6003
5849
5855
5855
5859
5859
LONG
15617
13519
13830
13830
13830
13830
13830
13830
13830
13830
13945
13945
13944
13944
15631
15020
14926
14926
13930
14918
14918
14905
14905
11123
11125
11125
11123
11317
11802
11404
12955
12240
12253
12335
12241
12235
12235
12235
12235
12243
12055
10900
10900
10900
13035
12840
12840
12855
13632
13632
136352
13629
12428
12434
12434
13406
13547
11550
10152
125435
9605
8318
8405
8550
11130
10908
10908
12542
13952
13928
13845
13907
13832
13822
13859
13307
13255
12840
13620
139035
139035
13606
13606
LOCALITY
BECHAROF L
SKAGWAY
TANIS LAKEe
TANIS LAKEe
TANIS LAKEe
TANIS LAKEe
TANIS LAKEe
TANIS LAKEe
TANIS LAKEe
TANIS LAKEe
YAKUTAT BAY
YAKUTAT BAY
YAKUTAT
YAKUTAT
STUYAHOK L
SKILAK L
MOOSE PASS
MOOSE PASS
HUBBARD GL
HATCHER PASS
HATCHER PASS
GIRDWOOD MINE
GIRDWOOD MINE
MCMURRAY
MCMURRAY
MCMURRAY
MCMURRAY
ATHABASCA LDG
MT EDITH CAVE
CAMERON LAKE
DEASE L VILLAG
FORT NELSON
FT NELSON
STEAMBOAT CR
SIKANNI CHF R
BEATON RIVER
BEATON RIVER
BEATON RIVER
BEATON RIVER
SIKANNI CHF R
KISKATINAW R
LITTLE GULL L
EITMEE GULE L
LITTLE GULL L
RANCHERIA
WATSON LAKE
WATSON LAKE
UPPER LIARD R
NADAHINI RIVER
NADAHINI RIVER
NADAHINI RIVER
THREE GUARDSME
N TETSA R
SUMMIT PASS
SUMMIT PASS
MACKENZIE DELT
CANOE LAKE
COPPERMINE
PELLY LAKE
NORMAN
THELON RIVER
CORAL HARBOR
SOUTHAMPTON IS
SOUTHAMPTON IS
TALTHEILEI NRS
FORT RELIANCE
FORT RELIANCE
FLAT RIVER
FIRTH RIVER
BLACK FOX CR
BABBAGE Re
BABBAGE CREEK
OLD CROW Re
KLUANE LAKE
KLUANE LAKE
ROSE LAPIE R P
TESLIN LAKE
WATSON LAKE
GLACIER BAY
MUIR INLET»?
MUIR INLET?
MUIR INLET?
MUIR INLET?
GL
GL
GL
GL
08
07
07
07
NN OO
949
934
967
967
967
967
967
967
967
967
899
899
916
916
951
939
941
941
899
947
947
S57
957
939
939
939
939
925
968
956
969
960
960
960
960
943
943
943
943
943
966
962
962
962
966
966
966
966
967
967
967
967
960
943
943
934
964
963
966
947
965
948
951
952
927
964
963
970
970
970
970
970
970
967
944
944
96B
966
928
967
967
967
967
COLLECTOR NAME
RAuP
ANDERSON
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
COVILLE
COVILLE
WALKER
WALKER
MULLER
SHARPLES
ANDERSON
ANDERSON
COVILLE
DUTILLY
DUTILLY
A VIERECK
VIERECK
TOCCEMMMUNNDVAGVDNDDAMAGDDCA
S{VUVUVELTVIES ZEZAZAZEZEAZZEVS
EG
>
RAUP
RAUP
RAuP
RAuUP
BALL
ARGUS
HERMANN
RIGBY
CALDER
CALDER
CALDER
CALDER
RAUP
RAUP
RAUP
RAuUP
RAUP
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
e€eese
=
MoOooQOOOOOT TILTIceceanzanoortriidsg
SEZELLTTETSC secs ts S£Pevrvnp
ARGUS
ARGUS
ARGUS
ARGUS
CALDER
RAUP
RAUP
PORSILD
LARSEN
LARSEN
LARSEN
PORSILD
KUYT
CODY
BROWN
IRVINE
RAuUP
LARSEN
LARSEN
SCOTTER
WELSH
RIGBY
WELSH
WELSH
RIGBY
MURRAY
RAUP
PORSILD
WELSH
ARGUS
ANDERSON
ARGUS
ARGUS
ARGUS
ARGUS
=<
P=et
=
=
NerPrMNne
MOOOCOYHMPIOCHNNCYVNOCCTIMWOVEZEMPCCCHPITIICOHDNDOD®D
ZEZLEVEC VEVALC AL £2 ees VAG
COL NO HERB + NO
96
16398
6299
6293
6336
6332
6334
6304
6202
6299
1141
1141
1061
1061
BRE
SeNe
6814
6814
1079
220R5
22006
2045
2045
ISC 224498
TSC 256241
GWA
GWA
GWA
GWA
GWA
GWA
GWA
GWA
US” 373567
A
US 887058
US 1086935
US 2176395
TSC 256236
TSC 256419
NA 319576
US 373566
DAO 32722
QAO 32742
ALA 8375
GWA
US 2347455
US 2347453
US 2347454
GWA
US 1287566
SWA
US 223340?
OTF 1143
GWA
GWA
GWA
SWA
US 24226156
SWA
GWA
SWA
SWA
GWA
GWA
SWA
GWA
GWA
SWA
GWA
GWA
GWA
GWA
SWA
SWA
ALA
ALA
ALA
SWA
GWA
SWA
GWA
GWA
GWA
msc
GWA
GWA
ALA
GWA
GWA
GWA
NTF
OTE
RARY
OTF
BRY
GWA
ALA
Ise
GWA
GWA
TS¢
GWA
GWA
SWA
SWA
17326
19740
19747
255845
14599
1200
1198
91150
1199
92303
19741
555896
255905
Specimens
TaxON SPC S
1673
1873
1673
1873
1873
1873
1873
1873
1873
1873
1473
1873
1873
1873
1873
1873
1873
1873
1873
1873
1873
1873
18673
1873
1473
1873
1873
1873
1873
1873
1873
1873
1873
1873
1873
1873
18673
1873
1873
1873
1873
1873
1873
1673
1873
1873
1873
1873
1873
1873
1873
1673
1873
1873
1873
1873
1873
1873
1873
1873
1873
1873
1873
1873
1873
1873
1873
1873
1873
1873
1873
1873
1673
1873
1873
1873
1873
1873
1873
1873
14873
1873
1873
1873
1673
1673
1673
1873
1873
122
122
122
122
122
122
122
122
lee
122 0
122 0
122
122
122
122
122
122
122
122
122
122
122
122
122
122
122
122
122
122
122
122
122
122
122
122
122
122
122
122
122
122
122
122
122
122
122
122
122
122
122
122
122
122
122
122
122
122
122
122
122
122
122
122
122
122
122
122
122
122
122
122
122
122
122
122
122
122
122
122
122
122
122
122
122
122
122
122
122
122
ooooco
ooooocoococoooecoeocoecoeceoocoeoceoececoooooceo
ooooococoooooococoeococooccocoeoocoocoeoo°e°ooe
Cited
V HYB PROV QUAD LAT
a ee
10
10
10
10
5854
5855
5859
5855
5451
5748
5805
5949
5952
5904
6000
6013
6003
6140
6140
6140
6136
6155
6152
6233
6231
6234
6234
6240
6310
6310
6337
6337
6337
6337
6337
6337
6355
6330
6330
6324
6331
6330
6332
6302
6430
6452
6549
6557
6536
6725
6757
6853
6806
6815
6800
6840
6839
6855
6803
6822
6822
6822
6822
6822
6839
0924
6924
6924
6925
6922
6922
6923
6923
6923
6923
6923
6922
6922
6953
6953
6956
6902
6902
6902
6902
7030
7030
7030
7030
7030
7030
7040
7010
LONG
13605
13903
13606
13903
16324
15404
15305
13638
13945
15455
16700
15048
15140
14215
14900
14900
14906
15425
154335
15337
15336
15049
15049
15230
14530
14530
14643
14643
14643
146435
14643
14643
14725
15000
15000
15020
15002
15000
15002
16320
16530
14750
14403
14526
16315
15007
15312
166135
16545
16600
16533
16610
166135
16110
14500
14355
14355
14355
14355
14355
14100
14608
14608
14608
15210
15208
15208
15210
15210
15210
15210
15210
15210
15210
15708
15708
15957
16350
16350
16350
16350
15730
15730
15730
15730
15730
15730
15655
14650
LOCALITY
GL
GL
MUIR INLET?
MUIR INLET?
MUIR INLET
MUIR INLET
FALSE PASS
UYAK BAY
RASPBERRY ISe
HAINES RD M 82
YAKUTAT BAYe
BATTLE L
DUCHILETHLUK R
BEAR CREEK
NINILCHIK
WRANGELL MTS
MATANUSKA VY
MATANUSKA VY
LTL SUSITNA CN
HEAD OF BIG Re
HEAD OF BIG Re
FAREWELL Le
FAREWELL
TALKEETNA MTS
TALKEETNA MTS
TONZONA RIVER
PHELAN CREEK
PHELAN CREEK
GLACIER CREEK
W FK LTL DELTA
W FK LTL DELTA
W FK LTL DELTA
W FK LTL DELTA
W FK LTL DELTA
DRY CREEK
TOKLAT R BR
TOKLAT R BR
THOROFARE R
TOKLAT R BR
TOKLAT RIVER
TOKLAT R BR
PASTOLIK RIVER
NOME
COLLEGE
CIRCLE
EAGLE CR CAMP
IMURUK LAKE
WISEMAN
EASTER CREEK
CAPE LISBURNE
OGOTORUK CREEK
CAPE THOMPSON
KUKPUK RIVER
KAPALOAK CREEK
KAPALOAK CREEK
DRIFTWOOD CR
OLD JOHN LAKE
SHEENJEK RIVER
SHEENJEK RIVER
SHEENJEK RIVER
SHEENJEK RIVER
SHEENJEK RIVER
FIRTH RIVER
CANNING RIVER
CANNING RIVER
CANNING RIVER
UMIAT
UMIAT
UMIAT
UMIAT
UMIAT
UMIAT
UMIAT
UMIAT
UMIAT
UMIAT
MEADE RIVER
MEADE RIVER
WAINWRIGHT
CAPE BEAUFORT
CAPE BEAUFORT
CAPE BEAUFORT
CAPE BEAUFORT
MEADE Re PeOe
MEADE Re PeOeo
MEADE Re PeOec
MEADE Re PeDOec
MEADE Re PeOe
MEADE Re PeDOde
MEADE RIVER
BULLEN
DATE
06
6
07
06
05
05S
08
08
6
07
08
8
06
06
06
COLLECTOR NAME
MADD ANDMNGMNANGANGDAODNOOVDA<<<<<YNONTFFTFTFMBWWAOWNArrIrac<c<ccc MOFrFWTrrroaOarrrrrrrgogogrececmzaezeeEenrramMmunoc<carezecrHQoo
rf Lr ze =e =
GrpervrvEszPePgcacaGcEeacte ev Vuriststrecseer
=
G
ZLTEZZELETELTEZE
ARGUS
ARGUS
ARGUS
ARGUS
NORBERG
ANDERSON
EYERDAM
VIERECK
ARGUS
HARMS
BOS
HAMMARSTROM
LITTLE» JR.
SCOTT
PALMER
PALMER
WELSH
DRURY
DRURY
DRURY
LITTLE»
ANDERSON
ANDERSON
VIERECK
ARGUS
ARGUS
PALMER
PALMER
PALMER
PALMER
PALMER
PALMER
VIERECK
ARGUS
ARGUS
VIERECK
VIERECK
VIERECK
WELSH
PALMER
HELLER
ARGUS
BEAUCHAMP
TRENT
SIEH
ANDERSON
STAENDER
ANDERSON
ARGUS
BELSON
MELCHIOR
VIERECK
VIERECK
PEGAU
SHETLER
KESSEL
KESSEL
KESSEL
KESSEL
KESSEL
LITTLE»
SPETZMAN
SPETZMAN
SPETZMAN
BORMANN
LINDSAY
SMITH
SUDA
SUDA
SUDA
SUDA
SUDA
WEST
WEST
GEIST
GEIST
SHUSHAN
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
WARD
ARGUS
JR.
JRe
COL NO 4ER8 +
6354
6389
6490
6389
358
3139
S24
8551
6107
4287
12
18306
1613
198
424
4190
423
3897
2°78
18439
1038
10238
5003
1146
1146
661
554
496
558
554
496
5759
689
629
1425
1793
1794
4857
394
1071
1200
SeNe
266
72
5873
61
4497
5954
547
4172
4277
16069
O7AAF
S55
S55
S19
$19
S19
19454
406
407
402
53-271
2253
1846
27966
28166
26166
24966
28066
Tite
7576
SeNe
SG eNe
55P0
5697
5626
54°9
5188
5189
5367
5239
5235
5190
1147
5822
NO
GWA
SWA
GWA
SWA
IISc 255878
TSC 255853
Isc 255850
SWA
GWA
ALA 32591
ALA 32294
OTF 1104
SWA
GWA
ALA 5976
ALA 5223
TSC 247217
CAN
CAN
CAN
SWA
ALA 27306
TSC 255851
ESLC
ALA 8390
SWA
ALA 5189
ALA 5159
ALA 5628
ALA 5157
Tsc 255902
rsc 255901
SWA
ALA 22494
SWA
ALA 11607
ALA 11597
SWA
TSC 247205
ALA 5955
ALA 27069
GWA
ALA 488
ALA 32696
TSC 255849
ALA 27555
ALA
ISC 255837
BWA
ALA 28876
SWA
ALA
ALA 13224
SWA
ALA 4269
ALA 5649
ALA 228671
ALA 22869
ALA 5048
GWA
GWA
TSC 255839
ISC 255838
TSC 255840
US 2385823
TSC 224879
ALA 10646
GWA
GWA
GWA
GWA
GWA
GWA
SWA
ALA 27709
ALA 28946
WIS
GwA
GWA
SWA
SWA
GWA
BWA
GWA
BWA
GWA
GWA
TSC 224878
SwA
265
Appendix B
TaxON SPC S V HYB PROV wUAnN LAT LONG
122 2 ALAXENSIS VAR. LONGISTYLIS
1873 122 2 2 2 5935 13629
1873 122 2 2 2 5935 13630
1873 122 2 2 14 5822 12955
1873 122 02 2 16 5849 12507
1873 122 0 2 2 22 5714 12243
1673 122 0 2 2 22 5714 12243
lu73 122 02 7 339 6205 12735
1473 122 2 7 368 6132 12520
1u753 122 0 2 7 388 6050 12337
ls73 122 0 2 12 4 68035 13936
1873 122 2 12 5 6734 13832
1873 122 0 2 12 22 6335 13554
1873 122 0 2 12 32. 6122 13859
1673 12202 12 35 6150 13300
1873 122 0 2 12 40 6047 13738
18673 12202 12 4G 6047 13738
1873 122 02 12 41 ©043 13500
1873 122 0 2 12 41 ©6043 13500
1873 122 0 2 12 41 6043 13503
1873 122 0 2 12 44 ©6003 12840
1873 122 0 2 12 44 6003 12840
1873 122 0 2 12 44 6003 12855
1873 122 0 2 14 3 5555 135001
1u73. 122 0 2 14 37 5555) 13001
1873 122 2 14 6 5648 13258
1873 «122 2 14 10 5859 13606
1673 122 2 14 10 5859 13606
18735 122 2 14 10 5859 13606
1875 12e 2 14 10 5859 13606
1873 122 2 14 10 5859 13606
1873 122 cs 14 10 5859 13606
1873 122 2 14 10 ©5859 13606
1873 122 2 14 10 5859 13606
1873 122 2 14 10 5859 13606
1673 122 2 14 10 65857 13602
18673 12202 14 10 5855 13605
1873 122 02 14 10 65855 13603
1873 122 02 14 lo 5855 13603
1873 122 02 14 lo 5855 13603
1873 122 02 14 10 5855 13603
1873 122 2 14 10 5855 13603
1873 122 2 14 lo 5855 13603
1873 122 2 14 10 5855 13603
1873 122 2 14 10 5855 13603
1873 122 2 14 lo 5855 13603
1873 «122 2 14 lo 5855 13603
1873 122 2 14 10 ©5855 13603
1873 122 2 14 10 5855 13603
1873 122 02 14 11 5818 13424
1673 122 02 14 11 «45818 13424
1873 122 02 14 11 5818 13424
1873 122 2 14 11 5826 13435
1873 «122 2 14 11 5826 13435
1873 «122 2 14 11 5825 13540
1873 122 02 14 34 5746 15312
1873 122 02 14 42 5820 15505
1873 122 rd 14 45 5925 136035
1873 «122 2 14 45 5928 13602
1673 122 2 14 45 5928 13605
18673 122 2 14 45 5907 13520
1873 122 0 2 14 45 5927 13518
1873 «122 2 14 46 5952 13945
1673 122 2 14 46 5915 13830
1873 122 2 14 46 5915 13830
1673 122 2 14 46 5952 13945
18735 122 2 14 46 5915 13830
1873 12202 14 62 6001 15141
1673 122 02 14 63 6055 14938
187353 122 02 14 63 6045 14915
1873 12202 14 67 6126 14255
1873 122 02 14 68 6107 14616
1873 12202 14 68 6150 14731
1873 1220 2 14 68 6157 14517
1673 122 02 14 69 6140 14900
1873 12202 14 69 6117 14858
1873 122 02 14 69 6157 14710
1673 122 02 14 70 «©6157 15111
1873 «122 2 14 71 6152 15433
1873 «122 2 14 73 6132 15927
1673 12202 14 73 6132 16018
1873 122 02 14 73 6132 16018
1673 122 0 2 14 79 6259 15604
1673 122 02 14 79 6259 15604
18673 122 2 14 60 6232 15541
1673 122 2 14 80 6257 15536
1673 122 2 14 80 6232 15545
1875 12202 14 80 6258 15536
266
LOCALITY
THREE GUAROSME
INSPECTOR CREE
DEASE L VILLAG
RACING RIVER
SIKANNI RIVER
SIKANNI RIVER
BRINTNELL L
Se NAHANNI
LIARD RIVER
OLD CROW R CN
OLD CROW Re
MAYO.
KLUANE LAKE
LOWER LAPIE R
ALSEK RIVER
ALSEK RIVER
WHITEHORSE
WHITEHORSE
WHITEHORSE
WATSON LAKE
WATSON LAKE
UPPER LIARD R
HYDER
HYDER
PETERS3URG.
MUIR INLET?
MUIR INLET?
MUIR INLET?
MUIR INLET?
MUIR INLET?
MUIR INLET?
MUIR INLET?
MUIR INLET?
MUIR INLET?
MUIR INLET?
MUIR INLET
MUIR INLET
MUIR INLET
MUIR INLET
MUIR INLET
MUIR INLET?
MUIR INLET?
MUIR INLET?
MUIR INLET?
MUIR INLET»
MUIR INLET?
MUIR INLETe
MUIR INLET»?
JUNEAU
JUNEAU
JUNEAU
MENDENHALL GLA
MENDENHALL GLA
GUSTAVUS.
TERROR BAY
UPPER KNIFE CR
KLEHINI RIVER
MOSQUITO LAKEe
MOSQUITO LAKE
HAINES+* MIe 6
SKAGWAY
YAKUTAT BAY.
TANIS LAKE
TANIS LAKE
YAKUTAT BAY.
TANIS LAKEe
DEEP CREEK
HOPE
BERTHA CREEK
MCCARTHY
VALDEZ
SHEEP MT
COPPER CENTER
MATANUSKA VY
EKLUTNA GL
EUREKA LODGE
SKWENTNA
HEAD OF BIG Re
ANIAK REGION
KALSKAG
KALSKAG
TAKOTNA
TAKOTNA
KUSKOK@IM Re
MCGRATH
KUSKOKWIM Re
MCGRATH
Re
GL
GL
GL
GL
GL
GL
GL
GL
DATE
14
12
13
“Tf
20
20
20
28
03
24
18
08
13
15
24
24
01
01
13
25
25
25
04
19
01
01
o oooooo0ooooco°cdeso oo oocoo
DONNNNNANNNN EF UUDADAAOAKAHOSOTHNOKMBDWIAAWHNVNDNN
06
967
967
969
960
943
943
939
970
961
952
970
944
944
944
944
O44
944
944
95R
966
966
966
924
924
967
967
967
967
967
967
967
967
967
967
967
967
967
967
967
967
967
967
967
967
967
967
967
967
967
941
967
967
967
967
950
954
967
967
967
967
961
967
967
967
967
967
967
941
967
961
935
948
961
940
965
965
961
950
949
930
930
941
941
949
949
949
948
COLLECTOR NAME
PZEZEEZECCTFTLFFEEMUYVYVEOTMCCMEKClFADADADADMAADAVN DVN TAN ONCCCAHAAAAVGAVAGVOANANVNAANDNMNANAVNMANOANVDVDOAAGQOOOZPFPPIITPICCcCozoTITcx~ono
z=
NevurzCZeeseD>
=
=
zzecznne
=
Ww
REEECVVV0EECLCSECZLEZEELTETEETCEEETCZEEE
FETT UVIAIVELZFrOCrTrCeVvurr vs za2 cz z£2 Et EELS
ARGUS
ARGUS
RIGBY
CALDER
RAUP
RAUP
RAUP
SCOTTER
CODY
GEIST
RIGBY
ANDERSON
RAUP
POPSILD
RAUP
RAUP
PORSILD
PORSILD
STIELL
ARGUS
ARGUS
ARGUS
WHITED
WHITED
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ANDERSON
ANDERSON
ANDERSON
ARGUS
ARGUS
ARGUS
CLARK
SCHALLER
ARGUS
ARGUS
ARGUS
ARGUS
LITTLE»
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
VIERECK
ANDERSON
HARMS
LITTLE»
ANDERSON
ANDERSON
LITTLE»
PALMER
WELSH
WELSH
LITTLE
DRURY
DRURY
MILLER
MILLER
ANDERSON
ANDERSON
DRURY
DRURY
DRURY
LAYDEN
JRe
JRe
JRe
JR.
COL NO HFRS + NO
6RO02
6735
O74A
24649
19156
10157
9149
12468
11945
14
3
12441
954
11887
11897
9145
9146
6
5076
5024
5028
1151
1152
6A38
6509
6496
6498
6491
6492
6494
6495
6478
6493
6421
6391
6392
4393
6397
6398
6392
6291
63A3B
6293
6388
6387
6387
6371
6422
6410
6422
6462
6666
6577
SG eNe
6689
6474
6683
6747
18118
6134
6275
6°76
6120
6307
A261
6536
6062
18377
1865
19670
18383
342
4250
4460
1A434
3898
1611
265C
265C
7416
7416
2149
1371
2136
200A3
1134
19745
19744
14806
27954
92302
255871
19750
255897
19751
255898
255879
124371
124370
255875
255900
1362
255854
2977
255842
34636
255877
255858
5163
247436
29981
255874
2856
255903
490
255857
Specimens Cited
TAXON SPC S V HYB PROV QUAD LAT LONG LOCALITY
1873 122 02 14% 84 6242 14358 SLANA
1873 122 0 2 14 84 6240 14130 ALA HWY M 1230
1673 122 0 2 14 84 6222 14300 NABESNA ROAD
1673 122 0 2 14 86 6347 14545 DONNELY DOME
1873 122 0 2 14 B86 +=6347 14430 GEORGE LAKE
1873 122 0 2 14 86 6345 14547 DONNELLY DOME
Mis 22 2 14 89 6315 15517 NIXON FORK
1h7s 122° 2 14 95 6434 16522 ANVIL MT.
1473 122 0 2 14 96 6445 16150 MOSES POINT
1673 122 0 2 14 95 ©6402 16055 EGAVIK
1673 122 0 2 14 96 6402 16055 EGAVIK
1673 122 0 2 14 97 6445 15857 GALENA
1473 122 0 2 14 97 6420 15843 KALTAG
1873 122 0 2 14 97 6420 15843 KALTAG
1873 122 0 2 14 97 6420 15843 KALTAG
1873 122 0 2 14 97 6420 15843 KALTAG
1673 122 0 2 14 97 6420 15843 KALTAG
1873 122 0 2 14 97 6420 15843 KALTAG
1673 122 0 2 14 97 6420 15843 KALTAG
1673 122 0 2 14 100 6451 14743 FAIRBANKS
1873 122 0 2 14 100 6452 14750 COLLEGE
1873 122 0 2 14 100 6455 14745 GOLOSTREAM CR
1873 122 0 2 14 100 6452 14750 COLLEGE
1673 122 0 2 14 100 6452 14747 COLLEGE
1873 122 0 2 14 100 6451 14759 COLLEGE
1873 122 0 2 14 100 6452 14747 COLLEGE
1673 122 0 2 14 100 6452 14747 COLLEGE
1873 122 0 2 14 100 6452 14747 COLLEGE
1873 122 0 2 14 100 6452 14747 COLLEGE
1873 122 0 2 14 100 6452 14747 COLLEGE
1873 122 0 2 14 100 6452 14747 COLLEGE
1873 122 0 2 14 100 6452 14750 COLLEGE
1873 122 0 2 14 100 6451 14748 COLLEGE
1673 122 0 2 14 100 6502 14740 ENGINEER CR
1873 122 0 2 14 100 6451 14743 FAIRBANKS
1673 122 0 2 14 100 6451 14743 FAIRBANKS
ia73 tee oS 14 100 6451 14743 FAIRBANKS
1873 122 0 2 14 100 6451 14748 FAIRBANKS
lo73 122 0 2 14 100 6451 14743 FAIRBANKS
1873 122 0 2 14 100 0452 14752 UA EXP. FARM
1073 1220 2 14 100 6452 14752 U A EXP. FARM
1873 122 0 2 14 100 6452 14752 U A EXP. FARM
1873 122 0 2 14 100 6452 14752 UA EXP. FARM
1673 122 0 2 14 100 6443 14710 MOOSE CREEK
1873 122 0 2 14 100 6443 14710 MOOSE CREEK
1673 122 0 2 14 100 6451 14752 SMITH LAKE
1473 122 0 2 14 100 6451 14752 SMITH LAKE
1873 122 0 2 14 100 6446 14816 NENANA MI 339
1873 122 0 2 14 100 6446 14816 NENANA MI 339
1673 122 0 2 14 100 6446 14816 NENANA 40 339
1873 122 0 2 14 100 6451 14748 COLLEGE
1673 122 0 2 14 100 6444 14809 FAIRBANKS
1673 122 0 2 14 100 6444 14809 FAIRBANKS
1873 122 0 2 14 101 6404 14507 GERSTLE R
1673 122 0 2 14 104 6545 14430 N EAGLE SUMMIT
1673 122 0 2 14 104 6545 14430 N EAGLE SUMMIT
1873 122 0 2 14 104 6545 14430 N EAGLE SUMMIT
1673 122 0 2 14 104 6527 14526 EAGLE CR CAMP
1873 122 0 2 14 104 6527 14526 EAGLE CR CAMP
1873 122 0 2 14 104 6527 14526 EAGLE CR CAMP
1673 122 0 2 14 104 6527 14526 EAGLE CR CAMP
1673 122 0 2 14 104 6557 14408 CIRCLE 4OKM S
1873 122 0 2 14 105 6507 14735 CHATANIKA R
1673 122 0 2 14 105 6507 14735 CHATANIKA R
1673 122 0 2 14 106 6527 14815 TOLOVANA R BR
1873 122 0 2 14 117 6654 15141 BETTLES
1673 122 0 2 14 121 6723 14350 SMALL LAKE
1873 122 0 2 14 124 6725 15007 WISEMAN
1873 122 0 2 14 124 6725 15006 WISEMAN
HYBRIDS
15 PEDICELLARIS 71 ATHABASCENSIS
187315 71 11 35 5237 10356 MCKAGUE
187315 71 12 44 6003 12840 WATSON LAKE
1873 15 71 12 44 6003 12840 WATSON LAKE
28 3 PLANIFOLIA SSP+ PULCHRA x 109 SCOULERIANA
1873 28 33109 14 80 6217 15612 KUSKOKWIM R
1873. 28: 33:109 14 89 6315 15442 NIXON MINE
1873 283 109 14 100 6452 14752 SMITH L
DATE
19
24
07
05
06
06
08
06
6
6
06
08
08
06
07
07
07
07
07
07
07
05
08
06
06
947
961
966
957
964
965
950
O71
965
931
931
961
931
931
931
931
931
931
931
940
957
966
S57,
966
957
966
966
966
966
966
966
957
932
964
931
931
933
963
953
964
964
964
964
964
964
967
967
967
966
967
959
966
965
963
951
951
951
965
966
965
966
965
B57
957
965
3939
957
962
949
93A
966
966
949
949
967
COLLECTOR NAVE COL NO
DUTILLY 216?7
E.G EITTEEe JR. 182?6
St WELSH 5652
G6 w ARGUS 10464
V Lt HARMS 3146
V L HARMS 3542
Ww H DRURY 3623
M WILLIAMS 2686
M L HENRY 10
C H ROUSE 19
C YH ROUSE 19
Ext EMRTLEsedR. 19504
C 4 ROUSE 44
C H ROUSE Gy
C 4 ROUSE Gy
G H ROUSE 9
G H ROUSE 39
G H ROUSE 40
G H ROUSE 40
J P ANDERSON 60P2
G wW ARGUS 1202
G W ARGUS 509?
G Ww ARGUS 1291
G w ARGUS 5120
G w ARGUS 1158
G w ARGUS 5116
G w ARGUS 5113
G6 Ww ARGUS 5108
G w ARGUS 5119
G wW ARGUS 5110
G W ARGUS 5114
G w ARGUS 12N2
G W GASSER 4Ag
V tL HARMS 2635
L J PALMER 10
L J PALMER 22
L J PALMER 169
1] SCHUCK V2
S 6 SMITH 143
L A VIERECK 7144
L A VIERECK 7193
L A VIERECK 7146
L A VIERECK 714
L A VIERECK 7148
L A VIERECK 7156
L A VIERECK 8285
L A VIERECK R1e5
L A VIERECK RORTA
L A VIERECK 7877
L A VIERECK R270
L A VIERECK 4806
L A VIERECK 7909
L A VIERECK 7540
L A VIERECK 7198
Jt BUCKLEY SeNe
J L BUCKLEY SeNe
J tL BUCKLEY SeNe
JN TRENT 4965
JN TRENT 366
JN TRENT 3565
JN TRENT 566
L A VIERECK 7793
G wW ARGUS 1155
G w ARGUS 1155
V L HARMS 4650
G T JOHNSON
S SHETLER ROSAF
R BROCKMAN
L H JORDAL 232
A J BREITUNG SeNe
G WwW ARGUS 5050
Y SUDA 13566
w H DRURY 2102
w H DRURY Z377A C
L A VIERECK BONG
H=28 + NO
TSC 255836
GWA
GWA
GWA
ALA 32606
ALA 32638
CAN
GWA
ALA 30404
ALA 25459
ALA 2058
SWA
ALA 25452
ALA 2059
ALA 2078
ALA 2077
ALA 25467
ALA 2075
ALA 25468
ISC 255876
SWA
GWA
GWA
SwA
SWA
GWA
GWA
SWA
GWA
GWA
SWA
ALA 6729
ALA
ALA 32625
ALA 5968
ALA 3975
ALA 5270
ALA 26175
GWA
FSLC 282
ESLEC 282
FSLC 270
FSLC 270
FSLC 272
ESLC 272
GWA
SWA
GWA
GWA
SwA
GWA
BWA
GWA
FSLC 55
GWA
ALA 26815
ALA 5030
ALA 30703
ALA 32697
ALA 306790
ALA 32694
GWA
GWA
ALA 6805
ALA 32598
TSC 255855
ALA 4227
ALA 28488
TSC 255856
CAN 49131
GWA
GWA
rAN
CAN
GWA
267
Appendix B
TaxON SPC S V HYB PROV
4S
18753
4S
1873
1873
14673
45
1675
45
1673
1873
1873
1873
1873
1873
1673
71
1873
1873
1873
1673
1873
1673
1873
1573
1873
1873
1873
1873
1873
1873
1873
1673
1873
18673
1873
1873
1673
1873
1673
1873
1873
1673
18673
1873
1873
1873
1873
1873
1673
1673
1873
1473
1873
1673
87
1873
87
1875
87
1673
1873
1675
268
ARCTICA X
45 49
ARCTICA X
45 83
45 63
4S 63
ARCTICA X
45 67
ARCTICA x
45 118
45 118
45 118
45 1168
45 118
45 118
45 118
ATHABASCENSIS X
71 15
71 15
71 15
7i 15
71 15
v1 15
71 15
71 15
a1 15
71 15
71 15
71 15
e 15
71 15
71 15
71 15
ret 15
71 15
71 15
71 2S
Lip ' 15
71 15
71 15
71 15
71 15
71 15
71 15
71 15
71 15
71 i5
71 15
71 15
71 15
71 15
71 15
71 15
71 15
71 15
BARCLAYI
87 s2
BARCLAYI
87 103
BARCLAYI
67 1168
87 118
8&7 118
49
14
83
14
14
14
67
14
118
02
02
14
14
14
14
14
x 116
14
14
14
wUAD LAT LONG
GLAUCA
13° S255 L7259
OVALIFOLIA
111 6546 16855
129 0806 16545
151 7010 14650
BARCLAYI
11 5818 15423
STOLONIFERA
2 5935 13629
295955 1135629
10 5859 13606
10 5858 13606
10 5859 13606
10 5859 13606
lo 65859 13606
36 5250 11804
36 5253 11805
52 4954 09920
391 6059 11715
4 5954 10205
4 5954 10205
35.5257 10556
35 5237 10356
35 5237 10356
35 5237 10356
35 5237 10356
35 5237 10356
35 5237 10356
35 5237 10356
35 5237 10356
35 5237 10556
35 5237 10356
35 5237 10356
35 5237 10356
35 5237 10356
35 5237 10356
35 5237 103556
35 5237 10356
35 5237 10356
35 5237 10356
35 5237 10356
35 5237 10356
44 6003 12840
44 6003 12840
44 6003 12840
44 6003 12840
44 6003 12840
44 6003 12840
44 6003 12840
44 6003 12840
44 6003 12840
85 6322 14339
85 6322 14339
COMMUTATA
11 5825 13432
LANATA
86 6330 14550
STOLONIFERA
010 5858 13606
010 5859 13606
U10 5859 13606
LOCALITY
ATTU ISLAND
LTL DIOMEDE IS
OGOTORUK CREEK
BULLEN
MT ROBERTS TR
MT GLAVE
MT GLAVE
MUIR INLET?
MUIR INLET?
MUIR INLET?
MUIR INLET?
MUIR INLET?
GL
GL
GL
GL
GL
15 PEOICELLARIS
ATHABASCA R
JASPER
CARBERRY
ENT=MACK HY 44
QUILLWORT LAKE
QUILLWORT LAKE
MCKAGUE
MCKAGUE
MCKAGUE
MCKAGUE
MCKAGUE
MCKAGUE
MCKAGUE
MCKAGUE
MCKAGUE
MCKAGUE
MCKAGUE
MCKAGUE
MCKAGUE
MCKAGUE
MCKAGUE
MCKAGUE
MCKAGUE
MCKAGUE
MCKAGUE
MCKAGUE
MCKAGUE
WATSON
WATSON
WATSON
WATSON
WATSON
WATSON
WATSON
WATSON
WATSON LAKE
ALA HWY M 1335
ALA HWY M 1335
LAKE
LAKE
LAKE
LAKE
LAKE
LAKE
LAKE
LAKE
MENDENHALL GL
FALLS CREEK
GL
GL
GL
MUIR INLET?
MUIR INLET?
MUIR INLET?
DATE
15
14
‘4
03
09
14
14
01
01
01
01
01
30
30
12
16
28
28
31
16
20
26
18
se
20
21
30
26
30
31
09
28
31
19
18
20
31
28
28
25
25
25
25
25
25
26
25
25
08
08
07
12
29
01
01
08
NNNNNNWNGN
07
07
06
07
07
07
05
08
06
05
08
06
06
06
07
05
07
05
06
05
05
06
08
06
05
05
05
06
06
06
06
06
06
06
06
06
08
08
08
06
NNO
945
926
966
966
967
967
967
967
967
967
967
967
917
917
906
959
962
962
939
939
939
939
939
939
939
939
939
939
939
939
940
939
939
939
939
939
939
939
939
966
966
966
966
966
966
966
966
966
944
944
967
951
967
967
967
COLLECTOR NAME
DOADONDANDD
GFerI<x<<<QOOOODOD PHF PPS FTP YPFPrPPPrrPrPYPPPrPrPPrPrrraqnctc sec
w
=<
cCEtEzzezEELE
o¢=
ZEZEZZZEZEAGCCCCCCCCCCCCCCaOCOCGKCOCE ES
<
SOULE
PORSILD
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
MACOUN
MALTE
MACOUN
THIERET
ARGUS
ARGUS
BREITUNG
BREITUNG
BREITUNG
BREITUNG
BREITUNS
BREITUNG
BREITUNG
BREITUNG
BREITUNG
BREITUNS
BREITUNG
BREITUNG
BREITUNS
BREITUNG
BREITUNG
BREITUNS
BREITUNG
BREITUNG
BREITUNG
BREITUNG
BREITUNG
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
SUDA
SUDA
SUDA
RAUP
RAUP
VIERECK
CODY
ARGUS
ARGUS
ARGUS
COL NO HFRR + NO
374
1676
5967
5765
6642
6798
6795
6525
6472
6500
6499
6516
SeNe
SG eNe
SeNe
54n2
8B0K2
8R0F2
in6
438
167
R2
44s
166
168
174
363
B82
363
196
542
94
104
166
44s
167
106
94
94
5051
5045
5048
5051
5045
5048
14446
13366
13466
12713
12713
8638
5114
6467
6500
6520
TSC 255823
TSC 256225
GWA
SWA
GWA
SWA
SWA
GWA
GWA
GWA
SWA
GWA
95377
122167
70266
CAN
CAN
CAN
GWA
NAO
GWA
SAO
AO
AO
DAO
DAO
AO
NAO
DAO
DAO
CAN
CAN
CAN
CAN
CAN
CAN
CAN
cAN
CAN
SASK
SASK
DAO
SASK
SASK
SASK
GWA
GWA
GWA
SASK 33993
SASK 33990
SASK 33992
CAN 278798
ALA 19484
49050
49057
49056
49135
49051
49052
49059
33706
33705
33988
33985
33996
GWA
CAN
GWA
GWA
GWA
Specimens Cited
loi 4
1875
1873
18675
1873
1873
1875
1873
1873
1873
18735
1673
1873
1875
18735
1875
1673
18735
1575
18753
1873
1673
1873
1873
1873
1873
1873
1673
1873
1873
1875
1873
92
14675
1875
1873
18735
1875
16735
1675
1873
1873
1873
1875
105 1
1875
18735
18735
1875
1875
1873
1873
1873
1873
1875
1873
1673
1873
1873
16735
1873
18675
1673
1875
1673
1873
122
TAXON SPC S V HY¥d PROV QUAD LAT LONG LOCALITY
BRACHYCARPA SSP» NIPHOCLADA X 49 GLAUCA
1014 49 12 4 6826 13845 DOG CREEK
1014 49 12 4 68338 13945 BABBAGE RIVER
101 4% 49 12 32 6103 13831 KLUANE L
1014 49 14 71 6155 15425 HEAD OF BIG Re
1014 49 14 80 6233 15336 FAREWELL L-
1014 49 14 86 6357 14547 RICH HWY M249
1014 49 14 687 6340 14930 TEKLANIKA R
1014 49 14 87 6340 14930 TEKLANIKA R
1014 49 14 687 6340 14930 TEKLANIKA R
101 4 49 14 87 6340 14930 TEKLANIKA R
101 4 49 14 87 6340 14930 TEKLANIKA R
1014 49 14 87 6340 14930 TEKLANIKA R
1014 49 14 87 6340 14930 TEKLANIKA R
101 4 49 14 88 6330 15002 TOKLAT R
1014 49 14 88 6328 15010 HIGHWAY PASS
101 4 49 14 88 6328 15010 HIGHWAY PASS
101 4 49 14 68 6©331 15004 TOKLAT R BAR
1014 49 14 88 6325 15035 MULDROw MORAIN
101 4 49 14 88 6325 15035 MULDROW MORAIN
1014 49 14 88 6325 15035 MULDROW MORAIN
1014 49 14 88 6325 15035 MULDROW MORAIN
101 4 49 14 88 6325 15035 MULDROW MORAIN
101 4 49 14 88 6325 15035 MULDROW MORAIN
101 4 49 14 100 6452 14750 COLLEGE
1014 49 14 100 6452 14750 COLLEGE
101 4 49 14 101 6412 14600 SHAW CR FLATS
1014 49 14 124 6725 15007 WISEMAN
1014 49 14 124 6725 15007 WISEMAN
1014 49 14 124 6725 15007 WISEMAN
1014 49 14 124 6725 15007 WISEMAN
101 4 49 14 124 6725 15007 WISEMAN
PHLEBOPHYLLA X 105 ROTUNDIFOLIA
92 105 14% 41 5842 15640 NAKNEK
92 105 14 104 6528 14527 EAGLE SUMMIT
92 105 14 104 6548 14639 VICTORIA MT
92 105 14 124 6725 15006 WISEMAN
92 105 14 145 6902 16350 CAPE BEAUFORT
92 105 14 145 6902 16350 CAPE BEAUFORT
92 105 14 152 7005 14335 BARTER ISLAND
92 105 14 153 7120 15640 BARROW
92 105 14 153 7120 15640 BARROW
92 105 14 153 7120 15640 BARROW
92 105 14 153 7120 15640 BARROW
ROTUNDIFOLIA SSP. ROTUNDIFOLIA X 92 PHLEBOPHYLLA
105 1 «92 O07 152 6813 13547 CANOE LAKE
105 1 92 14 866 6355 14730 DRY CREEK
1051 92 14 129 6806 16545 OGOTORUK CREEK
105 1 92 14 138 6950 14220 NUVAGAPAK PT
105 1 92 14 138 6950 14220 NUVAGAPAK PT
105 1 92 14 138 6950 14220 NUVAGAPAK PT
105 1 92 14 138 6950 14220 NUVAGAPAK PT
105 1 92 14 145 6902 16350 CAPE BEAUFORT
105 1 92 14 145 6902 16350 CAPE BEAUFORT
105 1 92 14 147 7030 15730 MEADE RIVER PO
105 1 92 14 151 7010 14650 BULLEN
105 1 92 14 151 7010 14650 BULLEN
105 1 92 14 151 7010 14650 BULLEN
1051 92 14 151 7010 14650 BULLEN
105 1 92 14 152 7005 14335 BARTER ISLAND
105 1 92 14 152 7005 14335 BARTER ISLAND
1051 92 14 152 7005 14335 BARTER ISLAND
105 1 92 14 152 7005 14335 BARTER ISLAND
105 1 92 14 153 7125 15630 POINT BARROW
1051 92 14 153 7125 15630 POINT BARROW
1051 92 14 153 7120 15640 BARROW
1 ALAXENSIS VAR. ALAXENSIS X 103 LANATA
122 01103 14 129 6815 16600 IBRULIKORAK CR 10
186735
DATE
20
25
il
09
09
07
970
970
944
950
949
948
956
956
956
956
956
956
956
956
956
956
960
939
939
939
939
939
939
957
957
964
939
939
939
939
939
949
964
953
949
966
966
966
966
966
967
966
964
962
966
966
966
966
966
966
966
966
966
966
966
966
966
966
966
966
966
966
966
960
COLLECTOR NAME
CCCCCrODP YEP PFrFPAMAAVVADODAOVOVOOCZE IVY
<DQOQDODODOFrVUC A
DOAODADADAIANGDNDANDNADAVNGVNGDAANYVADLY Cc
ZEEEZELEZCLZEAETVISE Serre
VUVUVUVUPESE
ZSze£cezztET ag
ZTETECECECCTECTEAZESTCECLEEECEP>yS
WELSH
WELSH
RAUP
DRURY
DRURY
ANDERSON
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
MURIE
NELSON
NELSON
NELSON
NELSON
NELSON
NELSON
ARGUS
ARGUS
VIERECK
ANDERSON
ANDERSON
ANDERSON
ANDERSON
ANDERSON
RAUP
HARMS
KLEIN
JORDAL
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
SUDA
LARSEN
VIERECK
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
BELSON
COL NO HFRS + NO
10964
10409
12403
4019A
241i
10859
607
605
67
87
OTF 1188
OTF 1191
ALA 19946
CAN
CAN
TSC 256687
ALA 4579
ALA 4577
ALA 22464
GWA
SWA
Ry
Qu
ALA 4695
GWA
Ry
SWA
ISC 256219
ALA 513
GH
RM
NA
US
ALA 16314
GWA
FSLC 286
1SC 256550
TSC 256548
ALA 27558
NIA
NA
US 2176353
32647
23092
256251
ALA
ALA
1sc
GWA
SWA
GWA
GWA
SWA
GWA
GWA
GWA
BWA
GWA
GWA
GWA
GWA
SWA
SWA
GWA
GWA
GWA
GWA
GWA
GWA
GWA
GWA
GWA
GWA
GWA
GWA
GWA
ALA 28687
269
Appendix B
TAXON
122
1873
1873
1873
1873
1673
1875
1873
1873
1873
1873
1875
1873
1873
1875
1875
1875
1873
1873
1875
1873
1873
1873
1875
1873
270
SPC S V HYB PROV QUAD LAT LONG LOCALITY DATE
2 ALAXENSIS VAR. LONGISTYLIS X 122 ALAXENSIS
122 e ice 2 4 5847 12502 MACDONALD CR jel
122 eee ie 10 6003 12840 WATSON LAKE 25: 16
122 eee” Lo 10 5837 13625 GEIKIE INLET rake )denales
122 eter 9 34 5713 15318 OLD HARBOR 6
122 eked 1% 46 5908 13822 ALSEK R FLATS 10 6
122 eutee (2S 69 6157 14710 EUREKA LODGE 28 6
122 ee Le 86 6320 14535 RAINBOW MT 30) "6
122 2i22° 14 94 6430 16525 NOME eG
122 2. 122 "34 94 6455 16508 SALMON LAKE 14 #7
122 2122 * 19 96 6402 16055 EGAVIK ah fs
122 enwte2 Ih 96 6402 16055 EGAVIK 11 8
122 2122 is 96 6402 16055 EGAVIK 5 Ut bes)
122 2 dae2 4 96 6402 16055 EGAVIK 118
122 2122 14 100 6452 14750 COLLEGE Lent /
122 21e2 14 100 6452 14750 COLLEGE ha axle
122 2122 14 100 6452 14750 COLLEGE Uren
122 21c2 14 100 6452 14750 COLLEGE VS ar,
122 2122 14 100 6452 14750 COLLEGE LS. 7,
122 21e2 14 100 6452 14750 COLLEGE AN 1
122 2122 14 100 6430 14905 NENANA DME ye uss
122 2122 14 101 6522 14556 TWELVE MI CR Li. 6
122 2i1e2 14 101 6410 14455 GERSTLE RIVER 16 7
122 21c2 14 101 6410 14455 GERSTLE RIVER 31 7
122 2122 14 101 6432 14703 LTL SALCHA R PAU iS)
COLLECTOR NAME
960
966
928
964
965
965
966
938
966
931
931
931
931
966
966
966
957
957
966
965
957
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957
966
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CALDER
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ROUSE
ROUSE
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
ARGUS
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SHETLER
SPETZMAN
SPETZMAN
VIERECK
COL NO HE%B + NO
24352
5925
754
29
7602
4460
RM19
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5900
16
18
18
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5109
5115
5117
intea
1012
5118
3720
BEAF
601
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ALA
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ALA
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GWA
ALA
ALA
ALA
ALA
GWA
GWA
GWA
6WA
BLA
GWA
ALA
ALA
ALA
ALA
SWA
17325
255872
26534
246037
32685
255859
2062
2057
25458
25456
6719
32583
3734
6840
6841
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274
Index to Salix
alaxensis, 5, 13, 14, 15, 18, 19,
161, .182, 186, #487, :t68, «169;
264
alaxensis, alaxensis, 5, 10, 14, 15,
16, 19, 20, 29, 172,.2481= 85,
264
alaxensis, longistylis, 5, 29,
182 —85, 187, 188, 189, 266
alaxensis, obovalifolia, 181, 185
alaxensis x lanata ssp. richardsonii,
8, 269
albertana, 175
aliena, 77
amplifolia, 131, 134
anadyrensis, 161
anglorum, 61, 81
anglorum, antiplasta, 81
anglorum, araioclada, 81
anglorum, kophophylla, 81, 84
Arbuscella (Section), 5, 10,
164
arbusculoides, 5, 8, 14, 15, 18,
19, 20, 33, 168-71, 188, 189,
210
arbusculoides, glabra, 169, 171
arbutifolia, 113
mGlieds)(5;° Wy ob8dcothn int he: 18;
19;2:26 50 S2en74;2 175,480 = 85,» 88;
Ba 93.0401 10384 10ze1 Tt, 164,
214
arctica,
arctica,
arctica,
180,
157,
antiplasta, 81, 84
araioclada, 81, 84
arctica, 84
arctica, crassijulis, 81, 83
arctica, kophophylla, 81, 84
arctica, torulosa, 81, 84
arctica x barclayi, 7, 268
arctica x glauca, 7, 85, 268
arctica x _ ovalifolia, 7, 71,
268
arctica
268
arctogena, 6
arctolitoralis, 66
85; 89;
< stolonifera, 7, 75, 76, 85,
arctophila, 5, 10, 19, 26, 85,
108 —11, 229
athabascensis, 5, 10, 18, 34, 99,
101, 102-07, 117, 119, 231
athabascensis x _ pedicellaris, 7, 8,
119, 267
aurita x repens, 6
babylonica, 5, 31, 41-43, 197
balsamifera, 147
Narelayi, 05, (513; 044, . 1Bpo 4832,
75, 120, 127, 128—31, 133, 134,
139; 172;:238
barclayi, angustifolia, 129
barclayi, commutata, 135
barclayi, grandifolia, 129
barclayi, padophylla, 125
barclayi, pseudomonticola, 125
barclayi, rotundifolia, 129
barclayi <x commutata, 8, 268
barclayi x lanata ssp. richardsonii,
8, 268
barclayi x stolonifera, 8, 75, 268
barrattiana, 5, 19, 20, 30, 174-77,
242
bebbiana, 5, 15, 18, 19, 33, 148-53,
197
bebbiana, bebbiana, 151
bebbiana, depilis, 149, 151
bebbiana, perrostrata, 149, 151
behringica, 53, 57
bella, 186, 189
bicolor, 163
blakii, 45
boganidensis, 171
brachycarpa, 5, 19, 93, 101, 247
brachycarpa, brachycarpa, 5, 10, 18,
34, 85, 89-94, 247
brachycarpa, glabellicarpa, 89
brachycarpa, mexiae, 92
brachycarpa, niphoclada, 5, 10, 14,
15, 18, 19, 26, 34, 90-94, 103, 247
brachycarpa, psammophila, 91
brachycarpa, sansonii, 91
brachycarpa ssp. niphoclada x glau-
ca, 7, 269
brachystachys, 153
brachystachys, scouleriana, 153
ealcicolasa73
candida, 5, 10, 18, 33,
183, 200
176 — 79,
275
Index to Salix
candida, denudata, 177
candidula, 177
caprea, 10, 151, 193,°755
capreoides, 153
Caprisalix (Subgenus), 164
Chamaetia (Section), 5, 10, 45
chamissonis, 5, 10, 18, 19, 25, 52,
85, 106-09, 111, 241
chlorophylla, 157
chlorophylla, nelsonii, 157
cinerascens, occidentalis, 149
cinerea, 10
(cinerea X purpurea) xX cCaprea, 6
(cinerea XxX purpurea) Xx (purpurea
x viminalis), 6
coaetanea, 155
commutata, 5, 8, 13, 18, 19, 20,
a1, 75,120, 131,.135— 39,209
commutata, denudata, 135, 139
commutata, mixta, 135
commutata, puberula, 135
commutata, sericea, 135, 139
commutata x myrtillifolia, 139
conjuncta, 129
cordata, 121
cordata, mackenzieana, 123
cordata, monticola, 124
cordata, rigida, 121
cordata x rostrata, 123
Cordatae (Section), 5, 10, 121
coulteri, 191
crassijulis, 81, 83, 84
cuneata, 85
cyclophylla, 67
depressa, 151
depressa, rostrata, 149, 151
desertorum, 95
desertorum, fruticulosa, 89
desertorum, stricta, 89
discolor, 105, 155, 157
drummondiana, 5, 10, 18,
184, 186-89, 264
drummondiana, bella, 186
drummondiana, subcoerulea, 186
19,°30;
exigua, interior, 43, 54
exigua, interior, pedicellata, 43
276
fallax, 103
farrae, 119, 120
farrae, microserrulata, 119
farrae, walpolei, 119
flagellaris, 65
flavescens, 153
flavescens, capreoides, 153
flavescens, scouleriana, 153
fluviatilis, 45
fragilis, 16
fulcrata, subglauca, 159
fumosa, 109
fuscescens, 5, 10, 15, 18, 19, 26,
28, 112-15, 220
fuscescens, hebecarpa, 115
fuscescens, reducta, 113
glacialis, 8, 68
glauca, 5, °‘10,° 14;975;:017, 34; a4
52, 85, 93, 94, 95-103, 105,
107, 164, 222
glauca, acutifolia (Western phase), 5,
18, 19, 95-103, 224
glauca, alicea, 95
glauca, desertorum, 95
glauca, desertorum, sericea, 95
glauca, glabrescens, 95
glauca, glauca (Beringia phase), 5,
18, 19, 95-103, 222
glauca, niphoclada, 91
glauca, perstipula, 95
glauca, poliophylla, 95
glauca, villosa (Rocky Mountain
phase), 5, 18, 19, 95-103, 228
Glaucae (Section), 1, 5, 10, 81
glaucops, glabrescens, 95
glaucops, villosae, 95
glaucosericea, 100
nastata, 5, 70, 15, 18, ‘9:
118-20, 131, 137, 213
hastata, farrae, 119
hastata, psiloides, 119
Hastatae (Section), 5, 10, 119
hebecarpa, 115
herbacea x polaris, 6
hookeriana, 5, 16, 18,
56, 131-35, 230
32,
#99°29; 32}
Index to Salix
humillima, 169
interior, 5, 10, 16,
200
18, 31, 42-45,
krylovii, 179
kurilensis, 88
lanata:-5e 151,ct#3
lanata, richardsonii, 5, 10, 14, 15,
18... 19, «29,. 129, 131; .470:=73,
177, 254
Lanatae (Section), 5, 10, 171
lancifolia, 35
lasiandra, 5, 10, 14, 16, 18, 19,
32, 35-38, 195, 230
lasiandra, lancifolia, 35
lasiandra, lasiandra, 38
lasiandra, recomponens, 35, 38
leiocarpa, 53
lingulata, 92
livida, 153
livida, occidentalis, 149
livida, rostrata, 149
longifolia, 43
longifolia, pedicellata, 43
Longifoliae (Section), 4, 5, 10, 16,
43, 45
longipetiolata, 88
lucida, 37, 41
maccalliana, 5, 10, 18, 20, 33, 38 -— 41,
232
Maccallianae (Section), 4, 5, 10, 38
mackenzieana, 123, 124
mackenzieana, macrogemma, 123
melanopsis, 45
monica, 157
monticola, 5, 10, 18, 19, 29, 124-29,
232
muriei, 92
myrsinites, 109, 111, 143
myrsinites, curtiflora, 140, 143
myrsinites, pseudomyrsinites, 140,
143
myrtillifolia, 5, 10, 14, 18, 27,
oly,c 120, t 12faetttakR) tadcerids:
143, 145-47, 233
myrtillifolia, curtiflora, 140
myrtillifolia, pseudomyrsinites, 8,
140, 143
myrtilloides, 115, 119
myrtilloides, pedicellaris, 115
Myrtilloides (Section), 5, 10, 113
Myrtosalix (Section), 5, 10, 107
nakamurana, 88
nelsonii, 157
niphoclada, 91
niphoclada, mexiae, 92
niphoclada, muriei, 92
novae-angliae, 5, 8, 18, 31, 120,
127, 131, 137, 138, 140-45, 246
novae-angliae, myrtillifolia, 145
novae-angliae, pseudocordata,
143
novae-angliae, pseudomyrsinites, 140,
143
nummularia, 5, 10, 19,
57, 59-61, 234
nummularia, tundricola, 59
nuttallii, 153
nuttallii, capreoides, 153
140,
| a
orbicularis, 45
ovalifolia, 5, 17, 52, 56, 69,
15, 76; 88. 234
ovalifolia, arctolitoralis, 5, 19,
66, 68, 69-71, 236
ovalifolia, camdensis, 65
ovalifolia, cyclophylla, 5, 19,
67—71, 237
ovalifolia, glacialis,
51, 68-71, 236
ovalifolia, ovalifolia, 5, 10, 19,
51, 64-71, 234
70,
28,
28,
5,698) AS pu2sy:
28,
pachnophora, 186
padifolia, 124, 125
padophylla, 125, 129
paleoneura, 61
parallelinervis, 159
pedicellaris, 5, 10, 18, 32,
114-19, 201
pedicellaris, athabascensis, 103
pedicellaris, hypoglauca, 115
277
Index to Salix
pedicellaris, tenuescens, 115
pellita, 186, 187
pennata, 157
pentandra, 5, 31, 35, 201
Pentandrae (Section), 4, 5,
41
perrostrata, 149
phlebophylla, 5, 10, 15, 18, 19,
25...62,, 56, 57, 61=63)74)243
phlebophylla x _ rotundifolia, 8, 56,
269
phylicifolia, 6, 10, 163, 164, 166
phylicifolia, pennata, 157
phylicifolia, planifolia, 157, 158
phylicifolia, subglauca, 159
phylicoides, 159
planifolia, 5, 10,
163, 201
planifolia, monica, 157
planifolia, nelsonii, 157
planifolia, planifolia, 5, 10, 18, 30,
157 — 67, 201
planifolia, pulchra, 5, 10, 14, 15,
holies Bi 30, G3IfLZ7G6iiliase:
159 —67, 172, 204
planifolia, pulchra, yukonensis, 5, 14,
18, 30, 161-67, 203
planifolia ssp. pulchra x scouleriana,
8, 267
polaris, 5, 10, 18,
5/1252
polaris, leiocarpa, 53
polaris, pseudopolaris, 49
polaris, selwynensis, 49, 52
pseudocordata, 140, 143, 145
pseudomonticola, 125
pseudomonticola, padophylla, 125
pseudopolaris, 49, 52
pulchra, 159, 163, 164
pulchra, anadyrensis, 161, 163
pulchra, kalarica, 159, 163
pulchra, looffiae, 159, 167
pulchra, palmeri, 159, 167
pulchra, parallelinervis, 159, 163
pulchra, yukonensis, 161
purpurea, 117
pyrenaica, 101
pyrifolia, 5, 19, 31, 147-49, 201
10, 35,
105 SWS Te O162)
19, 25, 49-52,
278
rectijulis, 107
(repens x phylicifolia) x (viminalis
x phylicifolia), 6
repens x viminalis, 6
reptans, 93, 101
reticulata, 5, 10, 13, 14, 15, 18,
492045; 547,48, 56) 75,9 76)0 250
reticulata, gigantifolia, 45
reticulata, glabellicarpa, 5, 8-9, 18,
25, 46-49, 254
reticulata, glabra, 45
reticulata, orbicularis, 45
reticulata, reticulata, 5, 18, 25, 45-
49, 250
reticulata, subrotunda, 45
reticulata, villosa, 45, 48
retusa, 79
Retusae (Section), 5, 10, 49
richardsonii, 171, 173
rigida, 5, 10, 18, 20,
121 —24, 127, 131, 201
rigida, mackenzieana, 123, 124
rigida, macrogemma, 123
rostrata, 149
rostrata, perrostrata, 149
rotundifolia, 5, 10, 15, 18, 53,
56, 256
rotundifolia, dodgeana, 5, 18, 19,
27, 54-58, 258
rotundifolia, pilosiuscula, 56
rotundifolia, rotundifolia, 5, 18, 19,
27, 53-58, 59, 63, 71, 256
29.932
Salix (Genus), 24
Salix (Subgenus), 164
saxatilis, 109
scouleriana, 5, 10, 17, 18, = 19,
30, 152—57, 192, 258
scouleriana, brachystachys, 153
scouleriana, coetanea, 154
scouleriana, flavescens, 153
scouleriana, poikila, 153, 157
scouleriana, thompsoni, 154
serissima, 37
setchelliana, 5, 14,
260
Sitchenses (Section), 1,
191
18, 27, 77-79,
4," 552° 10;
Index to Salix
sitchensis, 5, 10, 13, 14, 16, 18,
33, 75, 157, 190-92, 261
sitchensis, congesta, 191
sitchensis, coulteri, 191
sitchensis, denudata, 191
speciosa, 181
speciosa, alaxensis, 181
sphenophylla, 5, 10, 19, 20, 26,
28, 85-89, 210
sphenophylla, pseudotorulosa, 85, 88
stagnalis, 153
starkeana, 153
stipulifera, 101
stolonifera, 5, 13, 18, 26, 28, 52,
57, f2—77, 131, 262
stricta, 89
Subalbae (Section), 4, 5, 41
subcoerulea, 186
torulosa, 81, 83, 84, 88
tundricola, 59, 61
vagans, 153
vagans, cinerascens, occidentalis, 149
vagans, rostrata, 149
vestita, 49
Vetrix (Section), 5, 10, 149
villosa, 95
Villosae (Section), 5, 10, 177
viminalis, 10
viminalis x aurita, 6
viminalis xX caprea, 6
viminalis x cinerea, 6
walpolei, 119, 120
xerophila, 149, 153
279
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